A systematic study of the pulchra species group of Archips occurring in Japan (Lepidoptera, Tortricidae)

This paper reviews the pulchra species group of Archips, describes a new species, Archips stellata, and redescribes A. pulchra (Butler) and A. abiephaga (Yasuda). Although long confused with each other, these three species are distinguishable by differences in the wing markings, abdominal dorsal pits, and genitalia. To position the pulchra group systematically within the genus Archips, earlier related studies are reviewed and the phylogenetic relevance and polarities of traits are discussed. The forewing pattern, consisting of transverse black and glossy gray fasciae, is presumed to be the only synapomorphy of this group; affinity between the pulchra group and other groups of Archips in some traits may reflect symplesiomorphy.