The exceptionally diversified Neotropical genus Veturius Kaup: more new data on its phylogeny,taxonomy and distribution (Coleoptera: Passalidae)

New data are provided on 24 known and five new species in three subgenera of Veturius Kaup (Proculini), from Central and South America. Veturius s. str.: V. hincksi Boucher, 2006 (wing polymorphism, Colombia: Nariño), V. perecasi Boucher, 2006 (Colombia: Boyacá, Santander, perhaps Risaralda), V. montivagus Boucher, 2006 (first precise location, Colombia: Tolima), V. schusteri Boucher, 2006 (Costa Rica: Guanacaste, Cartago, Alajuela), V. platyrhinus (Hope & Westwood, 1845) (Colombia: Tolima), V. aspina Kuwert, 1898 (polymorphism, Colombia: Sucre, Cesar, Córdoba, Antioquia), V. yahua Boucher, 2006 (Colombia: Amazonas, Meta), V. ecuadoris Kuwert, 1898 (Colombia: Caquetá), V. muisca n. sp. (Colombia: Cundinamarca), V. nonuya Boucher & Salazar, n. sp. (Colombia: Amazonas), V. standfussi Kuwert, 1891 (polymorphism, Colombia: Caquetá; Peru: Piura), V. arawak Boucher, 2006 (Peru: Cuzco, Puno), V. fabieni Boucher & Salazar, n. sp. (Peru: Junín), V. dreuxi Boucher, 2006 (Bolivia). Publius Kaup: V. crassus (Smith, 1852) (Colombia: Cundinamarca, Boyacá, Meta, perhaps Tolima), V. rugifrons Boucher, 2006 (Colombia: Cundinamarca, Boyacá), V. concretus (Kaup, 1868) (Colombia: Cundinamarca, Boyacá, Meta), V. tectus Boucher, 2006 (second record, Colombia: Santander), V. dupuisi Boucher, 2006 (second record, Colombia: Cundinamarca), V. centralis Boucher, 2006 (second record, Colombia: Antioquia), V. taurus Boucher, 2006 (second record, Colombia: Norte de Santander), V. spinipes Zang, 1905 (topotype, Bolivia). Ouayana Boucher: V. paraensis Luederwaldt, 1927 (Brazil, Pará), V. negroensis Boucher, 2006 (Colombia: Vaupés; Brazil: Amazonas), V. amazonicus Boucher, 2006 (Colombia: Guainía), V. quaesitor Boucher & Salazar, n. sp. (Peru: Piura), V. uncinatus n. sp. (Colombia: no other indication), V. casalei Boucher, 2006 (Colombia: first precise location, Caldas, Antioquia), V. fanestus Boucher, 2006 (Colombia: Nariño). Veturius contains now 86 species, and is by far the most diversified recognized monophyletic genus in the Passalidae.     

A new Veturius (Veturius) of the “cephalotes” group from southern Colombia,Putumayo (Coleoptera: Passalidae)

A new species of Veturius (Veturius) Kaup from southern Colombia, Putumayo, lowland forests, is described and illustrated. V. paya n. sp. belongs to the South American “cephalotes” species group and is sister species of V. cephalotes (Le Peletier & Serville, 1825). Both species are distinguished by few characters of the head and thorax. They seem to be allopatric close to the western distributional limits of V. cephalotes. The endemism of V. paya n. sp. corresponds to a geographic vicariance in the axis of the Amazon Basin. An overview is given on the phylogeny and the chorology of the completed “cephalotes” species group.     

Three new species of Philocorydoras Suriano, 1986 (Monogenoidea: Dactylogyridae) infecting the gills of callichthyids (Actinopterygii: Callichthyidae) from the Peruvian Amazonia

During a research on gill ectoparasites of callichthyids fishes from the Peruvian Amazonia, the following monogenoideans were found: Philocorydoras peruensis n. sp. from Corydoras splendens (Castelnau); Philocorydoras multiradiatus n. sp. and Philocorydoras jumboi n. sp. from Brochis multiradiatus (Orcés, V.). All new species described herein are mainly differentiated from their congeners based on the morphology of the copulatory complex. In P. peruensis n. sp. the cirrus is “J”-like shaped tube slightly tilted to one side, while in P. jumboi n. sp. is “J”-like shaped tube in a straight position and in P. multiradiatus n. sp. the cirrus is an arced tube with inflated base and distally narrow. Brochis (Orcés, V.) represents a new genus hosting species of Philocorydoras. All new species presented in this work represent the first species of Philocorydoras reported for Peru.

     

New records of Polypedilum Kieffer (Diptera: Chironomidae) from India

Adult males of Polypedilum (Polypedilum) exterflexus n. sp., P. (Tripodura) rectangulum n. sp. and life stages of P. (Tripodura) pruina Freeman are described from India. In addition, one species, Polypedilum clavipennae n. sp. with all the life stages not assigned to any one of the existing subgenera is also described, and it may be justified to erect it as a new subgenus in future.     

Further study on the phylogeny,taxonomy and distribution of the genus Veturius Kaup in Central and South America (Coleoptera: Passalidae)

This study provides data on the phylogeny, taxonomy and distribution of 14 known and five new species of the Neotropical genus Veturius Kaup (Proculini), belonging to various subgenera and species groups: V. (Veturius) latissimus n. sp. (Colombia, Central Andes) and V. (V.) calimanus n. sp. (Pacific slope of the Occidental Cordillera) are separated from V. (V.) caquetaensis Boucher, 1988, which seems restricted to the Amazonian slope of the Oriental Cordillera (Caquetá, Putumayo); V. (V.) sinuatomarginatus Luederwaldt, 1941 (Costa Rica), n. syn. of V. sinuatocollis Kuwert, 1890; V. sinuatocollis aculeatus Luederwaldt, 1941 (syntype from Costa Rica); V. (V.) aspina Kuwert, 1898 (located in Occidente of Ecuador, Guayaquil); V. (V.) yahua Boucher, 2006 (located in Occidente of Ecuador, Pichincha and SW Colombia, Nariño); V. (V.) guntheri Kuwert, 1898 (located in Peru, SE Puno and Colombia, W Putumayo); V. (V.) cephalotes (Le Peletier & Serville, 1825) (citation from Guyana); V. (V.) sinuatus (Eschscholtz, 1829) (previous synonymy); V. (V.) libericornis Kuwert, 1891 (located in Peru, Cuzco); V. (V.) lepidus Fonseca, 1999 (revision; located in Colombia, Amazonas, Putumayo and Peru, Loreto); V. (V.) transversus (Dalman, 1817) [syntype; previous synonymy of V. trituberculatus (Eschscholtz, 1829) with V. assimilis (Weber, 1801) and located in Brazil, Mato Grosso]; V. (V.) sinuosus (Drapiez, 1820) (corrected reference for Colombia); V. (Publius) crassus (Smith, 1852) (new syntype); V. (P.) danieli Boucher, 2006 (holotype deposit); V. (P.) vazdemelloi Boucher, n. sp. (Andes of Ecuador, Azuay); V. (Ouayana) unicornis Gravely, 1918 (located in Colombia, E Vaupés); V. (O.) costaianus Boucher, n. sp. (Venezuela, Amazonas, NW Pacaraima Massif); Ticoisthmus Boucher, n. subg., for the species group of V. (O.) laevior (Kaup, 1868), of southern Central America; and V. (T.) brachypterus Boucher, n. sp. (Costa Rica, Sierra Talamanca). Ticoisthmus is considered the sister group of Ouayana. It belongs to the Meso-American low mountain dispersion pattern and demonstrates, especially in the genus Veturius, but also more generally in the Neotropical passalids, the hot-spot characteristics, with diversity and endemism, of the narrow land between the Depression of Nicaragua and the Isthmus of Panama.     

Revision of the species of the Tychus rufus group (Coleoptera: Staphylinidae: Pselaphinae)

The species belonging to Tychus rufus group are revised. Eleven species are recognized, described and illustrated and a key to their identification is provided. Nine taxa are new to science: Tychus carpathius n. sp. from Karpathos island (Greece); T. torticornis n. sp. from Lesbos Island (Greece); T. pisidicus n. sp. and T. inermis n. sp. from southwestern Turkey; T. antiocheus n. sp. and T. effeminatus n. sp. from southeastern Turkey; T. artvinensis n. sp. from northeastern Turkey, and T. sidonicus n. sp. and T. libanus n. sp. from Lebanon.     

Six new species of the subgenera Alloparadisa and Ashima (Diptera: Drosophilidae: Phortica) from Yunnan,China, with DNA barcoding information

Six new species of the genus Phortica are described from Yunnan, Southwest China: P. (Alloparadisa) kaibangya n. sp., P. (Ashima) semiannulata n. sp., P. (Ashima) dianmianensis n. sp., P. (Ashima) idiasta n. sp., P. (Ashima) menglian n. sp. and P. (Ashima) zhangyuanae n. sp. The 22 new DNA sequences of the mitochondrial COI gene with BOLD Process ID and GenBank accession numbers are provided for the six new species and six related known species from southern China: P. (Ashima) glabra, Chen & Toda, 2005 P. (Ashima) nudiarista Cheng & Chen, 2008, P. (Ashima) saltiaristula Chen & Wen, 2005, P. (Ashima) spinosa, Chen & Toda, 2005 P. (Ashima) symmetria Chen & Toda, 2005 and P. (Ashima) yiqini Zhu & Chen, 2017. The pairwise interspecific K-2P COI distances among all Chinese species of the subgenera Alloparadisa and Ashima are summarized. An identification key to males of all Chinese species of the subgenera Alloparadisa and Ashima is provided.     

Isotomiella (Isotomidae: Collembola) of China

Seven species of the genus Isotomiella Bagnall 1939 were collected in China, two of which from Hainan and Guangxi provinces (South China) are newly described and illustrated. Isotomiella longisensilla n. sp. has bidentate mucro and long sensilla on most tergites, a character never seen before in this genus. I. deharvengi n. sp. belongs to the minor group and is characterized by a pair of strongly curved sensilla on Abd. V.     

Two new species of the Empis (Leptempis) rustica group of the Mediterranean basin (Diptera: Empididae: Empidinae)

Two new species of the Empis (Leptempis) rustica group are described: E. (L.) tenuis n. sp. and E. (L.) lauriauti n. sp. Both were mainly collected in mountainous areas of the Mediterranean basin; the first species was collected in Morocco and Southern Spain; the latter was found in late summer in the Southern Alps in France. The key to species of the Empis (L.) rustica group is updated to include these two new species as well as a recently described Caucasian species.     

New Hypogastrura species and new distributional data for Hypogastrura distincta (Axelson 1902) from China (Collembola: Poduromorpha: Hypogastruridae)

Two new species, Hypogastrura analpapillata n. sp., on mushrooms from Yunnan Province, and Hypogastrura microspinata n. sp., on moss from Hunan Province, are described. Hypogastrura analpapillata n. sp. is the sixth species with highly modified mouthparts in the trybomi group. Hypogastrura microspinata n. sp. is close to H. yosii Stach 1964, but can be differentiated from the latter by body granules, tenent hair and lateral teeth on tibiotarsus. A key to Chinese species of Hypogastrura is provided and a list of Chinese Hypogastrura species and their distributional data are given.     

Revision of the western Palaearctic Mesoleptus (Hymenoptera: Ichneumonidae)

The western Palaearctic species of the genus Mesoleptus Gravenhorst 1829 are redescribed and keys are given for their identification. Also a key for the identification of the three stilpnine genera (Atractodes, Mesoleptus, and Stilpnus) is given and the biology of the genus is discussed shortly. The following species are described as new: Mesoleptus hispanicus n. sp., M. sawoniewiczi n. sp. and M. tunisiensis n. sp. The male of Mesoleptus tobiasi Jonaitis is described for the first time. New combinations is M. incessor (Haliday). New synonyms number 143 and 34 lectotypes are chosen.     

A revision of the Chinese Catops Paykull 1798 of the Catops fuscus species group (Coleoptera: Leiodidae: Cholevinae)

Five species of the Catops fuscus species group are reported from China. Three species are newly described: C. hlisnikovskyi n. sp. from the Beijing municipality and Jiangsu province, C. schuelkei n. sp. from Sichuan and Yunnan provinces, and C. smetanai n. sp. from Sichuan and Yunnan provinces. Female of C. sasajii Nishikawa 2007 is described and the species is reported from Hubei, Sichuan and Yunnan provinces for the first time. Catops nigricans (Spence 1813) is reported for the first time from China (Xinjiang Uygur autonomous region). Important morphological characters are illustrated and the distribution of all species in China is mapped. Preliminar phylogenetic analysis separates (C. hlisnikovskyi n. sp.+C. fuscus fuscus Panzer 1794) as a sister clade to C. nigricans+(C. sasajii+(C. schuelkei n. sp.+C. smetanai n. sp.)), with C. paramericanus Peck & Cook 2002 as outgroup.     

Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae)

A new subgenus, Huaius n. subgen. of the genus Pterostichus, is described to deal with the following three new species of ground beetles which were recently discovered from limestone caves in Central China: Pterostichus (Huaius) caoi n. sp. (Cave Lijia Dong in Yichang, Hubei Province), Pterostichus (Huaius) tiankeng n. sp. (Cave Xigou Dong in Hanzhong, Shaanxi Province) and Pterostichus (Huaius) yuae n. sp. (Cave Didonghe Dong in Hanzhong, Shaanxi Province). This new subgenus is not closely allied to any lineage of Chinese Pterostichus. The pronotal structure of Huaius has the most peculiar morphological characteristics, lacking lateromarginal borders and without grooves on the portion of basal foveae. All members of Huaius are de-pigmented, almost blind and extremely troglomorphic. This is the first report of cave-adapted pterostichine beetles from Asia.     

Phylogeny and systematics of the Acrapex apicestriata (Bethune-Baker, 1911) species complex (Lepidoptera,Noctuidae, Noctuinae,Apameini, Sesamiina) with the description of eight new species from the Afrotropics

Twelve morphologically similar species of Acrapex Hampson 1894, (Lepidoptera, Noctuidae, Noctuinae, Apameini, Sesamiina), from Western, Central and Eastern Africa are reviewed. Eight of these species are new to science and are described: Acrapex akunamatata n. sp. and A. incrassata n. sp. from Kenya; A. gracilis n. sp., A. iringa n. sp., A. lukumbura n. sp. and A. rungwe n. sp. from Tanzania; A. soyema n. sp. from Ethiopia; and A. zoutoi n. sp. from Benin. All 12 species belong to a species complex that we hereby define as the Acrapex apicestriata group. Host-plants for three of the new species are recorded: Setaria incrassata (Hochst.) Hack. for Acrapex incrassata; Cymbopogon pospishilii (K. Schum.) C.E. Hubb. for A. rungwe; and Andropogon perligulatus Stapf. for A. zoutoi. We also conducted molecular phylogenetic analyses (using maximum likelihood and Bayesian inference) on a six gene multimarker molecular dataset (four mitochondrial and two nuclear gene fragments; 4581 nucleotides in length) consisting of 15 Acrapex species (including seven species from the apicestriata group) and four outgroups species from the subtribe Sesamiina (from genera Busseola Thurau 1904, Sciomesa Tams & Bowden 1953, Pirateolea Moyal, Le Ru, Conlong, Cugala, Defabachew, Matama-Kauma, Pallangyo & Van den Berg 2010 and Sesamia Boisduval & Guenée 1852). Both maximum likelihood and Bayesian inference analyses yield a similar and well-supported topology, which supports the monophyly of the apicestriata group.     

Four new species of Cerambycidae (Coleoptera) from Brazil and updated identification keys

Four new species of longhorn beetles are described from Brazil: Coleoxestia diamantina n. sp. (Cerambycinae, Cerambycini), from Bahia; Mirador bravoi n. sp. (Cerambycinae, Ectenessini), from Bahia; Compsibidion antonietae n. sp. (Cerambycinae, Neoibidionini), from Goiás, Bahia and São Paulo; and Amphicnaeia quadrifasciata n. sp. (Lamiinae, Apomecynini) from Bahia. Coleoxestia diamantina and Compsibidion antonietae are included in previous keys.     

New and earlier unnoted mealybugs and felt scale (Homoptera: Coccinea: Pseudococcidae,Eriococcidae) from Morocco

Two new species of mealybugs (Pseudococcidae), Puto (Ceroputo) chetverikovi n. sp. and Fonscolombia toubkalensis n. sp., are described and illustrated. Thirteen other species are noted for the first time to fauna of Morocco or to Africa as a whole. Additionally, one species of closely related family Eriococcidae, Acanthococcus ericae (Signoret, 1875), is also reported for the first time to Morocco.     

A remarkable new Veturius (Veturius) of the Brazilian Central Amazonas,with an inventory of the genus in Brazil (Coleoptera: Passalidae)

A new species of Veturius (Veturius) Kaup, of Brazil, Central Amazonas, lower basin of Urucu, Tefé and Coari, is described and illustrated. V. urucuensis n. sp. belongs to the “cephalotes” species group, and has an external autapomorphy for the family Passalidae: a large tuft of setae on the medioposterior area of the metasternum. The species is closely allied to V. jolyi Boucher, 2006, from the Brazil-Venezuela-Guyana border. The peculiar endemism of the new species suggests an intra-continental island vicariance into the Amazon Basin. No similar process was known in the large genus Veturius. A complete inventory of the 22 Veturius species in Brazil is given, with their distribution, and following the previously established monophyletic species groups. From Brazil, two species are newly reported, V. (Ouayana) guyanensis Boucher, 2006 (Pará) and V. (O.) dominicae Boucher, 2006 (Roraima), and one species is confirmed, V. (O.) unicornis Gravely, 1918 (Amazonas). New details are given on some other Brazilian Veturius.