Affiliation: | Institute of Biological Sciences, University of Stirling, Stirling, FK9 4LA, Scotland, UK;;Integrative Ecology Unit, Division of Population Biology, Department of Ecology and Systematics, PO Box 17 (Arkadiankatu 7), FIN-00014 University of Helsinki, Finland;and;Department of Biological and Environmental Science, University of Jyväskylä, PO Box 35, FIN-40351 Jyväskylä, Finland |