Les foraminifères du Paléocène et de l'Éocène basal du sillon nord-pyrénéen (Aquitaine, France)

This paper presents for the first time the inventory of the Paleocene and Lower Eocene foraminifers located in the North Pyrenean trough, between the Atlantic Ocean and the neighbourhood of the town Pau. They have been studied from three outcrops. The Bidart Beach section shows the Lasseube Formation from the Cretaceous/Paleogene boundary to the base of the P 3a zone. The Loubieng Quarry section, near Orthez, represents the upper P 3a zone and the lower P 3b zone as well as the Lasseube Formation / Pont Labau Formation boundary. The interval between the upper part of the P 3b zone and the upperest part of the P 5 zone crops out along the Gan - Rébénacq road with a hiatus located at the Paleocene / Eocene boundary, the whole interval belonging to the Pont Labau Formation. 394 taxons of foraminifers are present in this formations: 349 benthic and 45 planktonic species. The Velasco type benthic foraminifers show a middle bathyal depositional environment, with a paleobathymetry included between 500-600 m and 1000 m: Nuttallides truempyi, Osangularia velascoensis, Bulimina trinitatensis. The Midway type species which were transported by the turbidite currents from the lower to middle neritic environments are frequent as well as the Cretaceous reworked species. The species number is low: 29 in the iridium layer of the Cretaceous/Paleogene boundary (P 0 zone). Fauna grows rich quickly in the Pα zone reaching 129 species. The diversity grows up progressively from P α to the P 4a zone (NP 8). The disappearances are rare until his horizon, but their number is bigger than the number of appearances from the P 4b zone. It reaches their maximum between the P 4c zone and the P 5 zone. It shows that the decline of the Paleocene fauna begins around 2 million years before the thermal event of the Paleocene / Eocene boundary in the Aquitaine sections. The disappearances stay important in the Lower Eocene - Ypresian, but the appearance of lots of Eocene species show that the fauna renewal is located in this stage. The Cretaceous taxa dominate in the Paleocene benthonic fauna. The appearance or the disappearance of some species has a stratigraphic value in the Aquitaine region: the disappearance of Coryphostoma incrassata in the P 1b zone, the appearances of Plectina dalmatina, Elongobula grata (P α); Bulimina tuxpamensis (P 2); Tritaxilina cubensis, Thalmannita madrugaensis (P 3a); Svenia bulbosa (P 3b); Discorbis perovalis (P 4a/NP 7); Elongobula pulchra, E. pupa, Asterigerina bartoniana, Neorotalia gr. tuberculata (P 4a/NP 8); Bigenerina pannonica, Pentellina pseudosaxorum (P 5/NP 9-10).     

La transgression thanétienne (Paléocène supérieur) dans l’Aurès occidental (Algérie), d’après les associations de Foraminifères de la coupe d’El Kantara

In the W-Aures (Algeria), the El Kantara pass displays about 50 m of Red Marls overlying the Maastrichtian limestones with Laffitteina. The Red Marls are, in their turn, overlain by the Thanetian marly limestones of the river El Haï. The micropaleontological inventory of these Red Marls establishes the co-occurrence of subaerian (Microcodium = Paronipora), fresh-water (Charophyta) and marine microfossils (Foraminifers). The stratigraphical significance of these microfossils is discussed. The upper part of the Red Marls, yielding abundant Valvulina and scarce Glomalveolina, is of Thanetian age. The age of the lower part is less established, but the occurrence of Microcodium suggests a Thanetian age too. Located at the margin of the northern opening of the trans-Saharan epeiric sea, the El Kantara section establishes a Thanetian age for the beginning of the Paleocene transgression. Danian deposits are missing, in spite of the absence of an obvious hiatus on the field. After general emersion during the early Paleocene, the Thanetian transgression starts with the set up of lakes at the depositional area of the Red Marls. Marine influences, limited at first, become gradually prevailing, and end with the deposition of neritic marly limestones of river El Haï.     

Charophytes du Crétacé/Paléocène du Moyen-Atlas (Maroc) : systématique et implications biochronologiques

Gyrogonites of charophytes from two localities of the Oudiksou syncline (Middle Atlas, Morocco) are studied. The Irbzer formation in the Achlouj-2 site yielded charophytes (Feistiella oblonga, F. globosa, Strobilochara apiculata, S. diademata) that suggest a Campanian-Maastrichtian age; this result is consistent with all previously published biochronological studies. The overlying Bekrit-Timahdit formation, supposed to be Thanetian to Lutetian in age, without paleontological evidence, yielded charophytes in the Saf locality (Peckichara cf. llobregatensis, Feistiella sp. 1, Maedleriella sp.). This association indicates a Paleocene age for the middle member of this formation.     

Lithostratigraphie et biostratigraphie des formations paléocènes et éocènes entre Bayonne et Pau (SW France)

The lithostratigraphy and biostratigraphy of the Paleocene and Eocene formations have been reconsidered on the basis of 18 dated boreholes that intersect the Basque Country, Peyrehorade, Sainte-Suzanne and Sauvelade allochthonous tectonic units, as well as the autochthonous units of the Pey-Saint-Lon anticline, the Arzacq Basin and the North Pyrenean Trough. Correlation of the formations identified in the boreholes and at outcrop has made it possible to outline the sedimentary evolution of this region during the Early Paleogene and to date the various stages of Pyrenean compression on the basis of sedimentary gaps and syntectonic detrital input. The Lasseube Formation is present over the entire study area. Sedimentation becomes differentiated during the Late Selandian-Early Ypresian period. In the North Pyrenean Trough and in the Peyrehorade, Sainte-Suzanne and Sauvelade units, the detrital-clayey Pont-Labau Formation is well developed and all its members are represented (Guillempau, Latapy, Néez, Libaros and Artigueloutan; zones upper P 3b-P 6b; upper NP 5-NP 11). A sedimentary gap showing erosion, and involving zones upper P 3b-upper P 4a and upper NP 5-NP 7, characterises the Basque units, the Pey-Saint-Lon anticline and the western part of the Arzacq Basin. Sedimentation recommences in zone P 8 with middle-bathyal deposits of the Cassoua Member and then continues with the Artigueloutan Member (transition facies towards the Louts Formation) or limestone resembling that of the Meilhan Formation. The Gan Formation shows an east-west polarity. In the east, between the Sainte-Suzanne Unit and the North Pyrenean Trough, the formation is argillaceous with a dominant neritic character, apart from a circalittoral to epibathyal passage in its lower part (zone P 7) containing numerous planktonic foraminifera. The formation is of a detrital and carbonate nature in the west (Sainte-Marie-de-Gosse 1). The bathyal Donzacq Formation appears to transgress a previously deformed surface. Two domains existed during the Lutetian: a continental to neritic domain in the south and a bathyal domain in the north (the Saint-Geours-d'Auribat and Miretrain formations) separated by a variably thick transition zone that is affected by thrust fronts. Two new formations - the Orthez and Sorde-l'Abbaye formations - are proposed, and the Marly limestone formation of Saint-Barthélémy (Douvillé, 1905) is redefined. The Orthez Formation (Sauvelade Unit) is composed of marl and continental conglomerate with neritic carbonate interbedding and belongs to the sub-zones NP 15a and b. The Sorde-l'Abbaye Formation (Peyrehorade Unit and the Basque units p. p.) is composed of neritic carbonate sediments (Lutetian-Priabonian). The Saint-Barthélémy Formation is made up of syntectonic conglomerate, turbiditic limestone and pelagic marl in the central part of the Basque units (NP 15a-NP 16). The Bartonian and Priabonian stages are characterised by carbonate sediments in the Arzacq Basin (Siest Formation). Its base is diachronous: Upper Lutetian (zone SB 16) in the east and Late Bartonian grading to Priabonian in the west (zones SB 17/18 and SB 19). The sediments are argillaceous (Côte des Basques Formation) along the Atlantic coast, in the central part of the Basque units and in the Bayonne-Cauneille syncline. Major thickness variations reflect tectonic phases at the top of the Lasseube Formation in the north (erosion pre NP 8), the Néez Member (NP 7/NP 8 boundary), the Artigueloutan Member (NP 11/NP 12 boundary), and the Gan Formation (P 8/P 9 boundary). The syntectonic conglomerate and breccia indicate the development of thrust fronts in the Libaros Member (NP 8), at the base of the Pillardou and Artigueloutan members (NP 9b, c-NP 10), in the Saint-Geours-d'Auribat Formation (NP 14b) and in the Orthez and Saint-Barthélémy formations (between zones NP 15a and NP 16).     

La microflore de diatomées au passage mio-pliocène en Corse

The “Aleria formation”, described in northeastern Corsica, consists of siliciclastic deposits referred to a deltaic environment and comprises diatomitic lenses exposed at Casabianda. The Aleria formation fills a depression of the Messinian erosional surface and is overlain by Early Pliocene sediments in several places. This suggests a late Messinian-earliest Pliocene age. Diatom microflora is well-preserved and contains 59 species. Pennates indicate a very large diversity with 54 species, while Centrics are represented only by five species. Diatom assemblages show a relative constant composition along the studied interval. The species Aulacoseiragranulata is the dominant taxon with about 90% in all samples. Taxa are of different salinity classes and different modes of life (planktic, tychoplanktic, benthic, epiphytic…), suggesting multiple environmental factor interactions. The environmental model suggested by sedimentological and biological data led as to assume a pond-like coastal receptacle common in deltaic system. This receptacle was slightly influenced by the close marine waters and was supplied by permanent freshwaters from continental run-off and alluvial sheet. This model based on diatom assemblages provides new data for palaeoenvironmental and chronological interpretations of the so-called Messinian Lago-Mare deposits.     

Les assemblages de diatomées du bassin messinien de Boudinar (Maroc nord-oriental)

The Messinian is one of the strongest biogenic silica accumulation periods in the world and more particularly in the Mediterranean, where it is marked by an important diatomitic sedimentation. In the Boudinar basin (Morocco north-Eastern, Western Mediterranean), the section of Sidi Haj Youssef, localised near the volcano of Ras Tarf, has approximately 100 m thickness of clayey-marly series in which 12 diatomitic levels of variable thickness are intercalated. The microfloristic study of diatoms on 86 samples, carried out in detail for the first time, made it possible to recognize 50 genus of diatoms (24 of centric and 26 of pennate) represented by 185 species (75 species of centric and 110 species of pennate). Four hundred individuals were taken from each sample to determine the relative frequency of each taxon within the diatoms assemblages. Thus, several assemblages were defined by the predominance of the following species: Coscinodiscusmarginatus, Actinoptychussenarius, Thalassionemanitzschioides, Actinocycluscurvatulus, Thalassiothrixlongissima, Rhizosoleniastyliformis and Actinocyclusehrenbergii. These diatoms assemblages display a littoral marine environment in communication with the opened sea. The abundance of the cold water species towards the base and the top of the section suggests broad exchanges of the basin with the Atlantic Ocean in Messinian. The predominance of the species Thalassionemanitzschioides and/or Thalassiothrixlongissima indicates periods of strong productivity that can be associated to upwelling systems.     

Pléistocène moyen et inférieur dans le Latium (Italie centrale)

Explorations and diggings of the Italian Institute of Human Palaeontology in Latium from 1950 to 2005, have brought out the following composite sequence: (1) for upper-middle Pleistocene of northern Latium: Travertine, gravels Acheulian-Mousterian transition, Riss. Homo (femur), Elephas antiquus, Hippopotamus, Bubalus murrensis, with upper Acheulian artefacts. (2) In middle Latium, middle Pleistocene: Volcanoclastic K-Ar 360 Ky. Below: Lower Acheulian complex and bone artefacts. Homo, Inuus, Elephas antiquus, Ursus deningeri, Dama clactoniana. Volcanic ash with Zelkowa, Buxus: caucasian flora. Hot pyroclastic flow about 15 m (50 feet) thick between 520 and 530 Ky. Limno-tuffite with Taxodiacea flora Lower-middle Pleistocene choppers artefacts below volcanic limit of 700 Ky. Southern Latium, lower Pleistocene: travertine reed Phragmites fragments. Ceprano hominid calvarium 800-900 Ky old. Gravel with chopper artefacts. Red sand with Unio shells. Lower palaeolithic gravelly sand, with very rough choppers artefacts, at Arce, Colle Marino, Colle Pece localities; at Castro dei Volsci chopper, assemblage is more evolved. Unconformity. Yellow sand layer with middle Villafranchian Anancus arvernensis and Mammuthus meridionalis fauna.     

Variabilité dentaire et crânienne de Numidotherium koholense (Mammalia, Proboscidea) de l’Éocène d’El Kohol, Algérie

The variability of one of the oldest known proboscideans, Numidotherium koholense from the Lower Eocene of El Kohol, Algeria, is here examined. The biometrical study of the jugal teeth shows that the population is homogeneous and corresponds to a single species. The important variations of the skull's shape are interpreted as resulting from sexual dimorphism. The dental variability is analysed and discussed from an evolutionary perspective regarding the adaptative radiation of the proboscidean order at the beginning of the Tertiary.     

Nouvelles observations sur les Miogypsinidés du Miocène inférieur et moyen de Provence et de Corse (France) et de Sardaigne septentrionale (Italie)

The detailed study of the Miocene strata of Bonifacio has revealed an atypical Miogypsinid assemblage. In addition to the three already cited genera, Miogypsinoides, Miogypsina and Miolepidocyclina from this region, Miogypsinodella is for the first time represented by two new species (Mdella corsicana nov. sp. et Mdella pillaria nov.sp.). The stratigraphical ranges of each genus in the Miogypsinidae are not verified here. However, Mdes bantamensis is still present in the Upper Burdigalian and the genus Miogypsinoides is also present in the Lower Langhian. Six species of Miogypsina coexist in the Upper Burdigalian (M. tani, M. globulina, M. intermedia, M. cf. sabahensis, M. cushmani, M. mediterranea) and two species are present in the Lower Langhian (M. antillea, M. digitata). This distribution is apparently an example of palaeoendemism resulting from the geographic isolation and the rotation of the Corso-Sardinian block and also from the palaeogeographical and palaeoecological favourable environment during that time.     

Signification paléoécologique des assemblages de diatomées du Messinien dans le bassin de Melilla-Nador (Rif Nord-Oriental, Maroc)

In the Melilla-Nador basin, the diatomites represent one of the main sedimentary components of the Messinian together with reefal carbonate platforms. Two vertical sections were realised in the diatomitic levels from distal platform. Diatom assemblages are for the first time described in this basin. A total of 125 diatom species were determined: 73 centrics and 52 pennates. Significant diatom assemblages defined from the statistical analysis indicate ecological variations concerning diatomitic levels or parts of diatomitic levels. Using the data sets we propose paleoenvironmental models showing the complexity of the ecological interactions. This study point the permanence of opened marine environment, the impulse of cold waters from Atlantic origin, a general decrease in bathymetry in the Messinian marls-diatomitic series, the influence of the carbonate platform progradation and the occurrence of upwelling systems.     

Stratigraphie, paléoécologie et foraminifères du paléogène des Alpes Maritimes et des Alpes de Haute-Provence (Sud-Est de la France)

Paleogene sedimentation in the Alpes Maritimes and Alpes de Haute Provence regions is composed of one complete sedimentary sequence. This sequence begins with the continental Formation from Microcodium and continues with the Cerithium layers, the Calcaires Nummulitiques and the Marnes Bleues, which are overlaid by the turbidites of the Grès d’Annot Formation. Sedimentation starts in the Nummulites perforatus zone close to the base of ‘the Truncorotaloides rohri zone (P 14) and ends in the Cassigerinella chipolensis-Pseudohastigerina micra zone (P 18) and the NP21 zone in the upper part of the Grès d’Annot. More biomarkers were used in order to define a more detailed local biozonation (biozones AMP 1 to AMP 7). Four local zones were also defined by the last occurrence of Nummulites millecaput and N. perforatus-N. ptukhiani and then by the first appearance of N. retiatus (AMGF 1-4). The evolution of benthic foraminiferal assemblages shows an increase in bathymetry from the internal platform in the Cerithium layers to the calcareous platform with large foraminifers, then to the external platform and the deep offshore environment of the Marnes Bleues and Grès d’Annot. Low faunal diversity in the Grès d’Annot together with the predominance of agglutinated species indicates a poorly oxygenated, organic rich and turbidite environment. Seven steps (SD 1 to SD 7) in the Eocene marine transgression are shown from East to West by detailed local biozonation and sequence analysis. Grès d’Annot sedimentation is also diachronous, beginning within the Truncorotaloides rohri zone (P 14, AMP 1) in the East and ending within the Pararotalia opima opima zone (P 20, middle Rupelian) in the West (Barrême). Small foraminifer Paleogene fauna from the Alpes Maritimes and Alpes de Haute Provence was studied from 400 samples. It is composed of 378 species. Two new taxa are proposed: Fissurina niceana n. sp. and Globocassidulina alpina n. sp. The species from the Escarène and Gorbio neighborhood described by M. von Hantken (1884) were re-examined.     

Groupe de Candona clivosa, nouveau groupe de Candoninae (Crustacea, Ostracoda) et sa diversification dans le Bassin de Turiec (Slovaquie) au Miocène supérieur

The Candona clivosa-group, a new group of Candoninae - is erected on the base of the folded inner lamella observed on the postero-ventral border of the female valve. The elongated sub-triangular and rectangular valves of this group are described from the Miocene and Quaternary deposits of Europe. A folded inner lamella is also observed in 11 species of Candona from the Upper Miocene of the Turiec Basin of which 8 are new - Candona fatrica nov. sp., C. tatrica nov. sp., C. lacustris nov. sp., C. palustris nov. sp., C. stagnosa nov. sp., C. laterisimilis nov. sp., C. jiriceki nov. sp. and C. incurva nov. sp. They show a higher variability of the outline, calcification of the valves and width of the zone of concrescence. The extrinsic and intrinsic aspects of this morphological diversification in the Turiec Basin are discussed.     

Données paléoenvironnementales sur le Vraconnien/Cénomanien de la région de Tébessa (Atlas Saharien,nord-est Algérie). Caractérisation de l’OAE2

The micropalaeontological content of five sections, located in Northeastern Algeria (Saharan Atlas) was investigated by means of washing/counting of microfossils in marly levels, and microfacies analyses of calcareous levels. In these levels of Upper Cretaceous age, hundred species of foraminifera were identified but only about 15 species of ostracoda and about ten of radiolaria. This work allows, first of all, to establish a rather precise stratigraphic frame, in particular by means of studying planktonic foraminifera. Two to five biozones were defined, between the Vraconnian (Th. Appenninica biozone) and the early Turonian (Whiteinella archaeocretacea biozone, then Helvetoglobotruncana Helvetica biozone for certain sections). Secondly, the quantitative analyses led on foraminifera allowed the definition of palaeoenvironment. The ratio P/P+B, generally very high, coupled with a little diversified benthonic microfauna, indicates a calm and deep environment, of external platform or slope type. Furthermore, at numerous levels, various indications give evidence of the existence of a strong surface productivity (presence of upwellings), responsible for the proliferation of radiolaria (late Vraconnian/early Cenomanian especially) or of globular planktonic foraminifera (hedbergellids/heterohelicids); being both associated with low-oxygen deep waters. Two anoxic events were also revealed, the first one at the end of Vraconnian (OAE1d) and the second at the end of Cenomanian (OAE2). This last event in particular was characterized on all the sections, in a more or less detailed way, thanks to the identification of certain indicator: Heterohelix “bloom”, “filament” event, disappearance of rotaliporids, presence of “blackshales” strongly enriched in organic matter (Bahloul levels).     

Les restes humains du Pliocène final et du début du Pléistocène inférieur de Dmanissi, Géorgie (1991-2000). I - Les crânes, D 2280, D 2282, D 2700

Since 1991, several human remains: 5 skulls, 4 mandibles and numerous postcranial fragments have been discovered on the Dmanissi prehistoric open site. It is an exceptional discovery due to the stratigraphical, paleontological and cultural context, which is well known and accurately well dated (Upper Pliocene-Early Pleistocene). Most of the hominids discovered in the level V and VI are dated between 1.81 My (level V) and 1.77 My (level VI) corresponding to a 40,000 years period. The assemblage of fossil human remains is peculiar due to (1) the quality of bone representation (distinct parts of the skeleton are preserved: skull, thorax, upper and lower limbs, belt), (2) the high degree of bone preservation (skulls and long bones are entire, rarely broken or crushed), (3) the diversity age at death estimated for each of the 5 individuals (3 adults, 1 young adult, 1 adolescent of both sexes). The study dealing with the first discovered mandibles and skulls has begun with Leo Gabounia since 1991 and represents several interests: 1) a paleoanthropological interest: the Dmanissi skulls are characterized by their small size; they are short, narrow and low. The skullcaps are less elevated than those of the Homo erectus group and even those of Homo ergaster. They are more elevated than those of Homo habilis and very close to Homo rudolfensis. The elevation and the transversal development of the middle part of the skull in the parietotemporal region are more significant: the Dmanissi specimens are intermediate between Homo habilis and Homo ergaster. In term of cranial capacity, a similar trend is observed. Generally speaking, the skull is slender. The vault is more flat than in Homo erectus, the frontal bone is less developed, divergent and the postorbital constriction is strong. The temporal bone is long, flat and the mastoid part is short. The upper part of the occipital bone is low and narrow. Crests are thin, less developed than in the Homo erectus group. The superior temporal crests are in a high position and a torus angularis is present on the adult-male specimen. The glenoid cavity is large with strong edges. The petrotympanic region is slender with a tympanic circle individualized and it shows a horizontal rotation in a posterior position, which is distinct from Homo erectus. The orthognathic trend of the face distinguishes the Dmanissi specimens from the early Pleistocene hominids (Homo habilis, Homo ergaster) and from the first Eurasian Homo erectus. Nevertheless, the subnasal region of the face is projected. The morphology of the mid-face, showing a developed pillar of the canine, an inframalar incurvation and an anterior position of the root of the zygomaticomaxillary crest, suggests strong masticatory stress. Considering the overall morphology, cranial and metrical features, the Dmanissi fossil skulls are intermediate to the Homo habilis-rudolfensis group and Homo ergaster while they are closer to the former and peculiarly to Homo rudolfensis (ER 1470). However, the Dmanissi fossil skulls are distinct from Homo rudolfensis by numerous features and among them: by their large maximum cranial width (Euryon-Euryon), the posterior rotation of their petrotympanic structure and the strong development of the pillar of their canine. Due to the gracility of their face, the narrowness of their occipital bone, and their cranial base pattern (mastoid region and petrotympanic structure), the Dmanissi fossil skulls are different from the Homo erectus group: 2) the abundance of the human fossils discovered in Dmanissi site provides information about the biodiversity of these hominids with the establishment of the morphological features related to either growth or sexual patterns: 3) compared to modern humans, the Dmanissi fossil skulls seem to follow a different growth pattern. The present study of the fossil skulls discovered is a pioneer step. Indeed, the Dmanissi site has yielded the oldest evidences of the first settlements in Eurasia, which were, until now, attributed to Homo erectus. The Dmanissi fossil skulls are close to the Homo habilis-rudolfensis African group. We attribute these hominids to Homo georgicus.     

Biostratigraphie,paléoécologie et zones d’acmé des foraminifères planctoniques au passage Crétacé-Paléogène dans la Téthys (Tunisie et Espagne) et l’Atlantique (France)

Biostratigraphical high-resolution analyses and quantitative data confirm that deposition is continuous across the K-Pg transition in several sections in Tunisia (El Kef, stratotype section) and Spain (Agost and Caravaca sections) located in the Tethyan realm and the Bidart sections in the Atlantic realm, without any relevant hiatus. The Upper Maastrichtian assemblages of planktic foraminifera from these sections are largely dominated by small biserial heterohelicids. They are associated to common species having planispiral test (i.e. globigerinelloids), trochospiral test (i.e. hedbergellids, rugoglobigerinids globotruncanids), to rare triserial heterohelicids (i.e. guembelitriids) and trochospiral species showing tubulospines (i.e. schackoinids). Stratigraphical ranges of these diverse taxa through the late Maastrichtian in the Tethyan and Atlantic realms show very few changes in the planktonic foraminiferal assemblages and most of the species are present in the Abathomphalus mayaroensis biozone. By our high-resolution sampling and the intensive research for the A. mayaroensis index species in the uppermost Maastrichtian samples, we confirm that this species is omnipresent up to the top of the Maastrichtian. Therefore, A. mayaroensis is present in almost all samples which are late Maastrichtian in age, but this species became very scarce in the uppermost Maastrichtian samples. This scarceness could be due to a climate cooling. A sharp decrease in relative abundance of the deep dwellers species, like as Abathomphalus intermedius and A. mayaroensis as well as in other keeled globotruncanids is observed at the studied sections from the Tethyan realm (indicative of low latitude) across the latest Maastrichtian. At the K/Pg boundary, all the globotruncanids disappeared. They are considered specialists living in tropical-subtropical deep seawater habitat. At this boundary, large and ornate heterohelicids also disappeared. Therefore, all the studied sections show that about 90 % of the Maastrichtian species became extinct according to a catastrophic mass extinction pattern. Only about 10 % crossed the K/Pg boundary and survived during the earliest Danian. The minor difference in the number of disappeared taxa is related to their latitude location or environment paleodepth. The changes in the species relative abundance, observed in the successive planktic foraminiferal assemblages, make it possible to recognize the Acme-stage 0 typical of the upper Maastrichtian interval. It is characterized by the highest species richness of Globotruncanids and heterohelicids specialists of tropical to subtropical marine conditions, the Acme-stage 1 typical of the Guembelitria cretacea Zone, and in particular of the Hedbergella holmdelensis Subzone dominated by “opportunists” species belonging to Guembelitria, the Acme-stage 2 which corresponds to the Pv. eugubina Zone dominated mainly by specimens belonging to Palaeoglobigerina and Parvularugoglobigerina and the Acme-stage 3 which characterizes mainly the Ps. pseudobulloides Zone dominated by biserial species belonging to Chiloguembelina and Woodringina.     

La macrofaune du Pléistocène supérieur d’El Harhoura 2 (Témara, Maroc) : données préliminaires

The El Harhoura 2 cave is situated in the commune of El Harhoura in the Temara region (Morocco). The entrance faces west, towards the ocean, around 200 m away from the current shoreline and about 16 m above sea level. The stratigraphy has currently 11 levels numbered from top to bottom. Level 1 is attributed to the early Neolithic. The archaeological material from level 2 places it within the Upper Palaeolithic (Iberomaurusian). Levels 3 to 11 are assigned to the Middle Palaeolithic (Aterian). Since 1977, a surface of 37 m² has been excavated, yielding particularly abundant remains of a large mammal fauna study in this paper.     

Réévaluation de la présence humaine au Pléistocène inférieur dans le Sud de l'Espagne

The Early Pleistocene locality at Venta Micena (Orce, Guadix-Baza basin, province of Granada, Spain) has provided four fossil remains - skull fragment VM-0, and long bone diaphyses VM-1960, VM-3691, and VM-12000 - which have been tentatively attributed to the hominids. Although several methodologies have been used to ascertain the human affinities of these specimens - including anatomical, morphometric and immunological analyses - the results obtained have not been conclusive, instigating a persistent debate. A taphonomic approach is used here for estimating the probability that a taxon the size of Homo sp. (~ 50 kg) could be represented in the fossil assemblage by four bone fragments and no tooth remain. A least-squares regression analysis between the percentage of teeth and the body mass estimated for each taxon of large mammals (N = 20) predicts a raw abundance of six teeth for Homo sp. in the assemblage. Given that up to the present moment no tooth remains attributable to the hominids has been unearthed during systematic excavations in the Venta Micena quarry, which has provided more than 15,000 fossils of large mammals, this argues strongly against the possibility that the three bone specimens could belong to Homo sp. The phalanx CV-0 from the Early Pleistocene site of Cueva Victoria (Cartagena, Spain) has also been attributed to the genus Homo. The taxonomic assignment of this specimen is biased, however, because it was not compared with Theropithecus oswaldi, the only primate species actually recorded from this karstic locality. A comparative anatomical and morphometric analysis of fossil and modern specimens of Theropithecus suggests that CV-0 can be attributed to T. oswaldi. As a result, Cueva Victoria does not contribute additional information concerning the first human settlements in Europe. By these reasons, apart from the paleoanthropological and archaeological findings from Atapuerca (TD lower levels and Sima del Elefante), the rich archaeological assemblages from Barranco León and Fuente Nueva-3 in Orce, dated 1.3-1.2 Myrs, which include fourteen hundred stone tools of Oldowan technology, constitute at present the only unequivocal evidence of human presence in Southeast Spain during Early Pleistocene times.     

Puycelci, nouveau site à vertébrés de la série molassique d’Aquitaine. Densité et continuité biochronologique dans la zone Quercy et bassins périphériques au Paléogène

After the previous lacking evidence allowed the interpretation of a non-deposition episod, the newly recorded mammal fauna from Puycelci (Tarn, SW France, close South to the Quercy paleokarstic area) now shows the MP 26 standard level (Late Oligocene, Early Chattian) being represented within the Eastern Aquitaine molassic basin filling. According to the evolutionary stage of known mammal lineages, the most significant Puycelci fossils are the Issiodoromys pauffiensis rodent and the Metriotherium cf. sarelense nov. sp. (described in this paper) artiodactyl ungulate. This new dating exemplifies the improvement of the mammalian biochronologic record as regards both the Quercy paleokarst and the surrounding lacustrine Tertiary basins W, S, and S-E to the Quercy, as well as, directly E and N-E, the smaller tectonic Tertiary basins within the crystalline Massif Central basement. The updated record shows the strong extension in time of the paleokarstic data, down to the Middle and Early Eocene, and up to the Early Miocene. Regarding the classically reported Late Eocene to Late Oligocene span, as well in the paleokarst and the peripheral basins, the available biochronological data now attains a valuable density level, the variation of which is significant. On one side there are periods (Late Eocene, Early Oligocene, Early Late Oligocene) with high biochronologic density, allowing improved time resolution using evolutionary stages within mammalian lineages (e.g. numerical ages). Such periods corresponds to almost continuous sedimentary processes, with only possible short breaks. On the other side, there are periods with unreported data, shared in the paleokarst and peripheral basins (Late Oligocene as best example). Such periods may well correspond to some non-deposition episods as once alleged by geologists. Then the surrounding biochronologic data would bear constraining value regarding these episods.     

Les Cervidae (Mammalia, Artiodactyla) du Pléistocène supérieur-Holocène ancien de la région du Parc National Serra da Capivara (Piauí, Brésil)

The Upper Pleistocene/Lower Holocene fossil-bearing sites of the Serra da Capivara National Park Region have yielded three cervid species: Mazama gouazoubira, M. americana and Blastocerus dichotomus, all currently living in South America, the two first in the region. A grand total of more than one hundred remains demonstrates the presence of Mazama gouazoubira in seven sites, mainly the Toca das Moendas, the Toca do Serrote do Artur, the Toca da Cima dos Pilão. This small species shows, since the Upper Pleistocene, a conspicuous tendency to reduce the average dimensions of its teeth and long bones. From the taller M. americana, only a dozen of remains were found in four sites, mainly the Sitio do Meio. In all of these it is sympatric with M. gouazoubira. It differs from this last one by its cheek teeth and its limb bones size and proportions. The oldest site where the species is known is Tarija (Bolivia, Middle Pleistocene) and it does not show any significant changes in size and proportions between recent and fossil samples. Sixteen remains of the large B. dichotomus were found in five sites, mainly the Toca das Moendas and the Toca da Barra do Antonião. The species is a rare fossil, but is frequently figured in the rock art painting of the region, where it is presently unknown.     

Fimbria lohani (Mollusca : Bivalvia) nomen novum pro Fimbria subpectunculus (d’Orbigny, 1850) du Lutétien (Éocène moyen) du bassin de Paris,nom préoccupé

Fimbria lohani (Mollusca: Bivalvia) nom. nov. pro Fimbria subpectunculus (d’ Orbigny, 1850) from the Lutetian (Middle Eocene) of Paris basin, name preoccupied. The names Fimbria magna ( Anton, 1838) and Fimbria subpectunculus (d’ Orbigny, 1850) are nomenclaturally invalid to designate the species from the Lutetian (Middle Eocene) of the Paris Basin. Fimbria lohani nom. nov. is proposed here as a replacement to correct the homonymy Corbis subpectunculus d’ Orbigny, 1850 non d’ Archiac, 1850.