黄河干流河岸带植物群落特征及其影响因子分析

作为河岸带生态系统的关键组成部分,河岸带植被为许多动植物提供了栖息地以及迁徙或扩散的廊道,并对非点源污染物有着缓冲和过滤作用。黄河是我国湿地的重要组成部分,也是生物多样性分布的关键地带。但是,目前黄河水资源的过度利用如农业灌溉和干流拦河水利工程的兴建在很大程度上改变了原有的水文情势,对河岸带植被发育带来了不利影响。另外,黄河两岸的加固硬化进一步破坏了河岸带植物的生存环境。然而,过去对于黄河河岸带植被仅有对个别河段的调查。为了解目前黄河河岸带植被现状,于2008年4—6月及2008年9—10月对黄河干流河岸带植被进行了两次系统调查,以期为黄河河岸带植被多样性的保护、河岸带的开发管理提供理论支持。对群落的物种组成、多样性进行了分析,并采用双向指示种分析法对黄河干流河岸带植被进行了数量分类。共采集到木本和草本植物169种,隶属于37科124属。区系分析表明黄河干流河岸带植被区系地理成分多样。空间分布方面,中游河段草本植物无论在种类数、密度、生物量上都较上游和下游河段丰富。TWINSPAN将植被划分为17个群落,论述了各群落的特征。环境分析表明,影响黄河干流河岸带植被空间分布的主要生态因子是海拔、年均气温、年均降雨量、年均径流量和平均最大流速。     

基于生物多样性保护的河岸带植被管理对策--以长白山二道白河为例

以生物多样性十分丰富的长白山二道白河上游河岸带为研究对象,在分析河岸带植被管理对生物多样性保护的重要作用的基础上,提出了河岸带及其植被的科学管理对策,为河岸带的生物多样性保护提供理论模式,对河岸带开发利用过程中生态保护的有效实施提供借鉴。     

河岸带生态系统植被与土壤对水文变化的响应研究进展

河岸带的植被与土壤是生态系统重要组成部分,对于维持河岸带的生态健康、生态系统服务与可持续性具有至关重要的作用。水文变化是河岸带生态系统的首要干扰因子,系统总结了水文变化对河岸带植被的特征以及植被形态、群落分布、繁殖、生存策略的影响,并阐述了河岸带水文和植被对土壤氮磷迁移转化的影响机制。根系作为土壤与植物地上部分之间物质、能量流动与信号传导的关键纽带,目前对根系的研究还较欠缺,需要加强水文变化对河岸带湿地植物根系形态、结构、功能特征的影响机理研究,以及湿地植物对水文变化的适应机制和耐受阈值方面的探究。在微观方面,应加强水文变化与植被等多因素耦合对土壤氮磷迁移转化过程的机理研究。河流形态和土壤的多样性决定着河岸带水文作用特征的复杂性,今后需注重河岸带个性特征与水文响应的关系研究。河岸带是横向的水陆生态过渡带和河流上下游的纵向生态廊道,亟需综合考虑和模拟流域土壤、植被与水文、人类活动之间的耦合关系,预测未来气候与社会经济情境下的河岸带生态系统演变规律,为河岸带生态系统的生态调节、生物多样性保护与生态恢复等提供理论依据与技术支撑。     

河岸带对陆地水体氮素输入的截流转化作用

氮素是陆地水体的重要污染物之一,相邻高地土壤中的氮素经非点源途径通过河岸带进入水体.河岸带是高地和水体之间的过渡带,健康的河岸带系统能够通过物理的、生物的和生物化学的过程,实现对氮素的截留转化.本文系统地介绍了氮素进入陆地水体的主要途径和河岸带对陆地水体氮素污染物截留转化的机制,并从河岸带水文过程、土壤特征、植被状况、人为活动等方面,论述了影响河岸带氮素截留转化作用的因素;在此基础上,对目前河岸带研究中存在的问题进行了讨论,并对我国开展河岸带的研究提出了建议.     

Changes in vegetation and soil properties following 6 years of enclosure in riparian corridors

河岸带是维持生物多样性的重要生态系统之一。然而,由于过度放牧引起的植被消耗和过度开垦等人类活动的干扰,河岸带植被多样性和植被盖度受到严重的破坏,甚至威胁了河道的稳定性。围栏封育在退化草地生态系统修复中被广泛应用,但对退化河岸带植被群落和土壤性质的影响尚不明确。本研究的目的旨在明确围封的实施是否会促进河岸带植被群落的物种组成、物种丰富度和物种多样性恢复,土壤氧分如何随围封年限的增加而变化。辽河干流自2012年起被围栏封育管理,本研究在辽河干流河岸带沿岸设置了20个草本群落长期观测样地,记录了2012–2017年样地中植被高度、盖度和个体数量等参数用于物种丰富度和物种多样性的统计分析。同时,分别测定了2012年和2017年植被群落土壤氧分含量,验证了植被群落和土壤氧分对围封的反馈,研究了2012–2017年辽河干流河岸带的围栏封育对物种多样性和土壤氧分的影响。结果表明,随着围封年限的增加,辽河干流河岸带草本群落植被丰富度和多样性显著增加。物种组成方面,菊科植物的优势度显著增加,禾本科植物优势度显著下降。围封后植被群落的恢复和禁止耕作,加速了土壤中磷和钾的消耗,表现为显著降低,土壤有机质含量对围封的响应表现的相对滞后,并没有显著变化。综上所述,本研究为河岸带植被群落物种多样性、物种组成对围封的响应提供了新的见解。     

塔里木河下游河岸带植被的空间结构特征

揭示我国内陆河流域下游河岸带植被的空间结构特征,对于了解我国西北干旱区荒漠河岸带植被的空间分布规律、指导荒漠化治理和内陆河水资源管理具有重要意义。该研究基于野外大范围植被调查数据支持下的遥感监督分类方法,利用Landsat-8 OLI遥感数字图像,辨识了塔里木河下游柽柳(Tamarix spp.)灌丛、胡杨(Populus euphratica)疏林和芦苇(Phragmites australis)草地3类主要的河岸带植被,并利用建立的叶面积指数(LAI)遥感反演经验模型反演了研究区柽柳灌丛和胡杨疏林的叶面积指数,旨在从区域尺度和总体趋势上分析荒漠河岸带植被的空间结构和分布特征。结果表明:在有详细地物资料的基础上,遥感监督分类可以作为一种干旱区荒漠河岸带植被分类的有效方法;遥感分类结果显示塔里木河下游胡杨疏林分布面积约336.4 km2,柽柳灌丛约为405.3 km2,胡杨疏林总体更靠近河道,柽柳灌丛分布范围更广;河岸带植被LAI整体很低,柽柳灌丛和胡杨疏林平均LAI值分别为0.253和0.252,LAI小于0.5的植被对应面积分别占柽柳灌丛和胡杨疏林总面积的92.4%和90.1%,表明了塔里木河下游荒漠河岸植被空间上稀疏分布的特征;统计结果显示,河岸带植被结构存在巨大的空间变异性,其中胡杨疏林比柽柳灌丛的空间变异性更大;河岸带植被LAI随距河道距离呈现显著负指数分布规律,在离河道1 km范围内LAI随离河道距离快速下降,而1 km外区域叶面积指数普遍低于0.1,表明植被主要分布在河道两侧1 km范围内。整体稀疏的空间分布、显著的空间变异性,以及由LAI体现的植被盖度随距河道距离的负指数下降规律是荒漠河岸带植被空间结构的3个基本特征。     

河岸带生态系统退化机制及其恢复研究进展

恢复和重建自然和人为干扰导致的退化河岸带生态系统是目前恢复生态学、流域生态学等学科研究的重要内容之一．对河岸带生态系统的干扰表现在河流水文特征改变、河岸带直接干扰和流域尺度干扰3个方面,分别具有不同的影响机制．河流水文特征改变通过改变河岸土壤湿度、氧化还原电位、生物生存环境以及沉积物传输规律对河岸带生态系统产生影响;河岸带直接干扰通过人类活动及外来物种入侵而直接影响河岸带植被多样性;流域尺度干扰则主要表现在河道刷深、河道淤积、河岸带地下水位降低和河流冲刷过程改变等．河岸带生态恢复评价对象包括河岸带生态系统各要素,评价指标已从单一的生态指标转向综合性指标．河岸带生态恢复应在景观或者流域尺度上进行考虑,识别对其影响的生物和物理过程以及导致其退化的干扰因子,通过植被重建与水文调控来进行．扩展研究尺度和研究对象及采用多学科的研究方法将是今后相关研究中的重要问题．     

广西桂林漓江河岸带植被配置类型与退化机制研究

河岸带植被结构与组成等配置状况直接影响其稳定河岸、净化水质、生物多样性、生物栖息地以及旅游景观等重要功能。查明漓江河岸带主要植被配置类型及其各自退化表现、原因与恢复途径,是开展河岸带退化植被恢复的基础与前提。该研究通过野外实地调查与测量,对漓江桂林-阳朔河段51个河岸带断面的地形、土壤、植被等进行了系统调查,分析了河岸带植被退化的主要表现、主导干扰因素,并通过流域与河岸带尺度综合的角度,提出了漓江河岸带植被恢复的思路。从岸坡地形与植被组合出发,按照自然、近自然、人工等干扰程度,广西漓江河岸带可划分为江心洲宽滩地疏林灌草自然型、丘陵陡坡林灌草自然型、平原农田缓坡灌草近自然型、平原滩地林草近自然型、聚落阶地疏灌林草人工-自然混合型、城镇岸坡林草人工型等6种典型植被配置模式。漓江河岸带植被退化主要表现为多样性差、景观破碎、生态功能弱,而城镇、交通、旅游等不合理开发建设是主导性的干扰因素。从生态稳定和景观美观的角度,提出了漓江河岸带植被恢复与优化的思路,以期为广西漓江河岸带生态修复提供科学依据。     

河岸缓冲带对氮污染物削减作用研究进展

氮(N)是水体的重要污染物之一。河岸缓冲带作为连接陆域和水域的生态纽带对N素去除效果显著,是有效控制N素向水体输出的重要途径。本文介绍了河岸带N素的迁移过程,从物理、化学及生物角度阐述了河岸带对N素的去除机理,归纳总结了河岸带对N素的去除效率、影响因素和研究手段。指出不同N素去除途径下N素去除效率的主要影响因素不同,植被结构、土壤渗透和持水特征、河岸宽度和坡度是通过影响物理作用去除N素的主导因素,植被类型、季节动态、植物生长发育是通过影响植物吸收去除N素的主导因素,土壤温度、有机碳含量、pH、硝态氮含量、氧含量是通过影响反硝化作用去除N素的主导因素。最后分析了当前研究中存在的问题,并提出了研究展望及今后研究重点。     

浙江灵山港滩区植被分布数量波动特征及数学描述

河岸带滩区植被是河岸带功能有效发挥的关键因素。以浙江省龙游县灵山港河岸带滩区为研究对象,运用双向指示种分析法和典范对应分析法,分析了2015—2019年滩区植被空间分布与数量波动特征,并构建了河岸带滩区植被波动测度公式。结果表明:2015—2019年灵山港河岸带滩区植物共有189种,隶属于53科158属,植物群落包括7种类型;相对水面高差是植物群落空间分布的主导因子,随相对水面高差增大,植物群落由湿生群落向中生群落变化,群落结构由单一草本结构向灌草和乔灌草复层结构变化;滩区优势植物种群波动明显,主要有上升波动型、下降波动型和复合波动型3种类型;植物波动具有明显的方向性,滩区植被波动率维持在-0.250～0.219,其中,2015年、2018年和2019年为正向波动,2016年、2017年为负向波动;构建了滩区植被波动率计算公式,为河道生态修复提供了一定参考。     

Impact of riparian buffer zones on water quality and associated management considerations

Recent attention has focused on riparian forest buffer systems for filtering sediment, nutrients, and pesticides entering from upland agricultural fields. This paper summarizes the results of a field monitoring study done in Tokachikawa watershed in Hokkaido, Japan, Cisadane, Cianten and Citamyang sub-watersheds in Indonesia and Cauvery watershed, India to quantify the impact of riparian buffer zones on changes in stream water quality. A watershed approach was used to compare land use indicators – uplands, forests, riparian forest, livestock areas – to a wide range of surface water physical and chemical properties. Stream water physical property values increased from upstream to the confluence point, influenced by the upland and livestock land use activities. The greatest reduction in impairment of water quality was observed in buffer zones located along higher order streams where the gradient is very low, leading to slow groundwater movement. The lower stream water temperature in riparian buffer zones suggests that the shading effect is most pronounced in this area of the watershed. The results demonstrate the positive impact of forest buffer zones in reducing the influence of agricultural nutrients and chemicals on surface stream waters. Design and management considerations for establishing riparian zone land use are discussed.     

Neighbourhood-scale urban riparian ecosystem classification

Urban riparian ecosystems are now recognized as essential components in determining the vulnerability of nature and human systems to climate change, but still remains uncertainty concerning how to stratify and classify urban riparian landscapes into units of ecological significance at spatial scales appropriate for management. Ecosystem classification presents a tool that allows for quantifying the social and ecological processes of ecosystems so as to shape them into relatively homogeneous management objectives, and provides the potential of offering decision support to urban planners based on urban ecological principles. The purpose of this study is to explore and develop a framework for urban riparian ecosystem classification. The classification framework integrates 9 ecosystem components and 29 spatially-explicit variables that characterized by the biophysical landscape, built environment and human population. Then this framework is applied at the neighbourhood scale in Huangpu River basin, Shanghai city, China, employing hierarchical cluster analysis. The results of the ecosystem classification show that the riparian ecosystems in the Huangpu River could be grouped into three principal categories and these ecosystems are aligned along a gradient of increasing urbanization. We highlight the conservation and payment for riparian ecosystem services delivered in the upstream of Huangpu River, and revegetation and enhancement of riparian ecosystems in the more urbanized downstream area. We also find that riparian vegetation connectivity and coverage were negatively related to land use mix, built environment density, and economic wealth. In terms of riparian vegetation ecology, neighbourhood scale land use mix and built environment density are more influential than socioeconomic background (such as average family income and immigrant status).     

Butterflies as an indicator group of riparian ecosystem assessment

Riparian ecosystems play an important role in modulating a range of ecosystem processes that affect aquatic and terrestrial organisms. Butterflies are a major herbivore in terrestrial ecosystems and are also common in riparian ecosystems. Since butterflies use plants for larval food and adult nectar sources in riparian ecosystems, butterfly diversity can be utilized to evaluate riparian ecosystems. We compiled butterfly data from 33 sites in three riparian ecosystem types across the country and compared butterfly diversity in terms of number of species and quality index in relation to riparian environmental variables. Number of butterfly and plant species was not different among three riparian habitat types. Additionally, there was no significant ecological variable to distinguish the butterfly communities on three riparian habitats. Non-metric multi-dimensional scaling ordination showed that butterfly communities in three riparian ecosystem types differed from each other, and butterfly riparian quality index was the main variable for butterfly assemblages. Five indicator species for moor and another five species for riverine riparian ecosystems were identified. Three and one indicator species for moor and riparian ecosystems, respectively, were plant specialists, while 44 butterflies were general feeders, feeding on a wide range of hostplants in several habitats. These results suggest that butterfly species use actively riparian habitats for nectar and larval food, and the butterfly riparian quality index can be employed to track faunal change in riparian habitats, which are frequently threatened by disturbances such as water level and climate changes, and invasive species.     

东风港滨岸缓冲带对水生生物群落结构的影响

选择苏州河的支流东风港为研究对象,选取其中一段人工建造滨岸缓冲带作为研究区,调查其水生生物群落并与对照区相比较,以具体说明滨岸缓冲带对水生生物群落结构的影响.通过对研究区与对照区内的浮游植物、浮游动物、底栖动物和大型水生植物的调查研究,结果显示,研究区浮游植物的种类数多于对照区,且以清水种占优势;浮游动物群落结构研究区明显比对照区复杂;底栖动物的种类数、密度和生物量研究区都多于对照区;另外研究区还有较为多样化的大型水生植物群落.这些都表明研究区的水生生物群落结构较对照区更为完善.说明滨岸缓冲带对提高水体生物多样性,完善水生生物群落结构具有较为明显的作用.     

Some approaches for measuring and modelling riparian shade

The radiation environment of streams is of major ecological importance because it controls stream thermal regime and light availability for photosynthesis. Therefore, methods are needed for measuring stream shade in practical riparian management. The quantity ‘diffuse non-interceptance’ (difn), defined as the proportion of incident lighting received under a sky of uniform brightness and best estimated from fish-eye images, is useful for general specification of light exposure. For routine measurement of difn along stream reaches we recommend using a matched pair of simple light sensors (e.g. photosynthetically available radiation sensors) under conditions of complete overcast (which has almost uniform brightness). Methods are also needed for predicting future light exposure as riparian plantings grow and increasingly shade the stream. A simple model is outlined for predicting difn at the channel centre as a function of channel dimensions (stream width, w) and riparian plant character (foliage density, canopy height, h). The model reproduces the broad empirical trend of increasing shade with increasing h/w ratio. Future model refinement will aim to quantify the increase in shade moving from channel centre to edge under an overhanging canopy.     

The effect of anthropogenic disturbance on the hydrochemical characteristics of riparian wetlands at the Middle Ebro River (NE Spain)

In natural systems, the chemistry of floodplain waters is a function of the source of the water, which is influenced by geomorphic features of riparian wetlands. However, anthropogenic disturbances may alter both geomorphic features and the natural balance of water mixing in the floodplain. The aim of this study was to classify riparian wetlands and characterize their water characteristics in one reach of the Middle Ebro River to assess the hydrochemical functioning of the system. In order to accomplish that goal, water samples were collected at 40 sampling sites during low-water conditions and two floods of different magnitude. Moreover, geomorphic characteristics of riparian wetlands were also analyzed to interpret the results at broader spatio-temporal scales. Three group of wetlands were identified using multivariate ordination: (1) major and secondary channels highly connected to the river by surface water, containing weakly ionized water with high nitrate levels during floods; (2) secondary channels and artificial ponds located in riparian forests near the river, most of which were affected by river seepage during the examined events. This type of sites had high major ions concentrations and elevated spatial variability with respect to nutrient concentrations during floods; (3) Siltated oxbow lakes, whose hydrogeochemical features seemed to be unaffected by factors related to river fluctuations. Total dissolved solids, major ion (sulfate, chloride, sodium, calcium, magnesium, and potassium) and nutrient (nitrate, ammonium and organic nitrogen, and phosphate) depended upon the relationships between surface and subsurface water flows. Seasonal changes and geomorphic characterization indicated that a strong functional dependence of floodplain wetlands close to the main river channel is established, whereas most of the floodplain area remains disconnected from river dynamics. Moreover, the effect of nitrate-enriched agricultural runoff seems to affect water quality and hydrochemical gradients of the system. Based on our results, we propose different types of actions for the management of the Ebro River flow to ensure a more natural ecological functioning of its floodplains. Handling editor: P. Viaroli     

神农架地区珍稀植物沿河岸带的分布格局及其保护意义

在神农架香溪河流域从源头到河口的不同海拔高度沿河岸带共设置 4 0个与河流方向平行的10 0m× 10m的样带 ,进行了植物群落学调查研究 .结果表明 ,沿河岸带分布有珍稀植物 14种 ,占神农架地区珍稀植物总数的 4 2 .4 % .这些珍稀物种主要分布在海拔 12 0 0～ 180 0m山地常绿阔叶落叶阔叶混交林带 ,珍稀植物种类的物种丰富度在中等海拔高度上最大 .珍稀植物的种类可划分为低海拔、中等海拔和高海拔 3组 .针对珍稀植物的分布特点 ,明确指出应重视河岸带在生物多样性、尤其是珍稀物种保护方面的重要作用 .     

Effects of pasture development on the ecological functions of riparian forests in Hokkaido in northern Japan

In the summer, the forest canopy lowers the water temperature, which is very important for anadromous fish, and its population density is significantly lower in grassland streams. Leaf litter and terrestrial invertebrates are the critical food resources for stream organisms. In a basin where the riparian forest is preserved, but other areas have been cut, the amount of leaf litter is almost equivalent to that in an intact natural basin. The annual input of terrestrial invertebrates falling into the forested reaches was 1.7 times greater than that in the grassland reaches, and fish biomass was significantly less in the grassland reaches. In-stream large woody debris creates storage sites for organic and inorganic matter and enhances habitat diversity for aquatic biota. However, the volume and number of large wood pieces decreased significantly with pasture development, because it clears the riparian forests and covers the riverbanks with grass. Fine sediment is a prominent by-product of agricultural development and adversely impacts periphyton productivity, the density and diversity of aquatic invertebrates, fish feeding, fish spawning and egg survival. We also examine the adequate width of a riparian buffer if it is to be able to satisfy its ecological functions.     

Predictions of stream nutrient and sediment yield changes following restoration of forested riparian buffers

Riparian tree planting is widely recognised as a means to improve water quality and stream habitat. However, shading of riparian pasture grasses can lead to channel widening, and riparian shade may limit the growth of macrophytes and algae that assimilate dissolved nutrients from the water column. We investigated concerns that riparian management could lead to increased yields of nutrients and sediments through a conceptual modelling exercise. A simple model of the trade-off between interception of nutrients in runoff by forest buffers versus reduction of in-stream uptake due to shade, predicted that a buffer strip alongside a small headwater stream would reduce nutrient export, while a buffer strip instigated as an isolated patch alongside a larger stream (c. >2.5 km² upstream catchment size) would increase nutrient export, as the relative amount of nutrients trapped by the buffer decreases as the nutrient load present in the stream water increases. However, in these larger streams with width exceeding approximately 6 m, sufficient light may reach the streambed for plant and algal growth, which in turn would promote instream nutrient processing. At the peak of streambank erosion after planting, predicted total sediment yield (hillslope plus bank sources) was appreciably higher than the hillslope pasture yield, but sediment yield stabilised c. 35–40 years after planting. When planting was extended over 40 years in the model, the sediment yield never exceeded that in pasture before planting. This conceptual modelling exercise shows that riparian tree planting programmes should commence in the headwaters and progress downstream to avoid nutrient yield increases. Significant sediment yield from bank stored sediment of small streams can be expected until the channel reaches the more stable, original forested width, but progressive planting may decrease the peak loads of sediment.     

Nutrient mobilization and processing in Sonoran desert riparian soils following artificial re-wetting

Research in river-floodplain systems has emphasized the importance of nutrient delivery by floodwaters, but the mechanisms by which floods make nutrients available are rarely evaluated. Using a laboratory re-wetting experiment, we evaluated the alternative hypotheses that increased nutrient concentrations in riparian groundwater during flash floods are due to (H1) elevated nutrient concentrations in surface floodwaters entering the riparian zone or (H2) re-mobilization of nutrients from riparian soils. We sampled soils from the riparian zone of a 400m reach of Sycamore Creek, AZ. Two sub-samples from each soil were re-wetted with a solution that mimicked the chemistry of floodwaters, with one sub-sample simultaneously treated with a biocide. We also measured structural characteristics of soils (texture, organic matter, moisture, and extractable nutrients) to investigate relationships between these characteristics and response to re-wetting. Riparian soils exhibited considerable variation in physical and chemical structure. Soil organic matter, moisture, and texture co-varied among samples. Re-wetting increased concentrations of nitrate and ammonium, and decreased SRP, relative to initial concentrations. Live soils were significantly lower in NO3-and SRP than biocide-treated samples. Extractable DIN pools were the best predictors of mobilization, and soil organic matter was strongly correlated with nitrate losses, probably via its relationship with microbial uptake. Nutrient mobilization and processing also varied considerably with depth, lateral position, and among plots. We estimate that 70–80% of N in riparian groundwater during flash floods is re-mobilized from riparian soils, but are unable to reject the hypothesis that flood inputs may be important sources of nutrients to riparian soils over longer time scales.