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A new species of Gracixalus (Amphibia: Anura: Rhacophoridae) from northern Vietnam
Authors:Truong Quang Nguyen  Minh Duc Le  Cuong The Pham  Tao Thien Nguyen  Michael Bonkowski  Thomas Ziegler
Institution:1. Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam
2. Department of Terrestrial Ecology, Cologne Biocenter, University of Cologne, Zülpicher Strasse 47b, 50674, Cologne, Germany
3. Faculty of Environmental Sciences, Hanoi University of Science, Vietnam National University, 334 Nguyen Trai Road, Hanoi, Vietnam
4. Center for Natural Resources and Environmental Studies, Hanoi National University, 19 Le Thanh Tong, Hanoi, Vietnam
5. Department of Herpetology, American Museum of Natural History, Central Park West at 79th Street, New York, NY, 10024, USA
6. Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam
7. AG Zoologischer Garten K?ln, Riehler Strasse 173, 50735, Cologne, Germany
Abstract:We describe a new species of small tree frog from northern Vietnam based on morphological differences and molecular divergence. Gracixalus waza sp. nov. is distinguishable from its congeners and other small rhacophorid species on the basis of a combination of the following characters: (1) size small (snout-vent length of males 27.1–32.9 mm, of females 37.6 mm); (2) head as wide as or wider than long; (3) vomerine teeth absent; (4) snout rounded and long (16–18 % of the snout-vent length); (5) spines on upper eyelid absent; (6) tibiotarsal projection absent; (7) dorsal skin smooth; (8) dermal fringes on forearm and tarsus absent; (9) dorsal surface of head and body greyish-green to moss-green with dark brown pattern forming an inverse Y marking; and (10) throat and chest with dark marbling. Our molecular data showed that the new species is nested in the same group with Gracixalus jinxiuensis sensu lato.
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