Affiliation: | (1) Department of Population Ecology, Zoological Institute, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen, Denmark;(2) Department of Ecology and Genetics, Institute of Biological Sciences, University of Aarhus, 8000 Aarhus-C, Denmark;(3) Present address: Section of Evolutionary Biology, Institute of Evolutionary and Ecological Sciences, University of Leiden, Leiden, The Netherlands |