Scleritome construction,biofacies, biostratigraphy and systematics of the tommotiid Eccentrotheca helenia sp. nov. from the Early Cambrian of South Australia

Abstract: Large collections of Eccentrotheca helenia sp. nov. from the lower Cambrian Wilkawillina and Ajax limestones in the Arrowie Basin, South Australia, contain abundant low, cap‐shaped and high, laterally compressed isolated sclerites in addition to partially articulated tubular specimens. The scleritome of Eccentrotheca helenia sp. nov. is fully described for the first time and shown to be formed by ontogenetic fusion of sclerites into successively stacked sclerite rings, forming a larger, tubular structure. The apical termination of the tube is highly variable, but is primarily constructed by low, cap‐shaped sclerites and characterised by a central aperture of variable inclination. The adapical portion of the tube is predominantly constructed by high, laterally compressed sclerites, but individual sclerite rings can contain both cap‐shaped and laterally compressed sclerites along with sclerites of intermediate morphology. The apical aperture presumably housed organic structures for attachment to a hard substrate, but the scleritome also occasionally preserves small lateral perforations between fused sclerites, which may have served to stabilise the scleritome by providing additional points of anchorage. In the Arrowie Basin, E. helenia is found in association with archaeocyath‐microbial‐spongiomorph‐dominated bioherms and most likely inhabited pendant or cryptic habitats within these bioherms. Eccentrotheca‐like sclerites form an integral part of the scleritomes of many tommotiids which may confuse taxonomic analysis. Sclerites previously assigned to ‘E.’guano, consistently occur together with sclerites of Kulparina rostrata in stratigraphic intervals consistently older than strata hosting E. helenia. Rare fused specimens indicate that the sclerites of K. rostrata and ‘E.’guano belong to the same scleritome.     

Miocene ostracods from the Tremiti Islands and Hyblean Plateau: biostratigraphy and description of new and poorly known species

The main taxonomic and stratigraphical results of a study of ostracods from the middle-late Miocene of the Tremiti Islands and the Hyblean Plateau are presented. The faunas are mostly typical of deep thermospheric or psychrospheric bathyal environments. Excluding a few, mostly juvenile shallow water contaminants, a total of 127 species were recorded. Twenty-four taxa are discussed herein; of these, 13 species (Cytherella dissimilis nov. sp., Cytherella parvula nov. sp., Cytherella subtilis nov. sp., Argilloecia triangularis nov. sp., Retibythere (Retibythere)? claudii nov. sp., Retibythere (Retibythere)? gibba nov. sp., Cytheropteron (Cytheropteron) joachinoi nov. sp., Cytheropteron (Aversovalva) consuetum nov. sp., Cytheropteron (Aversovalva) trimerumense nov. sp., Cytheropteron (Aversovalva) turgidum nov. sp., Loxoconcha julianii nov. sp., Buntonia whatleyi nov. sp., Pterygocythere nuda nov. sp.) and a subspecies (Costa tricostata dorsoarcuata nov. subsp.) are described as new, while six are left in open nomenclature. The stratigraphical distribution of ostracods is expressed in terms of planktonic foraminiferal zones. Information obtained from this study enriches biostratigraphical data on Mediterranean Miocene deep-sea ostracods, especially with respect to the Langhian.     

Palynology of the Dockum group (Upper Triassic), Texas,U.S.A.

A palynofloral assemblage is described from the Upper Triassic Dockum Group of Texas. Forty palynomorph species are recorded and these are correlated with European Upper Triassic assemblages. The Dockum Group assemblage is considered to be of late Karnian age.Two new genera, Tulesporites gen. nov., type species T. briscoensis sp. nov., and Daughertyspora gen. nov., type species D. chinleanus (Daugherty) comb. nov. have been erected; new species are Converrucosisporites matsenii sp. nov., Pyramidosporites traversei sp. nov., Callialasporites triassicus sp. nov., Plicatipollenites minutotriletus sp. nov., Triadispora dockumensis sp. nov., Bhardwajispora jansonii sp. nov., Protodiploxypinus americus sp. nov., P. ujhelyi sp. nov., Klausipollenites gouldii sp. nov., Pityosporites old-hamensis sp. nov., Abietineaepollenites bujakii sp. nov., Falcisporites oviformis sp. nov. and F. tecovasensis sp. nov.     

A new gorgonian genus from deep-sea Antarctic waters (Octocorallia, Alcyonacea, Plexauridae)

Mesogligorgia scotiae gen. nov., sp. nov. is described and illustrated from a colony collected in the Scotia Sea, 2,201–2,213 m in depth, on the ANDEEP-I cruise. The new taxon is placed in the family Plexauridae because of: 1) the presence of a horny axis with a cross-chambered central core and numerous loculi, 2) retractile polyps in calyces with distinct spicular components, and 3) armed polyps with large sclerites with a poorly- developed collaret and eight well-developed points. The irregularly distributed sclerites running along the axis, into a thick mesogloeal coenenchyme, and the elongated spindles with irregular ends are the most distinctive characters of the newly proposed genus.     

Systematics and taphonomy of an Early Kimmeridgian belemnite fauna from the Mediterranean Tethys (Monte Nerone, Central Apennines, Italy)

About three hundred belemnite rostra were collected from lower Kimmeridgian beds of a structural high sequence cropping out at Mt. Nerone (Central Apennines, Pesaro Province, Italy). The belemnite fauna is composed mainly of new species. Nine species were recognised, ascribed to five genera, that include Hibolithes semisulcatus M?NSTER, 1830; H. pignattii nov. sp.; Acutibelus sp. cf. acuariformis RIEGRAF, 1981; Belemnopsis neronensis nov. sp., Duvalia matteuccii nov. sp., D. nicosiai nov. sp., D. pallinii nov. sp., D. raymondi nov. sp. and Rhopaloteuthis massimoi nov. sp.; moreover a single specimen is treated in open nomenclature as Belemnopseidae incertae sedis. The stratigraphic and palaeobiogeographic significance of the new fauna is discussed. The taphonomy of the belemnite-rich level is described, with reference to borings found on the belemnite rostra.     

Genus Thubana Walker of the Malaysian Borneo (Lepidoptera,Lecithoceridae), with Descriptions of Seven New Species

The genus Thubana Walker, 1864 of the Malaysian Borneo is revised. Nine species are recognized including seven new species. The new species are: T. kumatai sp. nov., T. kubahensis sp. nov., T. kinabaluensis sp. nov., T. ochracea sp. nov., T. quadrata sp. nov., T. prolata sp. nov., and T. spinata sp. nov. With regard to the previously known species, photos of adults and the male or female genitalia are given, with some taxonomie notes. A key to the Thubana species of the Malaysian Borneo is provided.     

New radiolarian species of the order Nassellaria from the Coniacian-Santonian of the Perapedhi formation of southern Cyprus

Rich radiolarian associations are investigated. They include nine new species from the Upper Cretaceous (Coniacian-Santonian) metal-bearing beds of the Perapedhia Formation of southern Cyprus: Dorypyle concinna sp. nov., Gongylothorax mangaleniensis sp. nov., Heliocryptocapsa macroporus sp. nov., H. perapediensis sp. nov., Mita guttiformis sp. nov., Pseudoeucyrtis belogorskensis sp. nov., P. cypricus sp. nov., Rhopalosyringium mangaleniense sp. nov., and Sandovalella trihedra sp. nov.     

Three new species of genus Myrmica (Hymenoptera: Formicidae) from Himalaya

Three new species of the genus Myrmica are described from the Himalayas. Myrmica curvispinosa sp. nov., Myrmica kothiensis sp. nov. and Myrmica religiosa sp. nov. belong to the Myrmica inezae species group, which is earlier represented by 4 species. Myrmica curvispinosa sp. nov. is described based on worker and gyne, with a report of ergatoid as well. Myrmica kothiensis sp. nov. and Myrmica religiosa sp. nov. are described based on worker caste only. A key to the species of the Myrmica inezae species group has been provided in the following.     

Jurassic Scaphopoda from the Russian platform: Pulsellidae and Gadilidae

The scaphopod families Pulsellidae and Gadilidae from sandy and clayey beds of the Jurassic (Callovian?Volgian) of European Russia are investigated. In Pulsellidae, the following taxa are described: Annulipulsellum with two subgenera, Annulipulsellum (species A. ambiguum sp. nov., A. rugosum sp. nov.) and Mesopulsellum subgen. nov. (A. alternoides sp. nov., A. calloviense sp. nov., A. hirtistriatum sp. nov., A. medium sp. nov.), Gracilipulsellum gen. nov. (G. iodaense sp. nov.), Gardneridentalium gen. nov. (G. primitivum sp. nov., G. reticulatum sp. nov.), and Mesoentalina gen. nov. (M. fabulosa sp. nov.). New taxa of the family Gadilidae include the genus Mesoantalis gen. nov. with the species M. clava sp. nov., M. expolitum sp. nov., and M. volgense sp. nov.     

Sélaciens nouveaux du London Clay de l'Essex (Yprésien du bassin de Londres)

The Ypresian locality of Burnham-on-Crouch(Essex) has yielded numerous Selachian species, some of which are new: Hexanchus agassizi sp. nov., Scyliorhinus burnhamensis sp. nov., S. casieri sp. nov., S. pattersoni sp. nov., S. woodwardi sp. nov., Pararhincodon ypresiensis sp. nov., Triakis wardi sp. nov., Mustelus whitei sp. nov. The Hexanchid teeth classicaly refered to Notidanus serratissimusAg. include in fact two separate species belonging to different genera: Notorhynchus serratissimus (Ag.) and Hexanchus agassizi sp. nov. Lamna affinis (Casier) is arranged in the genus Isurolamna nov. in consideration of his dental peculiarities. The genus Burnhamia nov. is created for Rhinoptera daviesiWoodward that can be regarded as a Mobulid. Oxyrhina sheppeyensisCasier must be classified in the genus Anomotodon. The genus Triakis, Mustelus, Anomotodon and Pararhincodon are pointed out for the first time in the Ypresian.     

Systematics and taphonomy of an Early Kimmeridgian belemnite fauna from the Mediterranean Tethys (Monte Nerone, Central Apennines, Italy)

About three hundred belemnite rostra were collected from Lower Kimmeridgian beds of a structural high sequence cropping out at Mt. Nerone (central Apennines, Pesaro Province, Italy). The belemnite fauna is composed mainly of new species. Nine species were recognised, ascribed to five genera, which include Hibolithes semisulcatusMünster, 1830; H. pignattii nov. sp.; Acutibelu0s sp. cf. acuariformisRiegraf, 1981; Belemnopsis neronensis nov. sp., Duvalia matteuccii nov. sp., D. nicosiai nov. sp., D. pallinii nov. sp., D. raymondi nov. sp. and Rhopaloteuthis massimoi nov. sp.; moreover a single specimen is treated in open nomenclature as Belemnopseidae incertae sedis. The stratigraphic and palaeobiogeographic significance of the new fauna is discussed. The taphonomy of the belemnite-rich level is described, with reference to borings found on the belemnite rostra.La fauna a belemniti risulta composta quasi esclusivamente da specie nuove. Sono state riconosciute le seguenti specie: Hibolithes semisulcatusMünster, 1830; H. pignattii nov. sp.; Acutibelus sp. cf. acuariformisRiegraf, 1981, Belemnopsis neronensis nov. sp., Duvalia matteuccii nov. sp., D. nicosiai nov. sp., D. pallinii nov. sp., D. raymondi nov. sp. and Rhopaloteuthis massimoi nov. sp. Inoltre un esemplare è stato discusso con nomenclatura aperta come Belemnopseidae incertae sedis. Alcuni nuovi dati stratigrafici e paleobiogeografici sono stati analizzati ed integrati con i dati della letteratura. E’ stata anche esaminata la tafonomia del livello a belemniti e le perforazioni presenti sulla superficie dei rostri.     

New genera and species of coelacanths from the Bear Gulch Limestone (Lower Carboniferous) of Montana (U.S.A.)

Four new species of coelacanths (Actinistia) from the Namurian E2b marine Bear Gulch Limestone of Montana are described, and a fifth species, originally described as an actinopterygian, is redescribed. Two new suborders are diagnosed. The taxonomy is: Suborder Coelacanthocdei, family Rhabdodermatidae, Caridosuctor populosum nov. sp., suborder Hadronectoroidei nov., family Hadronectoridae nov., Hadronector donbairdi nov. sp., Polyosteorhynchus simplex nov. sp. and Allenypterus montanusMelton. Lochmocercus aciculiodontus nov. sp. is incertae sedis.     

A new subfamily of ichneumonids from the Lower Cretaceous of Transbaikalia and Mongolia (Insecta: Hymenoptera: Ichneumonidae)

A new subfamily of ichneumonids, Palaeoichneumoninae, is described from the Lower Cretaceous of Transbaikalia (Baisa locality) and Mongolia (Bon Tsagan and Kholbotu Gol localities). The new subfamily is intermediate between the archaic subfamily Tanychorinae and the Recent Ichneumonidae. It includes 12 new species, described in three new genera: Palaeoichneumon freja gen. et sp. nov., P. ornatus sp. nov., P. danu sp. nov., P. micron sp. nov., P. mirabilis sp. nov., P. tenebrosus sp. nov., P. townesi sp. nov., Rudimentifera mora gen. et sp. nov., R. suspecta sp. nov., Dischysma maculata gen. et sp. nov., D. similis sp. nov., and D. ramulata sp. nov.     

New radiolarian species from the Upper Cretaceous sections of the Crimean Mountains (Ukraine)

New radiolarians from the Upper Cretaceous beds (Middle Turonian-Upper Santonian) of four sections of the Crimean Mountains (Biuk-Karasu River, western peak of Ak Mountain, Ak-Kaya Mountain, and Chuku Mountain) are investigated. Seven new species are described, including Actinomma (?) belbekense sp. nov., Pseudoaulophacus kopaevichae sp. nov., Ps. trizonalis sp. nov., Phaseliforma turovi sp. nov., Triactoma karasuensis sp. nov., Patulibracchium belogorskensis sp. nov., and Paronaella (?) nikishini sp. nov.     

Discosorids and Oncocerids (Cephalopoda) of the Middle Ordovician Kunda and Aseri Regional Stages of Baltoscandia and the early evolution of these groups

Discosorid and oncocerid cephalopods of the Kunda and Aseri Regional Stage (Darriwilian, Middle Ordovician) from Öland Island, Sweden and North Estonia are described for the first time. We demonstrate that a high generic and specific diversity of Oncocerida was already established in the earliest Darriwilian. The new oncocerid species Neumatoceras borense nov. sp., N. breviborense nov. sp., Paramiamoceras breviventrum nov. sp., Richardsonoceras gastroscopium nov. sp., R. gerhardi nov. sp., R. goldmanni nov. sp., R. haelluddenense nov. sp., Richardsonoceroides kingi nov. sp., R. oelandense nov. sp., and R. rhytium nov. sp. are erected; Richardsonoceroides schiefferdeckeri (Dewitz) is revised. The new graciloceratid Kundoceras evansi nov. gen., nov. sp. is erected. A review of records of early oncocerids from China shows that Richardsonoceras Foerste is probably the most primitive oncocerid and supports the hypothesis that oncocerids evolved in the early Floian or latest Tremadocian from a Bassleroceras Ulrich and Foerste-like ancestor. The new discosorid genus Paldoceras nov. gen. and species Paldoceras paldiskense nov. sp. and P. neptunsakerense nov. sp. are erected. These discosorids record the morphological link between the late Darriwilian Ruedemannoceras Flower and the early Floian Apocrinoceras Teichert and Glenister, which is considered to be the earliest discosorid.     

Ten new species of caddisflies (Insecta: Trichoptera) from the Baltic amber

Ten new species of caddisflies of the families Polycentropodidae (nine) and Molannidae (one) are described from the Eocene Baltic amber: Holocentropus peregrinator sp. nov., H. proximorepertus sp. nov., H. dugoi sp. nov., H. fundamentalis sp. nov., H. telergon sp. nov., Plectrocnemia varigoria sp. nov., P. synthesia sp. nov., P. novokshonovi sp. nov., Nyctiophylax valideturbidus sp. nov., and Molanna okraina sp. nov.     

New ancyloceratidae (Ammonoidea) from the Aptian of the Ulyanovsk region,Middle Volga

A new genus Pseudoancyloceras gen. nov. with the type species P. calinovense sp. nov. and five new species of Ancyloceratidae (Australiceras (Proaustraliceras) elegans sp. nov., Audouliceras vyrystaykense sp. nov., A. brusyankense sp. nov., A. mordovense sp. nov., Tropaeum (Tropaeum) elaurkense sp. nov.) are described from the Aptian of the Ulyanovsk region, Middle Volga.     

Triassic radiolarians of Kotel’nyi Island (New Siberian Islands,Arctic)

Triassic radiolarians from Kotel’nyi Island (New Siberian Islands, Arctic Region) are studied. Radiolarians occur in the Middle and Upper Triassic beds, which are well dated by ammonoids, nautiloids, and bivalves. In the Middle Triassic beds, which are composed of dark gray and black clays and claystones with interbeds of combustible schist, siltstone, clayey limestone, and many phosphatic concretions, the following two assemblages are recognized: (1) a Late Anisian assemblage, including Glomeropyle clavatum Bragin, sp. nov., G. boreale Bragin, G. manihepuaensis Aita, G. insulanum Bragin, sp. nov., Triassospongosphaera multispinosa (Kozur et Mostler), Tetraspongodiscus hibernus Bragin, sp. nov., T. borealis Bragin, sp. nov., Eptingium abditum Bragin, sp. nov., E. manfredi Dumitrica, Spongostephanidium japonicum (Nakaseko et Nishimura), and Ladinocampe vicentinensis Kozur et Mostler; (2) Late Ladinian assemblage, with Muelleritortis firma (Gorican), M. kotelnyensis Bragin, sp. nov., Tiborella nivea Bragin, sp. nov., Pseudostylosphaera goestlingensis (Kozur et Mostler), P. omolonica Bragin, Triassospongosphaera multispinosa (Kozur et Mostler), Sarla cincinnata Bragin, sp. nov., S. obscura Bragin, sp. nov., S. prava Bragin, sp. nov., and Eonapora stiriaca Bragin, sp. nov. The Upper Triassic beds, composed of gray and black clays and claystones with lenses of clayey limestones, with sideritic and phosphatic-calciferous concretions have yielded (1) an Early Carnian assemblage, with Pentactinocarpus colum Bragin, sp. nov., Glomeropyle cuneum Bragin, sp. nov., G. algidum Bragin, sp. nov., G. aculeatum Bragin, sp. nov., Kahlerosphaera isopleura Bragin, sp. nov., Stauracanthocircus folium Bragin, sp. nov., Tetraspongodiscus uncatus Bragin, sp. nov., Poulpus costatus (Kozur et Mostler), Eonapora robusta Kozur et Mostler, Planispinocyrtis kotelnyensis Bragin, sp. nov., Annulotriassocampe baldii (Kozur), and Pseudoeucyrtis annosus Bragin, sp. nov.; (2) a Middle-Late Carnian assemblage with Pseudostylosphaera glabella Bragin, sp. nov., P. gracilis Kozur et Mock, P. voluta Bragin, sp. nov., P. gelida Bragin, sp. nov., Kahlerosphaera unca Bragin, sp. nov., K. aspinosa Kozur et Mock, K. fuscinula Bragin, sp. nov., K. acris Bragin, sp. nov., Capnuchosphaera kuzmichevi Bragin, sp. nov., C. triassica De Wever, C. angusta Bragin, sp. nov., Sarla intorta Bragin, sp. nov., S. compressa Bragin, sp. nov., S. aequipeda Bragin, sp. nov., Betraccium irregulare Bragin, B. kotelnyensis Bragin, sp. nov., Spongotortilispinus carnicus Kozur et Mostler, S. subtilis Bragin, sp. nov., Dumitricasphaera simplex Tekin, D. aberrata Bragin, sp. nov., D. arbustiva Bragin, sp. nov., Zhamojdasphaera epipeda Bragin, sp. nov., Z. proceruspinosa Kozur et Mostler, Vinassaspongus subsphaericus Kozur et Mostler, Palaeosaturnalis triassicus Kozur et Mostler, Paronaella concreta Bragin, sp. nov., P. aquilonia Bragin, sp. nov., Tetraspongodiscus cincinnalis Bragin, sp. nov., Annulotriassocampe baldii (Kozur), Canoptum zetangense Wang et Yang, Whalenella speciosa (Blome), Syringocapsa turgida Blome, Droltus gelidus Bragin, sp. nov., and D. niveus Bragin, sp. nov.; (3) an Early Norian assemblage with Pseudostylosphaera glabella Bragin, sp. nov., P. gelida Bragin, sp. nov., P. voluta Bragin, sp. nov., Kahlerosphaera retunsa Bragin, sp. nov., Capnuchosphaera deweveri Kozur et Mostler, Sarla globosa Bragin, sp. nov., Palaeosaturnalis mocki Kozur et Mostler, Paronaella aquilonia Bragin, sp. nov., Syringocapsa turgida Blome, and Droltus gelidus Bragin, sp. nov.; (4) a Middle Norian assemblage with Sarla globosa Bragin, sp. nov., Sarla sp., Syringocapsa turgida Blome, Canoptum sp., and Laxtorum? sp. The fact that these assemblages include taxa that are recorded more southerly, including paleotropical localities, and support dating based on mollusks, enables the use of these data for Boreal-Tethyan correlation. The greatest similarity to Tethyan associations is observed in the Late Carnian and Early Norian, that is, the periods of the greatest penetration of thermophilic mollusks into this basin. A total of 69 radiolarian species of 3 orders, 18 families, and 29 genera are described; 44 species are newly described; the stratigraphic and geographical ranges of the majority of taxa are improved considerably.