1. National Key Laboratory of Crop Genetic and Improvement, Hubei Hongshan Laboratory, College of Plant Science and Technology Huazhong Agricultural University Wuhan 430070 China;
2. Guangdong Laboratory for Lingnan Modern Agriculture Wushan Road Guangzhou 510642 China;Correspondences: Zhijuan Wang (wangzj@mail.hzau.edu.cn); Xia Li (xli@mail.hzau.edu.cn, Dr. Li is responsible for the distribution of the materials associated with this article)