Subfamily Dichomeridinae (Lepidoptera, Gelechiidae): Phylogeny, classification, and position in the system of gelechiid moths

On the basis of comparative morphological analysis, taking into account the male genital musculature, a cladogram of the tribes and subfamilies of gelechiid moths is proposed. In accordance with the main branches of the cladogram the new system of the family Gelechiidae consisting of 5 subfamilies (Physoptilinae, Anomologinae, Gelechiinae, Anacampsinae, and Dichomeridinae) is developed. The subfamily Physoptilinae is considered as a sister-group to other gelechiids, the monophyly of the latter is supported by a complex of synapomorphies (female retinaculum with anteriorly directed row of scales located on the base of radial stalk; muscles m ₃ running across the longitudinal body axis, connecting the lateral arms of vinculum with margins of juxta; protractors of aedeagus m ₅ divided into two bunches m _5a and m _5b). The subfamilies Anacampsinae and Dichomeridinae are regarded as sistergroups. The monophyly of the subfamily Dichomeridinae is based on the complex of synapomorphies (presence of parategminal sclerites, which are the apodemes for muscles m ₄; presence of distinct ventral wall in tegumen; intrategminal position of the muscles m ₂). The subfamily Dichomeridinae is considered to consist of three tribes, Anarsiini, Chelariini and Dichomeridini, with 34 genera in total. The cladogram for the genera of the subfamily Dichomeridinae is proposed.