Affiliation: | Hedmark University College, Department of Forestry and Wildlife Management, 2480 Koppang, Norway;;Centre for Ecological and Evolutionary Synthesis (CEES), Department of Biology, University of Oslo, PO Box 1066 Blindern, N-0316 Oslo, Norway;and;Norwegian Institute for Nature Research, Tungesletta 2, 7485 Trondheim, Norway |