陆地生物圈模型的发展与应用

陆地生物圈与大气圈和水圈之间能量、水和碳氮等元素的交换和循环对整个地球系统产生了深刻的影响。陆地生物圈模型(TBM)是研究陆地生态系统如何响应和反馈全球变化的重要方法和工具。通过对从生态系统到区域和全球陆地生物圈不同空间尺度的植被动态、生物地球物理和生物地球化学循环过程、水循环和水文过程、自然干扰和人类活动等过程时间动态的模拟, 陆地生物圈模型被广泛地应用于评估和归因过去陆地生物圈的时空变化和预测陆地生物圈对未来全球变化的响应和反馈。该文简要回顾了陆地生物圈模型的发展, 总结了模型对陆地生态系统主要过程的刻画和模型在生态系统生态学的应用, 并对未来陆地生物圈模型的发展和应用进行了展望。     

植被对近地面层水热交换影响的参数化模型

应用植被对地表面与大气之间水热交换影响的参数化模型,研究了不同植被类型的覆盖度、土壤湿度下垫面的土壤温度、叶温、近地面层气温及地表面与大气之间热量和水分交换,并一相应的观测资料进行了比较,结果表明,模拟不同植被的不同植被的温湿状况、辐射特征和能量平衡关系是合理的。因此,该参数化模型可于中尺度气象模型、气候模拟和环境生态学的研究。     

中国东北样带的梯度分析及其预测

陆地样带研究已成为国际地圈-生物圈计划(IGBP)全球变化研究的重要手段与热点。中国东北样带(NECT)已被列为IGBP国际全球变化陆地样带之一。该样带在东经112°与130°30＇之间沿北纬43°30＇设置,长约1600km,是一条中纬度温带以降水为驱动因素的梯度,具有由温带针阔叶混交林向温带草原的3个亚地带——草甸草原、典型草原与荒漠草原过渡的空间系列。该样带上有4个生态实验站。在大量的固定样地、实验调查研究资料与数据的基础上给出了样带的初步梯度分析及在全球变化图景下的预测,包括其地理位置、设置意义、地形地貌、气候梯度、土壤类型、植被类型和土地利用格局,一个遥感数据驱动的模型和NPP模型在整个样带上运行过。今后NECT将在生物地球化学循环(水、C、N、P等与痕量气体CO_2、CH_4等)、生态系统结构、功能与动态、生物多样性、土地利用与土地覆盖、动态全球植被模型(DGVM)以及高分辨率遥感数据应用等方面得到加强,将成为我国全球变化与陆地生态系统(GCTE)与其它IGBP核心项目研究的前沿阵地。     

全球气候变化的中国自然植被的净第一性生产力研究

本文根据已建立的植物生理生态学特点与水热平衡关系的植物净第一性生产力模型对中国自然植被的净第一性生产力现状及全球变化后的自然植被的净第一性生产力进行了分析,给出了中国陆地生态系统自然植被的净第一性生产力在全球气候变化条件下的变化图景,为合理开发、利用自然资源,以及监测和预测中国陆地生态系统自然植被净第一性生产力的变化及应采取的策略提供了科学依据。     

中国陆地植被氧气生产量变化模拟及其影响因素

大气中氧气主要来源于陆地绿色植物的光合作用,其含量高低直接影响大气质量。基于植被归一化指数(NDVI)、气温、辐射等要素,运用C-FIX模型,在模拟绿色植物净生态系统生产力(NEP)基础上,依据碳氧平衡原理,并结合Arc GIS空间叠加、裁切、栅格计算、空间统计等方法,估算了2001年、2005年和2009年中国陆地植被年氧气生产量,并通过控制实验分析了影响其变化的因素。结果表明:(1)2001年、2005年、2009年中国陆地植被年平均氧气生产量为531.618×107t,云南省、内蒙古自治区、黑龙江省、四川省年氧气生产量最多。中国植被氧气生产量随季节变化明显,夏季最多,其值是春、秋季的2倍,冬季的8倍。(2)中国陆地植被年氧气生产量空间分布为东南高、西北低,以福建省、浙江省、台湾省、云南南部、西藏东南部最高。各季节氧气生产量春季以云南省南部、西藏东南部和中国华东地区为最高,夏季以大、小兴安岭及长白山一带最高,秋季位于东南沿海、云南省、西藏东南部,冬季集中分布在云南南部与海南省。(3)2001—2009年中国陆地植被年氧气生产量呈增加趋势,增加率为7.886%。宁夏回族自治区增加最多,山西省次之。(4)植被覆盖变化是决定年氧气生产量增加的主要因素,贡献率约为60%,CO2浓度增加、气候变化分别承担了28%、12%的贡献率。     

西北干旱区河西走廊荒漠绿洲土地覆盖类型与蒸散的关系研究——基于Landsat 8和ZY3数据融合

蒸散是地表水热平衡的重要分量,也是陆地生态过程与水文过程之间的重要纽带,尤其在干旱区地-气相互作用、碳循环、水循环等过程所包含的物质与能量交换中占有极其重要的地位。基于Landsat 8遥感影像和资源三号影像(ZY3)的高分辨率植被信息,利用SEBS模型对西北干旱区河西走廊中段临泽绿洲北部区域地表蒸散量进行了估算,并用绿洲内部和绿洲-荒漠过渡带两个通量塔涡动相关数据对模型进行评估,分析了不同土地覆盖类型对蒸散量空间分布的影响。结果表明:(1)SEBS模型模拟值与实测日蒸散值之间拟合效果较好,且在均一地表时(绿洲农田区)估算精度更高(R~2=0.96,P0.001),RMSE、MAE分别为0.84 mm/d、0.56 mm/d;(2)从季节变化来看蒸散量与作物生长密切相关,夏季灌溉和降雨使得研究区水分充足,植被覆盖度高,蒸散量相应增加,在绿洲地区可达5.95 mm/d,而冬季最小仅为0.52 mm/d;(3)从蒸散量的空间变化来看,水体蒸散值最大,其余依次为农田、防护林、裸地和灌木丛,说明除水体外,随着植被覆盖的增大,蒸散量也逐渐增加。通过ZY3影像的高分辨率植被信息与Landsat 8影像热红外数据融合,提高了SEBS模型对该区域蒸散量的模拟效果,增进了我们对绿洲下垫面与大气间水热交换规律、水文过程、生态-水文相互作用的深入理解。     

菌根真菌在陆地生态系统碳循环中的作用

在陆地生态系统中,土壤、植被与大气之间有着可观的碳交换通量,陆地生态系统碳循环也和全球气候变化密切关联。菌根真菌可与绝大多数陆地植物建立菌根共生关系,通过矿质养分-碳交换连接起生态系统地上与地下部分,深度参与和影响陆地生态系统的碳循环过程。该文从碳的输入,土壤有机质的形成、稳定和分解等4个关键环节分别论述了菌根真菌在陆地生态系统碳循环中的作用。研究表明,菌根真菌在陆地生态系统碳的输入过程中扮演关键角色,其通过改善植物矿质营养,参与植物逆境响应,影响植物的光合作用强度,以及调控植物多样性与生产力之间的关系等多种途径,维持或提高植被初级生产力;大气中的CO₂被植物固定后,一部分碳经由菌丝网络输送到土壤中,随后经微生物的分解和转化,与矿物结合或被团聚体包裹而被稳定在土壤中;同时,菌根真菌通过影响根际激发效应和菌丝际生物化学过程,如分泌特定胞外酶,与菌丝际微生物互作,驱动芬顿反应,以及与腐生微生物竞争等,调控土壤有机质的分解和转化过程。考虑到菌根真菌对环境和气候变化的敏感性,该文还探讨了全球变化因子对菌根真菌介导的碳循环过程的影响。最后,该文对未来研究方向进行了展望,并提...     

陆地生态系统碳循环模型研究概述

陆地碳循环研究是全球变化研究中的一个重要组成部分,而碳循环模型已成为目前研究陆地碳循环的必要手段.本文针对有关碳循环研究方面的进展,介绍了陆地碳循环模型的基本结构、碳循环过程中涉及的两个基本模型以及目前陆地生态系统碳循环模型的两大类型,并通过对现有主要陆地生态系统碳收支模式的分析,指出了未来陆地碳循环模型的研究方向可能是发展基于动态植被的生物物理模型.这种耦合模型也可能是地球系统模式的重要组成部分.     

2005年中国科学院生命科学和医学学部新当选院士简介

方精云生态学家。北京大学教授。1959年出生于安徽怀宁。1982年毕业于安徽农学院林学系,1989年获日本大阪市立大学生物学博士学位。现任中国生态学会副理事长及国内外多个学术刊物的副主编或编委。建立了我国陆地植被和土壤碳储量的研究方法,系统研究了我国陆地生态系统的碳储量及其变化,较早地开展了碳循环主要过程的野外观测,构建了中国第一个国家尺度的陆地碳循环模式,为我国陆地碳循环的研究奠定了基础;系统研究了我国大尺度的植被动态及时空变化,揭示了我国植被生产力的变化趋势、空间分异及其对气候变化响应的规律;系统开展了我国植被分布与气候关系的定量研究,提出了基于植被气候关系的我国植被带划分的原则和依据,首次采用统一的调查方法,较系统地研究了我国山地植物多样性的分布规律。     

植被对多年冻土的影响研究进展

作为冰冻圈的主体,多年冻土是岩石圈与大气圈水热交换的产物,它的存在、分布及水热过程受到多种时空尺度环境因子的控制和影响.植被是生物圈的重要组成部分,是岩石圈与大气圈热量交换的媒介,它的存在和变化影响着多年冻土的水热过程和空间分布.文章综述了近几十年来植被对多年冻土影响的研究.首先,植被参与地气之间的水热周转过程,通过反射太阳辐射、贴地植被的吸水保水作用以及截留积雪作用等,对下伏冻土产生错综复杂的影响.但是不同植被类型的反射太阳辐射能力、保水与截留能力等各不相同,产生的影响大小有别.其次,同一植被类型的不同层次(如乔木层、灌木层等)对多年冻土的影响也不同,其中贴地植被产生的影响最显著.植被截留积雪,使得地面接收的太阳辐射和地表水分重分配复杂化,从而间接地影响多年冻土环境.因此当植被发生扰动后(如森林火灾和砍伐植被),就会对其所处的多年冻土环境产生各种不利影响,引发冻土灾害,甚至导致多年冻土消融.实际上植被与冻土同为寒区自然生态系统和环境的重要组成部分,它们在长期地质和生物演化中形成生态平衡.因此,植被还常常被用于指示多年冻土及其空间分布.最后提出目前研究中存在的问题,并对未来研究方向进行了展望.     

Response of macroinvertebrate communities to land use and water quality in Wudalianchi Lake

Macroinvertebrate assemblages are structured by a number of abiotic and biotic factors interacting simultaneously. We investigated macroinvertebrate assemblages along gradients of human disturbance and morphometric characteristics in five lakes connected by the same stream. We aimed to assess the relative effects of environmental gradients on macroinvertebrate assemblages and to investigate whether water quality effects on the assemblages were correlated with buffer land use. There were significant differences in macroinvertebrate community compositions among lakes, and our results indicated that oligochaetes (mainly Limnodrilus) and insects (mainly Chironomus) contributed highly to the differences. We used redundancy analysis with variation partitioning to quantify the independent and combined anthropogenic effects of water quality and land use gradients on the macroinvertebrate community. The independent effect of water quality was responsible for 17% of the total variance in macroinvertebrate community composition, the independent effect of buffer land use accounted for 6% of variation, and the combined variation between land use change and water quality accounted for 12%. Our study indicated that both the independent effects of land use and within‐lake water quality can explain the influence in macroinvertebrate assemblages, with significant interactions between the two. This is rather important to notice that changes in buffer land use generally may alter nutrient inputs and thus severely affect abiotic conditions encountered by macroinvertebrate. Our study demonstrates that considering buffer zone effects explicitly may be significant in the selection and application of conservation and management strategies.     

Effects of temperature and pH on the water exchange through erythrocyte membranes: Nuclear magnetic resonance studies

Summary The temperature and pH dependence of water exchange has been studied on isolated erythrocytes suspended in isotonic buffered solutions. At pH 7.4 a break in the Arrhenius plot of water exchange time at around 26°C was found. The mean value of the apparent activation energy of the water exchange time at temperatures higher than that of the discontinuity was 5.7 kcal/mole (±0.4); at lower temperatures the values of the apparent activation energy were below 1.4 kcal/mole. The pH dependence of water exchange time of isolated erythrocytes revealed a marked increase of the water exchange time values in the acid range of pH; a much smaller variation of the same parameter occurs between pH 7.0 and 8.0. These finding could be correlated with other processes involving erythrocyte membranes that showed similar pH and temperature dependence and were considered to indicate state transitions in the membranes. It is suggested that the temperature and pH effects on water diffusion indicate that conformational changes and cooperative effects are implicated in the mechanism of this transport process.Institute for Isotopic and Molecular Technology.     

Spatial variability and temporal trends in water‐use efficiency of European forests

The increasing carbon dioxide (CO₂) concentration in the atmosphere in combination with climatic changes throughout the last century are likely to have had a profound effect on the physiology of trees: altering the carbon and water fluxes passing through the stomatal pores. However, the magnitude and spatial patterns of such changes in natural forests remain highly uncertain. Here, stable carbon isotope ratios from a network of 35 tree‐ring sites located across Europe are investigated to determine the intrinsic water‐use efficiency (iWUE), the ratio of photosynthesis to stomatal conductance from 1901 to 2000. The results were compared with simulations of a dynamic vegetation model (LPX‐Bern 1.0) that integrates numerous ecosystem and land–atmosphere exchange processes in a theoretical framework. The spatial pattern of tree‐ring derived iWUE of the investigated coniferous and deciduous species and the model results agreed significantly with a clear south‐to‐north gradient, as well as a general increase in iWUE over the 20th century. The magnitude of the iWUE increase was not spatially uniform, with the strongest increase observed and modelled for temperate forests in Central Europe, a region where summer soil‐water availability decreased over the last century. We were able to demonstrate that the combined effects of increasing CO₂ and climate change leading to soil drying have resulted in an accelerated increase in iWUE. These findings will help to reduce uncertainties in the land surface schemes of global climate models, where vegetation–climate feedbacks are currently still poorly constrained by observational data.     

流溪河流域景观空间特征与河流水质的关联分析

人类活动影响或改变流域景观空间结构,并有可能对河流水质产生不同程度的影响,以流溪河流域为研究区,分析流域景观空间格局特征与水质指数之间的相关关系。将流域划分为27个子流域,采集水样分析水质状况,所选用的水质指标有氨氮(NH3-N)、硝态氮-亚硝态氮(NO3-N+NO2-N)、总磷(TP)、化学需氧量(CODCr)。结果表明:1)该流域土地利用结构与水质具有显著相关性,其中居住用地对水质的影响作用最强,林地对河流水质具有净化功能,与水质指标之间的关系表现为负相关,园地与水质指标关系具有不确定性;2)流域景观特征从上游到下游之间表现为城市化增强的梯度,水质状况响应这个梯度变化表现为上游优于下游,人类活动及城市化发展引起的土地利用变化及土地管理方式对水质变化有显著影响;(3)景观破碎度与水质呈现显著正相关性,是影响水质的重要指标,景观聚集程度和斑块形状复杂程度与水质有负相关关系;子流域尺度和河岸带尺度景观空间特征对水质的影响差异不明显。     

Water use and yield of bioenergy poplar in future climates: modelling the interactive effects of elevated atmospheric CO2 and climate on productivity and water use

Vegetation exerts large control on global biogeochemical cycles through the processes of photosynthesis and transpiration that exchange CO₂ and water between the land and the atmosphere. Increasing atmospheric CO₂ concentrations exert direct effects on vegetation through enhanced photosynthesis and reduced stomatal conductance, and indirect effects through changes in climatic variables that drive these processes. How these direct and indirect CO₂ impacts interact with each other to affect plant productivity and water use has not been explicitly analysed and remains unclear, yet is important to fully understand the response of the global carbon cycle to future climate change. Here, we use a set of factorial modelling experiments to quantify the direct and indirect impacts of atmospheric CO₂ and their interaction on yield and water use in bioenergy short rotation coppice poplar, in addition to quantifying the impact of other environmental drivers such as soil type. We use the JULES land‐surface model forced with a ten‐member ensemble of projected climate change for 2100 with atmospheric CO₂ concentrations representative of the A1B emissions scenario. We show that the simulated response of plant productivity to future climate change was nonadditive in JULES, however this nonadditivity was not apparent for plant transpiration. The responses of both growth and transpiration under all experimental scenarios were highly variable between sites, highlighting the complexity of interactions between direct physiological CO₂ effects and indirect climate effects. As a result, no general pattern explaining the response of bioenergy poplar water use and yield to future climate change could be discerned across sites. This study suggests attempts to infer future climate change impacts on the land biosphere from studies that force with either the direct or indirect CO₂ effects in isolation from each other may lead to incorrect conclusions in terms of both the direction and magnitude of plant response to future climate change.     

水分对林地和草地土壤氮初级转化速率的影响

水分含量是与土壤氮转化相关微生物活性的重要影响因素。本研究以黑龙江省北安市的草地和林地土壤为对象,通过室内培养试验,利用¹⁵N同位素标记技术和FLUAZ数值优化模型研究60%和100%田间持水量(WHC)条件下土壤氮初级矿化速率、初级固定速率、初级硝化速率和初级反硝化速率,以探讨土壤氮初级转化速率对水分含量变化的响应,阐明不同水分条件下土壤中氮的产生、消耗、保存机制及其生态环境效应。结果表明: 土壤水分变化不影响草地和林地土壤氮初级矿化速率和铵态氮固定速率,水分含量由60% WHC增加至100% WHC后显著增加了林地土壤的初级硝化速率,但对草地土壤的初级硝化速率没有显著影响。60% WHC条件下草地和林地土壤的初级反硝化速率可以忽略不计,水分含量增加至100% WHC后土壤初级反硝化速率显著提高,且草地土壤的初级反硝化速率显著低于林地土壤。100% WHC条件下林地土壤初级硝化速率与铵态氮固定速率比值(gn/ia)和N₂O排放量均显著高于60% WHC;100% WHC条件下草地土壤的N₂O排放量显著高于60% WHC,但两个水分条件下的gn/ia值无显著差异。表明短期内水分含量的增加可能会增加草地和林地土壤氮转化的负面环境效应,且对林地土壤的影响尤为显著。     

The boundary filtration effect of reed-dominated ecotones under water level fluctuations

The boundary filtration effect of land/water ecotones with reed-bed/ditch systems under water level fluctuations was studied in Baiyangdian Lake of North China. It was found that the changes in reed bed areas which were primarily affected by water level fluctuations and the root channels in the wetland soils together largely determined boundary filtration efficiency. The ecotones displayed the greatest boundary effect at a moderate water level of about 8 m above sea level. The massive root channels in the wetland soils promoted water flowing into the reed beds as far as 8 m horizontally by subsurface in wet years. In dry years, when the water level was below the root channel distribution zone, the lateral water exchange width of ecotones was limited to 0.5 m along the fringe area. It is calculated that, at 8 m water level, the total boundary length of ecotones is 7,273 km and the boundary exchange volume is 5.8 × 10⁶ m³. While at 6.5 m water level, the total boundary length of ecotones is reduced to 2,699 km and the boundary exchange volume is 1.1 × 10⁵ m³. The standard capacity for phosphorus retention was 105.9 and 2.5 tonnes at water levels of 8 and 6.5 m, respectively. This suggests that the boundary filtration effect of reed-bed/ditch wetlands is important for improving the water quality of inland waters, and this effect should be considered in regulating and managing lake water levels.     

黄土塬区苹果园土壤有机碳分布特征

以黄土塬区塬面和梁坡梯田5、10、15a和20a苹果园为对象,在行间距果树1.0、1.5m和2.0m处用土钻法分层采集0-100cm土样,LiquiTOCⅡ测定样品土壤有机碳(Soil Organic Carbon,SOC)含量,分析两种地形条件下各龄果园SOC的分布特征。结果表明:塬面5、10、15a和20a果园SOC分别为6.39、6.46、6.66g/kg和6.47g/kg,梁坡分别为5.83、6.05、6.54g/kg和6.09g/kg,两种地形条件下同龄果园SOC差异显著(P0.05),但龄间SOC消长趋势相似,均有15a的20a的10a的5a的;水平方向上,5a果园SOC沿树干向外增大,10a果园减小,15a和20a果园变化较小,塬面和梁坡同龄果园间SOC水平分布格局较一致;垂直方向上,同梁坡相比,塬面果园4个层次(0-10cm、10-20cm、20-50cm和50-100cm)的平均SOC较高;梁坡果园50-100cm土层的SOC龄间差异较大,20-50cm土层龄间差异较小,塬面果园50-100cm土层的SOC龄间变化较小;在"纯果园"利用阶段,果园利用方式并未引起SOC下降,深层SOC有明显的积累效应。     

土地利用变化对海南土壤水源涵养功能的影响

热带地区已有大量原始林转变为其他土地利用类型,影响了陆地生态系统的水源涵养功能.为了明确热带原始林转变为其他土地类型对土壤水源涵养功能的影响,在海南中部山区选择4种典型土地利用类型,包括林龄大于100年的原始林(VF)、10年生次生林(SF)、12年生槟榔林(AF)和35年生橡胶林(RF),评估土地利用变化对土壤持水性能和水源涵养功能指数(SWI)的影响.结果表明: 与原始林相比,表层土壤(0～10 cm)中,其他土地类型土壤持水性能指标均降低,12年槟榔林各土层指标均最低.土壤含水量和最大持水量与植被郁闭度、土壤有机质和土壤容重显著相关,表明郁闭度、土壤有机质和紧实度的改变是土壤持水性能变化的重要原因.与原始林相比,次生林、槟榔林和橡胶林土壤水源涵养功能分别减少27.7%、54.3%和11.5%,不同土层的差异各异,橡胶林仅表层土壤水源涵养功能显著降低.植被郁闭度、土壤有机质和土壤容重可解释土壤水源涵养功能变量的83.3%.土地利用转变显著改变了土壤持水性能和土壤水源涵养功能,相比12年槟榔林,35年橡胶林能更好地保持土壤水分,土地管理中增加土壤有机质和减少土壤紧实度可改善土壤持水性能及水源涵养功能.