塔里木盆地二叠系孢粉组合及生物地层学*

首次系统地研究了塔里木盆地二叠系孢子花粉，讨论了盆地各地层分区孢粉组合特征，建立了从早二叠世至早三叠世早期的6个孢粉组合带，阐述了各组合带孢粉学特征。依据对盆地二叠纪孢粉植物群组成分子的分析及其与国内外相关孢粉组合的比较研究，将盆地孢粉植物群分为3个发展阶段，即：早二叠世早、中期的欧美型植物群，早二叠世晚期至晚二叠世早期的安加拉欧美混生型植物群和晚二叠世中、晚期的安加拉型植物群。文中还从孢粉地层学角度论述了沙井子组、杜瓦组和普司格组的地质时代以及二叠/三叠系界线等问题。描述孢粉25属33种（含2新种）。     

山东聊城地区阿城镇煤田二叠纪孢粉组合

对山东聊城阿城镇煤田A19-6、A20-6、L104钻孔岩石样品进行了分析,发现从太原组中上部到上石盒子组地层中均有大量孢粉化石保存,经分析鉴定和统计,共有孢粉化石53个属168种及部分未定种.根据孢粉属种的纵向分布情况及含量的变化,在本区二叠系建立了4个孢粉组合,自上而下为:Ⅳ.Triquitrtites microgranifer-Macrotorispora media组合带;Ⅲ.Limitisporires rhombicorpus -Patellisporites meishanenisis组合带;Ⅱ.Gulisporites cochlearius-Sinulatis porites sinensis组合带;Ⅰ.Thymospora pseudothiessenii-Laevogatisporites vulgaris组合带.通过与邻区及华北部分地区对比,确定本区孢粉组合所在的地质时代应为:太原组中上部,早二叠世早期(组合Ⅰ);山西组,早二叠世中期(组合Ⅱ);下石盒子组,早二叠世晚期(组合Ⅲ);上石盒子组,晚二叠世早期(组合Ⅳ).     

徐州王庄煤矿山西组孢粉植物群及其地层意义

徐州王庄煤矿山西组中、下部产有丰富的孢粉化石,保存完好,共计55属132种（包括3新种）2个未定型孢子及1个疑源类未定类型。根据孢粉属种组成及其含量的变化,自下而上分为两套组合：Cyclogranisporites-Gulisporites-Laevigatosporites-Florinites组合（简称组合1）和Calamospora-Gulisporites-Vesiculatisporites-Florinites组合（简称组合2）。根据组合内一些重要分子的地质时限讨论及其与相关组合的对比,将组合1的地质时代归为早二叠世早期,组合2的时代归为早二叠世晚期。文中描述孢子3新种,图示孢子花粉91种。     

山东济阳坳陷二叠纪孢粉组合

为了划分对比地层,确定地层层位,对济阳地区油气勘探钻孔曲古2、曲古3、义136中的孢粉样品进行了分析,其中5块样品含有孢粉化石,分别是曲古23(2843.2m)、义1365(3772.6m)、曲古334(4059m)、曲古323(4041m)和曲古313(4013m)。通过分析鉴定,共鉴定出59属,119种和未定种,根据样品中孢粉成分和含量的变化,建立了如下三个孢粉组合,自下而上为:Ⅰ.Cyclogranisporites Granulatisporites Laevigatosporites组合(山西组下部),Ⅱ.Punctatisporites Cyclogranisporites Florinites组合(山西组上部),Ⅲ.Macrotorispora Vesicaspo ra Pteruchipollenites组合(上石盒子组)。通过与华北、华南部分地区对比,确定各孢粉组合的地质时代为早二叠世中期(组合Ⅰ和组合Ⅱ),晚二叠世早期(组合Ⅲ)。由于含孢粉样品较少,层位不连续,其中下石盒子组孢粉组合未建立,有待于进一步研究。     

新疆准噶尔盆地东北缘孔雀坪剖面二叠纪金沟组孢粉组合及其地层意义

准噶尔盆地东北缘孔雀坪剖面金沟组孢粉组合中Protohaploxypinus,Striatoabieites等具肋双气囊花粉非常发育,类型丰富,达7属32种之多;Cordaitina,Zonalasporites等单气囊花粉分异度较高,但含量较低;少量出现单沟花粉、具肋纹花粉、无肋双气囊花粉和蕨类植物孢子等。所见孢粉类型中约1/2的已知种在新疆北部只见于二叠纪地层中,1/3已知种在晚石炭世至二叠纪地层中均可见到。孢粉组合特征与盆地南缘下二叠统塔什库拉组上部组合相似,地质时代为早二叠世空谷期。     

河南义马地区晚三叠世—侏罗纪孢粉组合及其地质意义

重点讨论了谭庄组上段孢粉组合（包括１新种和１新组合）；简要描述了侏罗纪义马组砾岩段孢粉组合。谭庄组上段的孢粉组合主要以具双气囊的松柏类花粉为优势（占孢粉总数的６５％），同时又具许多晚三叠世的常见分子和孑遗分子Ａｒａｔｒｉｓｐｏｒｉｔｅｓ，Ｃｕｎｅｉｓｐｏｒｉｔｅｓ，Ｔａｅｎｉａｓｐｏｒｉｔｅｓ。该组合可与中国北方及欧洲晚三叠世孢粉组合进行对比。义马组砾岩段的孢粉组合系首次发现，以蕨类孢子为主（占孢粉总数６２．９％），裸子植物花粉次之（占孢粉总数３７％）。其中常见的类型为Ｃｌａｓｓｏｐｏｌｌｉｓ，其时代为早侏罗世。     

鲁西地区二叠系上、下石盒子组孢粉组合及其分界

为解决鲁西地区上、下石盒子组分界问题,在相关地层共采集处理孢粉样品100多块,经系统研究建立本区上、下石盒子组两个孢粉组合,自下而上为：1)Sinulatisporites sinensis-Patellisporites meishanensis组合;2)Anticapipollis reticor pus-Macrotorispora gigantea组合。根据所建立的孢粉组合特征,经与华北各地二叠系孢粉组合广泛对比,进一步阐述各孢粉组合的地质时代,并为本区二叠系上、下石盒子组的分界提出了较为可靠的意见。该界线与区内中期华夏植物群晚期和晚期华夏植物群早期植物群界线基本一致。     

江苏句容上二叠统龙潭组孢粉组合及孢粉热变指数和有机质类型

通过对苏南句容A10井的孢粉研究,首次发现了上二叠统龙潭组孢粉化石。根据孢粉组合特征。与国内同时期的孢粉组合进行了比较和讨论,并用孢粉热变指数和有机质类型,对龙潭组进行了油气评价。为苏南地区晚二叠世的地层对比,提供了新的古生物依据。     

塔里木盆地二叠纪孢粉植物群区系性质及塔里木地块演化探讨

塔里木盆地二叠系孢粉分为五个孢粉组合带,即：ＳＱ、ＰＶ、ＡＶ、ＬＰ和ＰＧ带。据对盆地孢粉植物群性质的讨论及其与国内外相关孢粉植物群的对比研究,本文将塔里木地块二叠纪孢粉植物群的演化进程分为三个阶段,即：早二叠世早、中期的欧美区型植物群,早二叠世晚期至晚二叠世早期的安加拉区－欧美区混生型植物群和晚二叠世中、晚期的安加拉区型植物群。古地磁研究结果支持这一推论,即孢粉植物群的演化历史基本反映了塔里木地块的漂移、拼合进程。     

新疆吐哈盆地北缘黄山街组孢粉组合

新疆鄯照壁山沟口和柯柯亚剖面黄山街组的孢粉化石,经研究,共有52属77种。命名为Dictyophylli-dites-Aratrisporites-Parataeniaesporites组合（简称DAP组合）。根据组合主要分子的已知时代分布、含量高低变化,以及与有关已知时代孢粉组合进行对比,鉴定其地质时代为晚三叠世早中期。根据孢粉组合恢复古植被,反映当时的气候为温暖潮湿的温带类型。     

山东滕县煤田石炭纪、二叠纪孢粉植物群及古气候

根据前人在原位孢子方面已积累的大量资料,并参考现代孢子花粉形态研究成果,大致推测了本区石炭-二叠纪分散孢子花粉的大多数属的植物亲缘关系。通过已建立的含煤地层孢粉组合带推测相应时期的古植被类型,并与前人有关地质时期及地层的大植物化石带进行了对比,不仅验证了各地质时期植被的兴起、发展、繁盛以及衰退绝灭的历史过程,更大意义上将微体形态的孢子花粉与宏观上的植被进行了对比归类,从而使得对研究区晚古生代含煤地层的古植被的形态面貌有了更直观的了解。最后,利用孢粉分析恢复相应地质时期的古气候。     

Callistophytalean pteridosperms from Permian aged floras of China

Abstract: Recent investigations into Permian aged floras from China have highlighted the widespread occurrence of callistophytalean pteridosperms that challenge previous understanding of their spatial and temporal distribution and diversity. In China, the group spans the Permian period and constitutes a distinctive but rare component in many peat‐forming environments. The stratigraphically earliest callistophytalean occurs in the Asselian‐Sakmarian stages with fossils from the Taiyuan Formation of northern China including ovules of Callospermarion undulatum in coal ball assemblages, and ovulate fronds of Norinosperma shanxiensis and synangiate fronds of Norinotheca shanxiensis in adpression assemblages. More abundant in the fossil records are adpression remains from the Roadian‐Wordian stages with the Lower Shihhotse Formation preserving abundant vegetative and ovulate remains of Emplectopteris triangularis that is now considered to represent a callistophytalean. The youngest callistophytalean recognised is from the Wuchaipingian‐Changhsingian stages with the Xuanwei Formation of southern China containing a single stem of Callistophyton boyssetii that provides indisputable evidence of the group in the lead up to the end‐Permian mass extinction. These accounts are augmented by analysis of pollen records that demonstrate the callistophytalean pollen genus Vesicaspora to be widespread through palynological assemblages from the Permian period in both North and South China, including the Upper Shihhotse Formation, Shihchienfeng Group, Xuanwei Formation, and possibly also in the mid‐Pennsylvanian Benxi Formation. Although macrofossil specimens are uncommon elements in the assemblages that contain them, they demonstrate the continuity of callistophytalean pteridosperms from the Pennsylvanian sub‐period into the early Guadalupian epoch of the Permian in North China and into the Lopingian epoch of the Permian in South China. Of the species present, both Callistophyton boyssetii and Callospermarion undulatum are known from the Pennsylvanian–earliest Permian age floras of Euramerica, whereas Norinosperma, Norinotheca and Emplectopteris appear to represent endemic Cathaysian elements. Results imply that callistophytalean pteridosperms can no longer be excluded from theories of post‐Carboniferous plant evolution and floristics, appearing to have played an important role in both Permian and Carboniferous aged plant communities. The presence of Vesicaspora in several formations from which macro‐remains have not been identified is a hopeful indicator that further callistophytalean pteridosperms are yet to be found.     

Early and Middle Jurassic Sporo-Pollen Assemblages of Qiquanhu Coal-Field of Turpan Xinjiang

One hundred and thirteen species of spores and pollens belonging to 58 genera were found from 66 core samples taken from two boreholes of the Early and Middle Jurassic deposits in the Qiquanhu coal-field of the Turpan Basin, Xinjiang have been investigated in this paper. Two of the species are described as new. Three sporo-pollen zones from lower to upper are described as follows: Sporo-pollen zone Ⅰ (equivalent bed: Badaowan Formation) . The pteridophytic spores are slightly more abundant than the gymnospermous pollen in this assemblage. The most abundant spores are Cyathidites minor, C. australis and Undulatisporites pflugii, U. taenus and so on. The second important fern spores are Osmundacidites, Lycopodiumsporites, Apiculatisporis and Duplexisporites. The most important gymnospermous genus is Cycadopites which ranks highest in the assemblage and includes C. nitidus and C. typicus.The other gymnospermous pollen include Classopollis, Pinuspollenites, Protoconiferus Piceites. Besides, there are a few pollen grains of Taeniaesporites and Chordasporites surviving from the Late Triassic in this assemblage. The present assemblage may be compared with the Early Jurassic sporopollen assemblage from the Daling Formation of the North-East Guangxi. Therefore, the Badaowan Formtion may be referred to Early Jurassic in age, probably Early Lias. Sporo-pollen zone Ⅱ (equivalent bed: Sangonghe Formation) In this assemblage the gymnosperms are predominant.Of them, Podocarpidites and Pinuspollenites increase obviously, and Quadraeculina and Protoconiferus are common. Few surviving elements are observed. The most abundant spores are those of Cyathidites, and the particularly abundant pollens are those of Cycadopites, This assemblage may be compared with the Late Lias sporo-pollen assemblage from the Fuxian Formation of the Northern Shangganning Basin. Therefore, the Sangonghe Formation should belong to Early Jurassic, probably Late Lias in age. Sporo-pollen zone Ⅲ (equivalent bed: Xishanyao Formation). The contents of the spores and pollens are almost equal in this assemblage. Cyathidites and Cycadopites are the most important spores and pollen. Osmundacidites and Quadraeculina are common. Neoraistrickia, Lycopodiumsporites and Eucommiidites troedssonii are present. Ancient striate bisaccate pollen disappears. The characteristics of this assemblage show some resemblances to those of the Middle Jurassic Yanan Formation in Chongxin county of Gansu province. So the Xishanyao Formation should belong to Middle Jurassic.     

四川宣汉晚三叠世孢粉组合及其古植被与古气候意义

对四川盆地东北缘宣汉七里峡剖面须家河组进行了详细的孢粉学研究,发现丰富的孢粉化石,共计59属113种,其中包括蕨类植物孢子35属62种,裸子植物花粉22属47种,苔藓类植物孢子1属3种,以及藻类化石1属1种。据此建立一个孢粉组合,命名为Dictyophyllidites-Lunzisporites-Chasmatosporites组合(简称DLC组合),并进一步分为上、下两个亚组合。根据孢粉组合内一些重要分子的地质时代延限及与国内外有关孢粉组合的对比,提出其地质时代属于晚三叠世诺利期—瑞替期(Norian—Rhaetian)。该孢粉植物群反映了一个以蕨类植物的真蕨类、石松类和裸子植物的松柏类和苏铁类群等占主导且较为繁盛的植被面貌,揭示出温暖潮湿的亚热带—热带气候环境。     

滇西保山地区丁家寨组生物群的时代

首次系统描述保山地区丁家寨组的类化石,详细讨论丁家寨组所产类和腕足类动物群及孢粉植物群的时代和性质,提出丁家寨组的时代应与华南早二叠世的紫松阶相当,即相当于国际上的Ａｓｓｅｌｉａｎ阶至Ｓａｋｍａｒｉａｎ阶。确认丁家寨组不存在不同时代化石的再沉积混杂或大量化石时代倒置的异常现象。丁家寨组出现特提斯型Ｅｏｐａｒａｆｕｓｕｌｉｎａ动物类;腕足类组合属介于西澳区和泰马区之间的过渡动物群,西澳区的色彩较浓;     

The Paleocene-Eocene Palynoflora from the Tantou Basin in West Henan

On the basis of palynological data the Paleoeene-Eocene palynoflora from the Tantou Basin in west Itenan are divided into three sporo-pollen assemblages. They are given ill ascending order as follows: 1. The first assemblage (Dazhang Formation) characterized mainly by the predominance of palynoforms of Ulmaeeae anal various Proteaeeous pollen, being assigned to Late Paleoeene and indicating a dry-hot climate. 2. The second assemblage (Lower part of the Tantou Formation) marked by the presence of various Triporopollenites and Tricolporopollenites, belonging to the Early Eocene. A moist-hot climate is reflected. 3. The third assemblage (the Upper part of the Tantou Formation) represented by aboundant pollen grains of saeeated elements (consisting of Pinaeeae and Podocarpaeeae) and the occurrence of Aquilapollenites pollen which plays an important part in this assemblage. It is interpreted as belonging to the middle to early Eocene. The climate was moist and warm. As indicated by the sporo-polleu assemblage the present Paleoeene to Eocene flora in the whole phytoprovienee may belong to a flora transitional from western to eastern China. In this paper the distribution of Aquilapollenites pollen and its stratigraphicai significance are also discussed and it is considered that this kind of pollen distributed in the Paleoeene of China is eertainly not redeposited. At the same time, sonm palynoforms of certain significance for delimitation of Paleoeene and Eocene are provided.     

The Sporo-Pollen Assemblage of the red Beds of Nandian in Lancang of Yunnan

The Red Beds of Nandian consists mainly of red mud stone, marlite, quartzitic sand- stanc, Sabulous psephyte and fine psephyte. Predecessor divided them into three formations referring to Upper Triassic-Jurassic. The marlite in Upper Formation of the Red Beds are rich in fossil Sporo-pollen. The fossil spores and pollen contain 17 genera and 15 species, including 3 new species and 2 new combinations. The Sporo-pollen assemblage is characterized by the absolut predominance of the pollen of gymnospcrms (84.8%) compared with the spores of pteridophyta (15.2%), The pollen are specially characterized by the abundance of Classeopollis Pfiug (75.3%) with a certain amount of Psophosphaera (4.5%), Monosulites (2.4%) and a minority of Pinuspollenites (0.6%). The spores grains are characterized by the abundance of Cyathidites (4.1%), and also by presence of Calamospora (2.1%), Reticulatisporitcs (2.4%), Indeterminable form (1.2%), and rare Schizaeisporites (0.3%). Based on the sporo-pollen assemblage of Upper Jurrassic of the Eurasia continent is characterized by the absolute abundance of Classpollis Pflug (>50% of the total number), the Upper Formation oi the Red Beds of Nandian may be referred to Upper Jurassic age. The climate of the Nandian region in the Late Jurassic period is supposed to be as arid as that of Africa at the present time. Bundary of the eastern termination of southern part of the Eurasia continent in- ferred by Vakhrameev (1970) should be complemented.     

江汉盆地渔洋组上部的古新世孢粉组合

20世纪70至80年代,几乎所有关于江汉盆地跑马岗组和渔洋组的古生物文献都将其时代确定为晚白垩世。李伟同在深入地研究了跑马岗组标准剖面上的轮藻化石后提出跑马岗组大部分属于古新统。取自层位与跑马岗组相当的渔洋组上段上部的岩芯样品经分析后获得了丰富的孢粉化石。这一组合总的面貌与王大宁等获得的晚白垩世渔洋组的第三组合区别明显,而与古近纪的新沟嘴组的第四组合相似处很多。在该孢粉植物群中,常见于当地的晚白垩世分子,如希指蕨孢属、克拉梭粉属、江汉粉属、刺参粉属、鹰粉属等没有或极少见到。而在本组合常见的南岭粉属、五边粉属、漆树粉属等多出现在我国南方古新世地层中。因此,其时代很有可能为古新世。如果是这样,渔洋组上部有可能跨越了白垩系和古近系界线。从当前孢粉植物群含有很多亚热带成分以及麻黄粉属以及胡颓子粉属、忍冬粉属等看,它反映了江汉盆地当时偏干旱的亚热带气候。     

湖北秭归盆地中侏罗世泄滩组孢粉组合

本文记录了鄂西秭归盆地东缘泄滩组上部21个属36个种的孢粉植物群,其中包括3个新种,讨论了主要属种组合特征和地层延限,根据中侏罗世孢粉植物群组合在国内外的对比,提出泄滩组上部Cyathidites minor-Classopollis-Callialasporites组合的时代属于中侏罗世,这个组合以桫椤科,掌鳞杉科植物的繁盛,和出现大量Cal-lialasporites母体植物为特点,暗示泄滩组上部形成于炎热而干旱的气候环境。