蒙古新发现的渐新世山河狸和松鼠类化石(英文)

蒙古中部地区的三达河组(HsandaGolFormation)是亚洲渐新世陆相地层的经典地层之一。三达河组的沉积岩层现被分成两个岩性段:上部早渐新世晚期-?晚渐新世山地段(ShandMember)和下部早渐新世早期塔塔尔段(TatalMember);两岩段间有时夹有KharKhoroo玄武岩层(BryantandMcKenna,1995;MengandMcKenna,1998)。该玄武岩层的时代用40Ar/39Ar法被确定为距今31.5Ma(Hcketal.,1999,他们称其为BasaltI)。三达河组中产有丰富的哺乳动物化石,特别是啮齿类化石(MatthewandGranger,1923;Kowalski,1974;RussellandZhai,1987;Hcketal.,1999)。近年来,本文的后一作者在三达河组中又采集到一批啮齿类化石,其中包括松鼠和山河狸2科的化石。本文是对这两类化石新材料的记述。现生的松鼠科分异很大,而且大多数都分布在亚洲。然而,松鼠科化石在古近纪时在北美和欧洲却很丰富。在北美出现于晚始新世,在渐新世时已明显分异;在欧洲出现于早渐新世,在中新世时分异很大。相反,亚洲古近纪的松鼠化石却发现得很少,标本也很破碎。已报道的只有3件标本。2件产自党河下游地区:Bohlin(1946)报道的晚渐新世的Sciurussp.,标本只是2枚单个臼齿(M3和m2);王伴月、邱占祥(2004)报道的早渐新世的党河鲜松鼠(Oligosciurusdangheensis),标本只是一段带m1～2的下颌骨。还有一件是Minjin(2004)报道的蒙古渐新世的三达河克热姆鼠(Kheremhsandgoliensis),标本也只是一段带m1～3的下颌骨。本文报道的松鼠化石产自蒙古国的巴彦洪戈尔省的塔石盖贝(TashgainBel)地点的三达河组上部山地段,时代为早渐新世晚期-?晚渐新世。这批松鼠化石标本包括1件具P3～M3的上颌骨(PC369)和2段下颌骨(PC3610和PC3611)。这是亚洲古近纪目前已知的最好的一批松鼠化石标本,代表松鼠科Kherem属的一新种:亚洲克热姆鼠(Kheremasiaticasp.nov.)。其主要特征是颊齿较大,较宽;下内尖的高度和下次尖的大小从m1到m3变化不大,彼此相近;下内脊较明显;下内尖较明显,与下后边脊间有浅沟分开等。K.asiatica的形态特征表明,Kherem属应归入北美的Cedromurinae亚科。新的发现表明,亚洲渐新世的松鼠已发生了分异,至少可分为3支。以Kherem为代表的一支,与北美早期的cedromurines有较近的系统关系。它可能是由类似于北美Oligospermophilus的种类演化并迁徙到亚洲来的。现生的山河狸只有一种(Aplodontiarufa),仅生活在北美西部沿海地区。但从始新世到中新世,山河狸却遍布全北区。它们在北美较繁盛,分异大,但在亚洲发现得很少。在亚洲渐新世地层中虽曾报道过3属4种山河狸化石,但材料都很少(Prosciurusordosicus和P.?shantungensis均只有一枚单个的牙)。山河狸化石在蒙古渐新世地层的新发现表明,亚洲渐新世的山河狸化石比已知的要丰富。本文描述的山河狸化石分别采自蒙古南戈壁省Shunkt地点的山地段和前杭爱省UlaanKhongil(=TatalGol)地点的塔塔尔段。它们代表原松鼠属的3类:蒙古原松鼠(新种)(Prosciurusmongoliensissp.nov.)、蒙古原松鼠(相似种)(P.cf.P.mongoliensis)和小原松鼠(新种)(P.pisinnussp.nov.)。蒙古原松鼠的主要特征是下颊齿的下中附尖与下后附尖脊连,下中尖发育较弱,下外中脊弱或无;p4无下前边尖和下前齿带;m1具从下外脊伸向下次脊的小刺;m2和m3的下后脊II较发育;下次脊在p4和m1后弯,与下后边脊连,在m2和m3横向,伸达下外脊等。蒙古原松鼠(相似种)与蒙古原松鼠的区别是:颊齿尺寸较小,p4具明显的下外中脊,p4和m1的下次脊弯曲度较缓,m1缺从下外脊伸向下次脊的小刺,m2～3的下后脊II较短等。小原松鼠的主要特征是:个体较小;齿冠较低;下臼齿的下中附尖孤立;下颊齿的下次脊横向、较低,与下外脊连,下中尖和下外中脊均较明显,下次尖明显向前颊侧延伸,具明显的齿带等。蒙古渐新世啮齿类化石的新发现表明,亚洲渐新世的松鼠和山河狸与北美的关系要比与欧洲的近。

NEW OLIGOCENE SCIURIDS AND APLODONTIDS (RODENTIA, MAMMALIA) FROM MONGOLIA

A new species of Kherem, K. asiatica, and three new forms of Prosciurus (P. mongoliensis sp. nov., P. cf. P. mongoliensis and P. pisinnus sp. nov.) from Oligocene of Mongolia are described here. Kherem asiatica, known from Shand Member (late early Oligocene?-?late Oligocene) of Mongolia, is characterized by: cheek teeth larger and wider than those of K. hsandgoliensis; m1～3 with subequally high metaconid and large hypoconid, with more distinct entolophid and entoconid separated from posterolophid by shallow groove. Kherem is referred to the North American subfamily Cedromurinae. It might be derived from an Oligospermophilus-like form. The three genera so far described from Asia (Sciurus, Oligosciurus and Kherem) may represent three different lineages, indicating the Oligocene diversity of the sciurids in Asia. Prosciurus mongoliensis is known from Shand and Tatal Members of the Hsanda Gol Formation, Oligocene, in Mongolia. Its main features are: brachydont cheek teeth have mesostylid connected with metastylid crest, weak mesoconid, and weak or no ectomesolophid; anteroconid and anterior cingulum are absent on p4; an accessory crest extends from ectolophid to hypolophid on m1; metalophid II is well developed on m2～3; hypolophid bends posteriorly to join with posterolophid on p4～m1, but straight to join with ectolophid on m2～3. Prosciurus cf. P. mongoliensis from Shand Member is different from P. mongoliensis in having smaller cheek teeth in size, p4 having distinct ectomesolophid, p4 and m1 having gently curved hypolophid, m1 lacking accessory crest from ectolophid to hypolophid, and m2～3 having shorter metalophid II. Prosciurus pisinnus, collected from Tatal Member, is small in size. Its cheek teeth are brachydont with distinct cingulum. The metalophid II is short, and the hypolophid is transverse, low, and joins with ectolophid. The mesoconid and ectomesolophid are well developed, and the hypoconid extends more anterobuccally. The protoconid of p4 has more developed anterior arm. The new material of the Mongolian Aplodontidae not only has confirmed the early existence of Prosciurus in Asia, but also shows that Prosciurus experienced considerable diversity during the Oligocene in Asia. Both the sciurids and aplodontids found from Oligocene of Asia have closer relationships with those of North America than with Europe.