^aSchepens Eye Research Institute, 20 Staniford Street, Boston, MA 02114, USA

^bDepartment of Ophthalmology, Harvard Medical School, Boston, MA, USA

^cDepartment of Physics, University of Massachusetts Boston, Boston, MA, USA

^dCenter for Neurologic Disease, Brigham and Women's Hospital, Cambridge, MA, USA

^eDepartment of Oral Medicine, Infection, and Immunity, Harvard School of Dental Medicine, Boston, MA, USA