Affiliation: | 1. Centre for Biomolecular Sciences, School of Biology, Biomolecular Sciences Building, University of St. Andrews, North Haugh , St. Andrews , KY16 9ST , UK gai@st-andrcws.ac.uk;3. Centre for Biomolecular Sciences, School of Biology, Biomolecular Sciences Building, University of St. Andrews, North Haugh , St. Andrews , KY16 9ST , UK;4. Plant Research Unit. School of Life Sciences, University of Dundee at SCRI, Invergowrie , Dundee , DD2 5DA , UK |