Mi Young Noh and Yong Hun Jo equally contributed to this work.;2. Department of Agricultural Biology, Division of Plant Biotechnology, College of Agriculture and Life Science, Chonnam National University, Gwangju, Korea;3. Department of Parasitology, College of Medicine and Frontier Inje Research for Science and Technology, Inje University, Busan, Korea;4. Division of Applied Life Science, Gyeongsang National University, Jinju, Korea;5. Department of Biological Science and the Research Institute for Basic Sciences, Hoseo University, Asan, Korea;6. Jeollanam-do Agricultural Research & Extension Services, Naju, Korea