New Middle Permian insects from Salagou Formation of the Lodève Basin in southern France (Insecta: Pterygota)

Summary

New insects are described from the Middle Permian (Guadalupian) of Salagou Formation in the Lodève Basin (Languedoc region, France). Palaeopterans are represented by new odonatopterans viz. Huangiopterum lodevense n. gen., n. sp. and Huangiopterum species B, attributed to clade Panodonata (Huangiopteridae n. fam.), already recorded through the family Saxonagrionidae in the same formation, and a diaphanopterodean Martynovia cf. halli Beckemeyer 2004 (Martynoviidae) known from the Lower Permian of Oklahoma. Archaeorthopterans are supplemented by Bethouxia ariegensis n. gen., n. sp., Paralongzhua elongata n. gen., n. sp., Lodevolongzhua incompleta n. gen., n. sp., the caloneurodean Jarmiloptera mouralensis n. gen., n. sp., and a cnemidolestodean cf. Tococladus sp. (Tococladidae). Permophyllum rotundatum n. sp. (?Tettigoniidae) demonstrates the first presence of Orthoptera with wings probably mimicking Peltaspermales reproductive organs in Late Palaeozoic possibly analogous to the known crypsis of roachoid tegmina with pteridosperm pinnules. Frequent occurrence of hardly identifiable grylloblattid wing fragments support their significant role in term of abundance and diversity of the Permian faunas. New taxa based on wing venation pattern as Genera and species undetermined A, B (Liomopteridae), Permobaharellus salagousensis n. gen., n. sp. (family uncertain), Lodevophlebia reticulata n. gen., n. sp. (Sylvaphlebiidae), Oborella lodevensis n. sp. (Euryptilonidae), and Genus and species undetermined C, were described. Paoliida is represented by a paoliid gen. et sp. indet. and three anthracoptilids Strephocladus gandi n. sp., Strephocladus mouralensis n. sp., Lodevocladus subtilis n. gen., n. sp. The first record of miomopteran Palaeomantis sp. (Paleomanteidae) from Salagou Formation is presented. Acercarian insects are represented by the first record of Thripida and diverse new hemipterans, namely Euroscytina lutevanorum n. gen., n. sp. (Cicadomorpha: Scytinopteridae), Prosbole dio n. sp., Prosbole garrici n. sp., Pervestigia cf. veteris Becker-Migdisova 1961 (Prosbolidae), Archescytina species A (Paleorrhyncha: Archescytinidae) and Permopsyllidium lesclansis n. sp. (Sternorrhyncha: Psylloidea: Protopsyllidiidae). Entomofauna from the Salagou Formation demonstrate highly diverse insect communities with some taxa linked to early and late Permian ones known from assemblages in Euramerica and Russia, such as Obora (Czech Republic), Wellington Formation (Kansas and Oklahoma, USA), and Tshekarda (Ural, Russia).     

Latest occurrences of the Mesozoic family Elcanidae (Insecta: Orthoptera), in Cretaceous amber from Myanmar and Spain

Mesozoic orthopterans of the family Elcanidae are reported (as nymphs) in amber, from the latest Albian-Cenomanian of northern Myanmar and the Albian of northern Spain. Four distinct new species in two new genera occur, Burmelcana longirostris n. gen, n. sp. in amber from Myanmar and Hispanelcana arilloi n. gen, n. sp., H. alavensis n. sp. and H. lopezvallei n. sp. from Spanish amber. Detailed preservation reveals the fine structure of the tibial spurs and spines that are so distinctive to Elcanidae, as well as details of the abdominal styli, cerci, tarsomeres, and mouthparts. Elcanidae and their stem group, Permelcanidae, are known from the Early Permian to the Early Cretaceous (Aptian), so the amber fossils represent the latest known occurrence of this clade.     

New genera and new species of leaf beetles (Coleoptera: Polyphaga: Chrysomelidae) from Lowermost Eocene French amber

Three new monotypical genera of leaf beetles are described: Aoriopsis eocenicus n. gen, n. sp. and Acolaspoides longipes n. gen., n. sp. from the subfamily Eumolpinae, and also Crepidocnema yantarica n. gen., n. sp. from the subfamily Alticinae. Short reviews of known fossil records of the mentioned subfamilies are given and position of the new genera discussed.     

Anomala trapezifera species-group: a burst of diversity (Coleoptera: Scarabaeidae: Rutelinae)

The species Anomala aereiventris n. sp., A. aspersa n. sp., A. atrivillosa n. sp., A. clarivillosa n. sp., A. contusa n. sp., A. eusticta n. sp., A. hiata n. sp., A. latifalculata n. sp., A. leopardina n. sp., A. levicollis n. sp., A. longisacculata n. sp., A. m-fuscum n. sp., A. perspicax n. sp., A. piccolina n. sp., A. globulata n. sp., A. stillaticia n. sp., A. subridens n. sp., A. subusta n. sp., A. tenoriensis n. sp., A. tuberculata n. sp. and A. vallisneria n. sp. from Costa Rica are described. These species and A. polygona Bates 1888, A. trapezifera Bates 1888 and A. vulcanicola Ohaus 1897 are placed in a new species-group, named after A. trapezifera, whose diagnosis is provided. Their distribution patterns are discussed.     

Family Lasiosynidae n. fam., new palaeoendemic Mesozoic family from the infraorder Elateriformia (Coleoptera: Polyphaga)

Lasiosynidae n. fam. is proposed for the genera Lasiosyne Tan, Ren & Chih 2007 (transferred from Archostemata to Polyphaga), Anacapitis Yan 2009 Tarsomegamerus Zhang 2005 (proposed in the superfamily Chrysomeloidea) and Bupredactyla n. gen. The new family is regarded in composition of the infraorder Elateriformia without a more detailed attribution, because it demonstrates a mixture of characters of different families and superfamilies, i.e. somehow an intermediate position between the superfamilies Dascilloidea, Elateroidea, Buprestoidea and Byrrhoidea with most resemblance to Dascillidae, Schizopodidae, Eulichadidae, Ptilodactylidae and Callirrhipidae and probable more close relationship to the last three families. Four new fossil species of the genus Lasiosyne: L. daohugouensis n. sp., L. fedorenkoi n. sp., L. gratiosa n. sp., L. quadricollis n. sp., and also Bupredactyla magna n. sp. are described from the Middle Jurassic Jiulongshan Formation of eastern Inner Mongolia, China. A probable generic composition of the new family is considered. The synonymy of generic names Anacapitis Yan 2009 and Brachysyne Tan & Ren 2009, n. syn. as well as synonymy of species names Lasiosyne euglyphea Tan, Ren & Chih 2007, Pappisyne eucallus Tan & Ren 2009, n. syn. and Pappisyne spathulata Tan & Ren 2009, n. syn. are proposed.     

New and little known species of the genus Zorochros Thomson 1859 (Coleoptera: Elateridae) from Palaearctic and Oriental Region

Sixteen new species of the genus Zorochros Thomson 1859 from China, Indonesia, Malaysia, Nepal and Thailand are described and illustrated, a key to the species of the Zorochros indicus group is given, and new records on already known species are provided. New species: Z. ahrensi n. sp., Z bingkorensis n. sp., Z. dabieshanensis n. sp., Z. dolini n. sp., Z. hartmanni n. sp., Z. hubeiensis n. sp., Z. karnaliensis n. sp., Z. magnificus n. sp., Z. naniensis n. sp., Z. nigredos n. sp., Z. platiai n. sp., Z schawalleri n. sp., Z. schmidti n. sp., Z. senaroensis n. sp., Z. theodori n. sp., and Z. tongshanensis n. sp.     

A revision of the genus Paronthobium from New Caledonia (Coleoptera: Scarabaeidae: Scarabaeinae: Epilissini)

A revision of the New Caledonian genus Paronthobium Paulian 1984 is presented. Anonthobium Paulian 1984 is synonymized with Paronthobium Paulian 1984 after evaluating the diagnostic characters originally introduced by Paulian to separate them. Upon examination of the type specimens, Onthobium caledonicum Paulian 1935 is separated from O. simplex Fauvel 1903 and restored as valid. Anonthobium moui Paulian 1984 is synonymized with Anonthobium micros Paulian 1984. Fifteen new species are described: P. adio n. sp., P. daghfousi n. sp., P. dierkensi n. sp., P. farino n. sp., P. iac n. sp., P. julieni n. sp., P. memaoya n. sp., P. minutum n. sp., P. nasutum n. sp., P. orientale n. sp., P. peckorum n. sp., P. petchecara n. sp., P. subdentatum n. sp., P. taom n. sp. and P. tchingou n. sp. The genus Paronthobium is now composed of 22 species. Illustrations of parameres and of male protibiae are provided for each species. Distribution maps and a key to species are given.     

Review of subtribe Pterellina Rohdendorf, 1967 (Diptera: Sarcophagidae) of Middle East with descriptions of twelve new species

Summary

Based on a revision of large recent collections housed at the Natural History Museum (London), Tel Aviv University and in the private collection of Yu. Verves and L. Khrokalo (Kyiv, Ukraine), new distributional data and an updated list with comments of 28 species of Middle Eastern Sarcophagidae of the subtribe Pterellina are presented. Seventeen of them are recorded for Israel, 11 for Egypt, four for Iran, two each for the Palestinian Authority, Saudi Arabia and Turkey, and one each for Jordan and the UAE. No data are known for the territories of Bahrain, Cyprus, Iraq, Kuwait, Lebanon, Oman, Qatar, Syria and Yemen. Twelve species new to science are described, including nine from Israel (Chaetapodacra montana n. sp., Eremasiomyia bicolor n. sp., E. deserta n. sp., E. fuscata n. sp., Protomiltogramma bidentata n. sp., P. serrata n. sp., Pterella hermonica n. sp., P. indigens n. sp., P. zefatica n. sp.), two from Egypt (Chaetapodacra griseola n. sp., Eremasiomyia alba n. sp.) and one from Saudi Arabia (Protomiltogramma arabica n. sp.). The male of Eremasiomyia nigra Rohdendorf, 1935 is described for the first time. One new combination of specific name is proposed ? Protomiltogramma immunita (Villeneuve, 1923), n. comb. Original keys for the identification of genera and species are proposed. An annotated list of all world genera and species and compiled data on larval habits of Pterellina are included.     

Descriptions of new Neotropical Lycoperdininae (Coleoptera: Endomychidae) and their phylogenetic placement

Two new genera and three new species of Lycoperdininae (Coleoptera: Endomychidae) from South America, are described and illustrated based on adults: Hylaperdina n. g., along with H. brevicornis n. sp. from Ecuador and H. costariciensis n. sp. from Costa Rica, and Chileanus n. g., along with C. talca n. sp. from Chile. Their most likely placement within the Lycoperdininae is discussed. A key to genera of the neotropical Lycoperdininae is provided.     

Les Cryptotrogus Kraatz de la faune d’Iran et des régions limitrophes (Coleoptera : Scarabaeoidea : Melolonthidae)

Le genre Cryptotrogus Kraatz, 1888 est étudié. Après étude des spécimens-types des espèces associées et de nouveaux matériels, ce genre est redéfini et Cyphonoxia Reitter, 1889 en est à nouveau considéré comme synonyme. Les nouvelles combinaisons et synonymies suivantes sont établies ou restaurées : Cryptotrogus buettikeri (Sabatinelli & Pontuale, 1998) n. comb., Cryptotrogus delhiensis (Anand, 1988) n. comb., Cryptotrogus haarlovi (Petrovitz, 1955) n. comb., Cryptotrogus kermanensis (Semenov & Medvedev, 1936) n. comb., Cryptotrogus kircheri (Balthasar, 1930) n. comb., Cryptotrogus praestabilis (Reitter, 1889) n. comb., Cryptotrogus tatianae (Semenov & Medvedev, 1936) n. comb., Cryptotrogus zarudnyi (Semenov & Medvedev, 1936) n. comb., Cyphonotus bicoloratus Petrovitz, 1962 n. syn. de Cryptotrogus mesopotamicus (Petrovitz, 1962), Cryptotrogus miksici Petrovitz, 1965, et Cryptotrogus orita (Reitter, 1902) n. syn. de Cryptotrogus niveus (Hampe, 1852), Cyphonoxia glasunowi Semenov, 1897 n. syn. de Cryptotrogus brenskei (Reitter, 1895) comb. rev., Cyphonoxia kermanensis Semenov & Medvedev, 1936 n. syn. de Cryptotrogus zarudnyi (Semenov & Medvedev, 1936) n. comb., Cryptotrogus zemindar (Sharp, 1876) n. comb. Par ailleurs, huit espèces sont décrites et comparées aux espèces proches : sept d’Iran, Cryptotrogus borumandi n. sp., Cryptotrogus ebrahimii n. sp., Cryptotrogus miesseni n. sp., Cryptotrogus mirzayansi n. sp., Cryptotrogus monodi n. sp., Cryptotrogus morgani n. sp. et Cryptotrogus parallelus n. sp., et une, Cryptotrogus irakanus n. sp., d’Irak. Enfin, de nouvelles données sont présentées pour quelques espèces de la faune iranienne appartenant aux genres Melolontha Fabricius, 1775, Anoxia Castelnau, 1832, Polyphylla Harris, 1841 et Microphylla Kraatz, 1890. Une espèce est nouvelle pour la faune d’Iran : Melolontha aceris Faldermann, 1835.     

Ecnomus of Madagascar,species without modified spur (Trichoptera: Ecnomidae)

The 11 Malagasy species of the genus Ecnomus McLachlan, 1864 that do not belong to the psychomyiellodes species group are studied. One of them (Ecnomus salgiel Malicky, 2015) was recently described from Nosy Be Island. Ten are new and described: Ecnomus felix n. sp., E. dobignyi n. sp., E. matsaboryi n. sp., E. granjoni n. sp., E. commersoni n. sp., E. menavavae n. sp., E. ikopaensis n. sp., E. rubermortis n. sp., E. evidens n. sp., and E. tendroensis n. sp. All are endemic to Madagascar. Their distributions on the island are briefly described according to ecological data.     

Descriptions of the first semi-aphenopsian troglobiotic Patrobini and of a new anophthalmic cave-dwelling Trechini from central Sichuan,China (Coleoptera: Caraboidea)

Summary

The first known troglobiotic species belonging to the ground beetle tribe Patrobini is reported from a limestone cave called Lianhua Dong in Pengzhou, a suburb county-level city in northern Chengdu, Sichuan Province, China. Troglopatrobus zhouchaoi n. gen., n. sp., is an eyeless and large-sized patrobine beetle with extremely elongated body (esp. head and elytra) and appendages. Duvalioblemus (Shublemus) liyuani n. subgen., n. sp., a minute anophthalmic trechine species, is also discovered from the same cave. It is the first cave-dwelling species belonging to this genus already known as endogean from Sichuan and Yunnan provinces.     

Austrini - a new tribe of Tropiduchidae planthoppers from the Eocene Baltic amber (Hemiptera: Fulgoromorpha)

A new tribe Austrini n. trib. of Tropiduchidae planthoppers from the Eocene Baltic amber is described to comprise Austris raffelis n. gen., n. sp. Fossils ascribed to family Tropiduchidae are discussed.     

Six new species of the subgenera Alloparadisa and Ashima (Diptera: Drosophilidae: Phortica) from Yunnan,China, with DNA barcoding information

Six new species of the genus Phortica are described from Yunnan, Southwest China: P. (Alloparadisa) kaibangya n. sp., P. (Ashima) semiannulata n. sp., P. (Ashima) dianmianensis n. sp., P. (Ashima) idiasta n. sp., P. (Ashima) menglian n. sp. and P. (Ashima) zhangyuanae n. sp. The 22 new DNA sequences of the mitochondrial COI gene with BOLD Process ID and GenBank accession numbers are provided for the six new species and six related known species from southern China: P. (Ashima) glabra, Chen & Toda, 2005 P. (Ashima) nudiarista Cheng & Chen, 2008, P. (Ashima) saltiaristula Chen & Wen, 2005, P. (Ashima) spinosa, Chen & Toda, 2005 P. (Ashima) symmetria Chen & Toda, 2005 and P. (Ashima) yiqini Zhu & Chen, 2017. The pairwise interspecific K-2P COI distances among all Chinese species of the subgenera Alloparadisa and Ashima are summarized. An identification key to males of all Chinese species of the subgenera Alloparadisa and Ashima is provided.     

New Tridactyloidea in Miocene amber from the Dominican Republic (Orthoptera: Caelifera)

Two new Tridactyloidea of the families Ripipterygidae and Tridactylidae are described from the Early Miocene (Burdigalian) amber of the Dominican Republic, namely: Archaeoellipes engeli n. gen., n. sp. (Tridactylidae: Tridactylinae) and Mirhipipteryx antillarum n. sp. (Ripipterygidae). These new taxa represent the fi rst record of Tridactyloidea in Dominican amber and the fi rst fossil record of Ripipterygidae, which are otherwise known only from the extant fauna.     

New Hypogastrura species and new distributional data for Hypogastrura distincta (Axelson 1902) from China (Collembola: Poduromorpha: Hypogastruridae)

Two new species, Hypogastrura analpapillata n. sp., on mushrooms from Yunnan Province, and Hypogastrura microspinata n. sp., on moss from Hunan Province, are described. Hypogastrura analpapillata n. sp. is the sixth species with highly modified mouthparts in the trybomi group. Hypogastrura microspinata n. sp. is close to H. yosii Stach 1964, but can be differentiated from the latter by body granules, tenent hair and lateral teeth on tibiotarsus. A key to Chinese species of Hypogastrura is provided and a list of Chinese Hypogastrura species and their distributional data are given.