化石直脉蝎蛉科再分类(长翅目)

本文对化石直脉蝎蛉科(Orthophlebiidae)的分属做了再分类,仍保留该科内的 2个属:直脉蝎蛉属 Orthophlebia和原直脉蝎蛉属Protorthophlebia,解体中蝎蛉属Mesopanorpa。中蝎蛉属被解体后,其包含所有的种,如Rs1主干带3支或3支以上支脉,归入直脉蝎蛉属;如Rs1仅带2支支脉,归入原直脉蝎蛉属;原中蝎蛉属以Rs主干两倍长于MA主干作为属的分类根据,但这一属征常常导致分类上的混乱,故本文不作为属或亚属的分类根据。上述保留的2个属,再以Rs分支先于、后于或等于MA的特征,即Rs主干短于、长于或等于MA主干作为亚属的分类根据。以此,在直脉蝎蛉属Orthophlebia和原直脉蝎蛉属Protorthophlebia之下各自设立3个亚属(Hong et Zhang, 2004)。在已建立的6个亚属中,Orthophlebia(Brachyphlebia)Hong et Zhang, 2004和Protorthophlebia(Mecynophlebia)Hong et Zhang,2004分别修改为Orthophlebia(Orthophlebia)Westwood, 1845和Protorthophlebia(Protorthophlebia)Tillyard,1933的新异名。中生代直脉蝎蛉中脉的前支(MA)并非在古生代之后就完全消失。根据对中生代不同时期、不同层位、不同地区直脉蝎蛉和本目其他科在内化石标本的观察,中脉及其前、后支脉(MA、MP)仍然继续存在与发展。基于这种事实,本文认为直脉蝎蛉科脉序命名中沿用已久的Rs后支(Rs3 4)应为中脉的前支(MA),原来沿用的中脉(M)为中脉的后支(MP)。这样,有助于我们对长翅目中脉及其前、后(MA、MP)支脉的准确辨认,尤其是对MA脉在地史上变革、发展,以及与Rs脉的演化关系的认识可能有所裨益。     

长蝎蛉幼期形态附生物学记述

【目的】蝎蛉科(Panorpidae)是长翅目(Mecoptera)最大的科,是重要的生态指示昆虫。然而,由于对环境条件要求苛刻,饲养困难,其幼期研究很不充分。【方法】本研究通过人工饲养成虫获得了长蝎蛉Panorpa macrostyla Hua的卵、幼虫和蛹等全部虫态,运用光学显微镜和扫描电子显微技术观察了其超微形态,并简要记载了其生物学特性。【结果】长蝎蛉每年发生1代,成虫发生于6月末至8月初。卵椭球形,卵壳表面覆盖一层隆起的网状结构。幼虫蠋型,具3对分4节的胸足和8对不分节的腹足;头壳高度骨化,具1对由26个小眼组成的复眼和1对3节的触角,口器咀嚼式;腹部第1-9节背面具有成对的背毛突,第10节仅有1根背毛突,腹部末端具有一个可伸缩的吸盘;呼吸系统为周气门式,具1对前胸气门和8对腹气门。幼虫共4个龄期,以预蛹期在土室内越冬。蛹为强颚离蛹,外形接近成虫,雄蛹腹部末端膨大。【结论】基于幼虫形态特征,长蝎蛉明显区别于新蝎蛉属Neopanorpa、华蝎蛉属Sinopanorpa、双角蝎蛉属Dicerapanorpa以及单角蝎蛉属Cerapanorpa幼虫。然而,长蝎蛉幼虫头部刚毛L2和SO2,腹部末节刚毛D2,SD1和SD2端部均膨大呈棒状,与蝎蛉属Panorpa其他种类区别明显,表明长蝎蛉的属级地位需要进一步研究。     

黄脸油葫芦消化道和马氏管的组织学观察

应用石蜡切片技术对黄脸油葫芦Teleogryllus emma(Ohmachi and Matsumura)成虫消化道和马氏管的显微结构进行观察。消化道由前肠、中肠和后肠3部分组成:前肠由内向外可分为6层:内膜、肠壁细胞层、底膜、纵肌、环肌和围膜;中肠组织结构也分为6层,即由内向外依次为围食膜、肠壁细胞层、底膜、环肌、纵肌和围膜;后肠的组织结构与前肠基本相似,但内膜比前肠的薄,且肌肉的排列较前肠不规则,与中肠的肌肉排列相似,即环肌在内,纵肌在外。消化道各部位的结构差异与功能有密切关系。马氏管管壁由8个左右形状多变并具有显著细胞核的大形的单层上皮细胞组成。     

蝎蛉科(长翅目)昆虫雌性生殖系统构造及其在分类学上的意义

蝎蛉科Panorpidae6种蝎蛉的雌性生殖系统间存在显著差异,尤其是卵巢管数目、受精囊包膜颜色、附腺大小、及各部分相互间位置。卵巢管为多滋式,每个卵巢所含卵巢管数目在长瓣蝎蛉Panorpa longihypovalva Hua et Cai和路氏新蝎蛉Neopanorpa lui Chou et Ran中为10根,在大蝎蛉P.magna Chou中为16根,在太白蝎蛉P.obtusa Cheng中为18根,在染翅蝎蛉P.tincta Navas中有28根,在长白山蝎蛉P.changbaishana nom.nov.中,一个卵巢一般由8根卵巢管组成,而另一个经常为10根,表明在同一个体中有不对称性。长白山蝎蛉的受精囊包膜为红色,而其它种类透明。大蝎蛉附腺在6种蝎蛉中最大,几乎与卵巢等粗。表明雌性生殖系统可用于蝎蛉科的种类鉴别,并简要讨论了长翅目Mecoptera的系统发育关系。此外,长白山蝎蛉Panorpa changbaishana Hua是为Panorpa choui Hua,1998所提订的新名。     

河南省伏牛山蝎蛉记述：长翅目：蝎蛉科

本文报道了１９９６年７月河南省伏牛山昆虫考察中采到的蝎蛉科６种昆虫,其中蝎蛉属Ｐａｍｏｒ ｐａ４种,即大蝎蛉Ｐａｎｏｒ ｐａ ｍａｇｎａ Ｃｈｏｕ,六刺蝎蛉Ｐ．ｓｅｘｓｐｉｎａ Ｃｈｅｎｇ,华山蝎蛉Ｐ．ｅｍａｒｇｉｎａｔａ Ｃｈｅｎｇ和伏牛山蝎蛉Ｐ．ｆｕｎｉｕｓｈａｎａ Ｈｕａ ｅｔ Ｃｈｏｕ,ｓｐ．ｎ。新蝎蛉属Ｎｅｏ ｐａｎｏｒ ｐａ２种,即河南新蝎蛉Ｎｅｏｐａｎｏｒ ｐａ ｌｏｎｇｉｐｒｏｃｅ     

南五台蝎蛉成虫复眼的超微结构

采用扫描电镜和组织切片法,观察南五台蝎蛉Panorpa nanwutaina Chou成虫复眼的超微结构。南五台蝎蛉复眼近半椭球形,包括1500～1600个小眼。小眼表面光滑,由角膜、晶体、2个初级和12个次级色素细胞、视杆、以及基膜组成。角膜为多层片状纤维结构;晶体含有4个晶锥细胞;视杆由若干个视网膜细胞组成。晶体、视杆周围、和色素细胞内含有大量的色素颗粒,基膜两侧也有色素颗粒分布。南五台蝎蛉的复眼属于并列像眼。与普通蝎蛉P.communis L.小眼的次级色素细胞数目不同。讨论了南五台蝎蛉角膜的功能以及感觉毛和次级色素细胞在分类中的作用。     

蚊蝎蛉捕捉足构造

依据光学显微镜和扫描电镜观察,对扁蚊蝎蛉Bittacus planus Cheng和缠绕蚊蝎蛉Bittacus implicatus Huang et Hua成虫足的外部形态、肌肉组织及超微结构进行了描述和绘图。蚊蝎蛉成虫三对足结构相似,均为捕捉式,以高度特化的跗节捕捉猎物,各跗分节间具发达的关节、凹槽和爪缩肌肌腱膨大区,第4和5跗分节的齿形成嵌合构造,在昆虫纲中为该类群独有特征。首次发现胫节伸肌b由腿节基部发出的一小块三角形肌肉组织和一条长肌腱组成; 爪缩肌肌腱在第5跗分节凹槽末端有一突起,收缩时突起羁绊于凹槽内的“Y”字形底托。分析了成虫捕食行为与捕捉足构造之间的关系,并简要比较了蚊蝎蛉捕捉足与螳螂捕捉足的结构特点。     

拟蝎蛉科--中国长翅目新记录科

报道中国长翅目Mecopteral新记录科——拟蝎蛉科Panorpodidae。介绍了拟蝎蛉科的主要特征,编制了中国长翅目分科和拟蝎蛉科分属检索表：提出了1个新组合——短拟蝎蛉Panorpodes brevicaudata(Hua,1998),移自蝎蛉属Panorpa Linn．,分布于吉林省长白山。     

蝎蛉科与蚊蝎蛉科昆虫触角感器超微形态比较

触角是昆虫重要的感觉器官,其感觉功能主要是通过其上的各类感器来实现的。本文以染翅蝎蛉Sinopanorpa tincta(Nav偄s)和中华蚊蝎蛉Bittacus sinensis Walker为代表,采用扫描电子显微镜观察了蝎蛉科Panorpidae和蚊蝎蛉科Bittacidae触角感器的类型及其超微结构,并简要讨论了各种感器的功能。结果显示:染翅蝎蛉触角上共发现7种感器,分别为短刺形感器、Bhm氏鬃毛、毛形感器、短毛形感器、锥形感器、刺形感器、和微毛形感器。中华蚊蝎蛉触角上也发现7种感器,分别为Bhm氏鬃毛、芽形感器、毛形感器、刺形感器、畸形毛感器、腔锥感器和钟形感器。蚊蝎蛉触角与蝎蛉的最大区别是着生有大量的腔锥感器,初步推测众多的腔锥感器可能与蚊蝎蛉对湿度的高度敏感有关。     

蝎蛉属Panorpa东北一新种（长翅目：蝎蛉科）

本文描述产自中国东北辽宁和黑龙江的蝎蛉１新种－刘氏蝎蛉Ｐａｎｏｒｐａ ｌｉｕｉ,ｓｐ．ｎｏｖ．。模式标本保存在西北农业大学昆虫博物馆。     

桃小食心虫幼虫的消化道和马氏管的显微结构观察

利用光学显微镜和扫描电子显微镜,在形态学和组织学水平上研究_『桃小食心虫 Carposina sasakii 幼虫消化道和屿氏管的结构.桃小食心虫幼虫消化道由前肠、中肠和后肠组成.前肠细短,肌肉层薄.前肠与中肠交界处有突出的胃盲囊.中肠长且粗大,内有围食膜,肠壁细胞较大,外层为发达的环肌和纵肌.后肠上皮细胞内陷很深.6根念珠状的马氏管位于中、后肠分界处.     

Characterization of the alimentary canal of the aphidophagous ladybird,Adalia bipunctata (Coleoptera: Coccinellidae): anatomical and histological approaches

The alimentary canal of the two‐spot ladybird Adalia bipunctata (Linnaeus) (Coleoptera: Coccinellidae) presents the foregut (stomodeum), the midgut (mesenteron) and the hindgut (proctodeum). The shortest region is the foregut and the longest is the midgut. The relative proportions of the main regions were found to be similar for males and females. In the foregut it was possible to distinguish the pharynx, the esophagus and the proventriculus but no crop. The hindgut is composed of the ileum, rectum and rectal canal. Generally the organ width is similar for males and females, but females presented a wider proventriculus. The epithelium of the foregut varied from squamous to simple cuboidal and columnar. In the midgut the epithelium is simple columnar with goblet and regenerative cells. The epithelium of the hindgut varied from simple cuboidal to squamous. Females presented thicker midgut epithelium whereas males presented thicker epithelium in the esophagus. The anatomy of the alimentary canal of A. bipunctata seems to conform to its carnivorous and recent phylogenetic status within the family Coccinellidae.     

普通齿蛉幼虫的消化系统结构和取食分析

研究了普通齿蛉Neoneuromus ignobilis Navás幼虫消化系统的解剖和电镜扫描结构,并结合肠道内食物残渣的成分分析了其取食特性.结果表明,普通齿蛉幼虫消化系统较为简单,无特殊的变异,消化道细长,由前肠、中肠和后肠构成,前后肠较长,分别占体长的42%和47%,中肠最短,仅占总长度的11%.前肠由食道、嗉...     

光滑足距小蠹消化道的解剖结构

采用体视显微和扫描电镜技术研究光滑足距小蠹Xylosandrus germanus(Blandford)消化道的解剖结构。结果表明,光滑足距小蠹消化道由前肠、中肠和后肠三部分组成。成虫前胃由8个骨化的前胃板组成,呈灯笼状结构。前胃板由板状部和片状部组成,板状部短而简单,片状部甚长,由斜面、咀嚼刷和关闭刚毛组成。胃盲囊着生在中肠近后端,有细管状和囊状2种,成虫分别有1对,幼虫有1对细管状和3~5对囊状。6根马氏管分成2组,1组4根,另一组2根。6根马氏管与后肠肠壁形成隐肾系统。消化道具有1对囊状和1对细管状的胃盲囊可作为光滑足距小蠹成虫的识别特征。     

粉尘螨消化系统的形态学观察

光镜下观察了粉尘螨Dermatophagoides farinae消化系统结构,其组成包括：口前腔、前肠、中肠、后肠、肛门和唾液腺。口前腔由颚体围绕而成;前肠包括一个肌肉的咽和食道,食道从脑中穿过;中肠分为前中肠（包括一对盲肠）和后中肠,中肠的上皮细胞呈现多种形态; 后肠包括相对大的结肠和狭窄的直肠;消化腺为不规则形,位于脑前方。本文阐述了消化道的分支情况、显微结构及细胞形态。     

Morphological and ultrastructural characterization of the alimentary canal in larvae of Streltzoviella insularis (Staudinger) (Lepidoptera: Cossidae)

Streltzoviella insularis (Staudinger) is an important tree‐boring pest, that primarily damages Sophora japonica (Linnaeus) and Ginkgo biloba (Linnaeus), as well as other common species, at great economic cost to the urban landscape construction industry in China. In the present study, the alimentary canal morphology of S. insularis was observed using light microscopy, and its ultrastructure was investigated by scanning and transmission electron microscopy. The foregut of S. insularis can be divided into the pharynx, esophagus, crop, proventriculus, and cardiac valve. The well‐developed crop forms the longest section of the foregut. It is able to store large amounts of food and is lined with a monolayer of epithelial cells. Many sclerotized microspines occur on the surface of the anterior intima and there are dense spines on the posterior intima of the proventriculus. Epithelial cells of the midgut include columnar cells, goblet cells, and regenerative cells, but endocrine cells are absent. The hindgut consists of the pyloric valve, ileum, and rectum. There is no clear distinction between the ileum and colon. The intima surface of the pyloric valve carries many microspines, whereas the intestinal wall of the rectum is thin with well‐developed rectal pads. The rectal epithelial cells form a squamous monolayer. A cryptonephric excretory system is located in the hindgut. There are six spiral Malpighian tubules, in which a cellular layer on a basement membrane encloses a lumen. These results will provide the basis for further studies of the structure and function in S. insularis larvae.     

韭菜迟眼蕈蚊幼虫消化系统的解剖学和组织学

本利用石蜡切片对韭菜迟眼蕈蚊幼虫其消化系统的组织学进行了研究。结果表明,幼虫的消化道无特殊变异,但中肠亚端部的一对胃盲囊长而发达,是消化道的突出特征。中肠和胃盲囊不同部位的细胞学特点有明显差异。根据中肠细胞形状及其分布将中肠分为4个区域,对中肠不同区域的细胞学特点进行了描述。     

Fine Structure of the Midgut and Hindgut in Lepidocampa weberi (Insecta,Diplura)

The fine structure of the alimentary canal, especially the midgut and hindgut of Lepidocampa weberi (Diplura: Campodeidae) is described. The general organization of the canal is similar to that of Campodea. The midgut epithelium is composed of columnar apical microvillated cells. Each nucleus contains a single intranuclear crystal. Close to the pyloric region, the posterior midgut cells are devoid of microvilli and intranuclear crystals. There is no special pyloric chamber as in Protura or pyloric cuticular ring as in Collembola but a morphological transformation from midgut to hindgut cells. Eight globular Malpighian papillae, consisting of distal microvillated cells and flat proximal cells, open into the gut lumen via ducts formed by hindgut cells. The structure of the hindgut is complicated and can be divided into three segments. The anterior hindgut cells have an irregular shape and compact cytoplasm. A striking interdigitation between the large bottle-shaped epithelial cells and longitudinal muscle cells occurs in the middle segment of the hindgut. The thick cuticle gives rise to long spikes projecting into the gut lumen. The posterior hindgut cells possess the morphological features for water reabsorption. Some hypotheses are advanced about the function of the different regions of the gut.