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Fossil liberation: a model to explain high biodiversity in the Triassic Cassian Formation
Authors:Vanessa Julie Roden  Imelda M Hausmann  Alexander Nützel  Barbara Seuss  Mike Reich  Max Urlichs  Hans Hagdorn  Wolfgang Kiessling
Institution:1. GeoZentrum Nordbayern, Section 2. Palaeobiology, University Erlangen-Nürnberg, Loewenichstr. 28, 91054 Erlangen, Germany;3. SNSB – Bayerische Staatssammlung für Paläontologie und Geologie, Richard-Wagner-Str. 10, 80333 München, Germany

Department of Earth & Environmental Sciences, Ludwig-Maximilians-Universität München, Richard-Wagner-Str. 10, 80333 München, Germany;4. SNSB – Bayerische Staatssammlung für Paläontologie und Geologie, Richard-Wagner-Str. 10, 80333 München, Germany;5. Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany;6. Muschelkalkmuseum Ingelfingen, Schlossstraße 11, 74653 Ingelfingen, Germany

Abstract:With 1429 animal species, the Triassic Cassian Formation in the Dolomites, Southern Alps (Italy), yields the highest species richness reported from any spatially constrained pre-Quaternary formation known to science. The high preserved diversity is partly attributable to a high primary diversity governed by the tropical setting, increasing alpha diversity, and the breadth of habitats spurring beta diversity. More important is the excellent preservation of fossils and the ease with which they can be extracted from the poorly lithified sediments. We propose the term ‘liberation Lagerstätten’ to capture this preservational window. In contrast to conservation Lagerstätten, liberation Lagerstätten like the Cassian Formation originate from normal marine conditions but low-grade diagenesis. Molluscs contribute substantially to species richness, comprising 67% of all invertebrate species in the Cassian Formation. The gastropod dominance (39% of all species) is nearly as great as in Recent tropical settings, contradicting the concept of a substantial Cenozoic rise.
Keywords:diversity  Cassian Formation  gastropod dominance  aragonite preservation  fossil Lagerstätten
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