长江三峡中坝遗址地层的孢粉学初步研究

对重庆忠县中坝遗址剖面82个地层样品和遗址附近的12个现代表土样品进行孢粉分析,结果表明:表土样品中孢粉含量较高,沉积地层中频繁出现蕨类植物孢子(相对于木本和草本花粉,其数量多,比例高),揭示出中坝地区植被长期处于次生化状态。表土花粉中,松在乔灌木中占明显优势;旱生草本花粉以蒿属、十字花科和菊科等为主;蕨类孢子以水龙骨、三缝孢子、凤尾蕨、石松孢子和单缝孢子为主,植被处于次生化。地层花粉中,蕨类孢子以凤尾蕨属、卷柏属、其它三缝孢子和单缝孢子为主;乔灌木植物花粉以松属和榆属占优势;草本植物花粉中,禾本科、菊科、藜科和蒿属等含量较高。研究点孢粉分析显示,人类活动对中坝遗址影响较大:1)新石器晚期:先人栽培种植活动已经开始;2)新石器时代末期至六朝,受人类活动影响,植被覆盖度较低;3)商代至周初至秦汉时期,人类活动不断加强;4)唐宋时期,人类栽培种植活动较活跃;5)明清至现代,人类砍伐当地植被日趋严重,导致植被次生化现象增强。     

东北长白山垂直林带下现代表土花粉与植被关系

在长白山北坡5个垂直植被带的林下采集29个表土(苔藓)样品进行孢粉分析,同时将其结果作对应分析.结果表明,除个别样品,在长白山垂直植被带的孢粉组合中乔木花粉占优势,其中以松属和桦木属最多,在各植被带均有分布.松属花粉最高含量出现在以红松为主的针阔混交林带的孢粉组合中;桦木属花粉则大量见于亚高山岳桦林带和以栎桦为主的阔叶落叶林带,但前者伴有耐寒、早的小灌丛杜鹃花属花粉,而在后者中则见有大量蕨类孢子.阔叶落叶林带的孢粉组合是以类型多、百分含量高的落叶乔木花粉和大量蕨类孢子为特征.亚高山针叶林带的孢粉组合中出现数量众多的云杉属花粉.在高山苔原带孢粉组合中虽然乔木花粉略占优势,但极少或不见蕨类孢子,与其他植被带比较,草本植物和小灌丛花粉明鲜较多,其中杜鹃属花粉含量在草本中占优势,其中出现苔原带的指示花粉--仙女木属.     

东北长白山垂直林带下现代表土花粉与植被关系

在长白山北坡5个垂直植被带的林下采集29个表土(苔藓)样品进行孢粉分析，同时将其结果作对应分析。结果表明，除个别样品，在长白山垂直植被带的孢粉组合中乔木花粉占优势，其中以松属和桦木属最多，在各植被带均有分布。松属花粉最高含量出现在以红松为主的针阔混交林带的孢粉组合中；桦木属花粉则大量见于亚高山岳桦林带和以栎桦为主的阔叶落叶林带，但前者伴有耐寒、旱的小灌从杜鹃花属花粉，而在后者中则见有大量蕨类孢子。阔叶落叶林带的孢粉组合是以类型多、百分含量高的落叶乔木花粉和大量蕨类饱子为特征。亚高山针叶林带的孢粉组合中出现数量众多的云杉属花粉。在高山苔原带孢粉组合中虽然乔木花粉略占优势，但极少或不见蕨类孢子，与其他植被带比较，草本植物和小灌丛花粉明鲜较多，其中杜鹃属花粉含量在草本中占优势，其中出现苔原带的指示花粉——仙女木属。     

四川螺髻山表土和化石孢粉揭示的环境生态意义

通过四川螺髻山（1984、2017年）表土和化石孢粉与植被的关系分析发现：孢粉组合中木本植物含量（91.0%）占绝对优势,松属、冷杉属、青冈属、常绿栎类、落叶栎类、桤木属、杜鹃花科、禾本科和蒿属为主要花粉类型;人工次生林花粉组合能很好地反映母体植被的群落特征,并可指示优势种的存在和人类活动痕迹;针阔混交林花粉组合能较好地反映群落整体特征,花粉类型能与母体植被中优势种较好地对应;常绿阔叶林、针叶林和灌丛草甸的花粉组合难以反映母体植被的群落特征。DCA表明,花粉谱的百分含量,能较好地区分人工扰动植被和天然植被,但人工次生林、常绿阔叶林和针阔混交林之间以及针叶林和灌丛草甸之间未能区分。30年前后表土孢粉组合变化明显,1984年样品以松属、青冈属和桤木属等花粉为主,而2017年采样分析中松属、桤木属花粉占绝对优势。随着人类活动的加强,部分地区松树和桤木大面积飞播或种植,表土孢粉组合对当地植被指示意义与扰动强度呈负相关关系。现生植被与表土孢粉组合的差异能为恢复古植被和古环境提供参考,但原生植被已经被破坏地区的孢粉图谱,就很难作为重建历史时期植被的根据。本研究可以为亚热带山地利用孢粉学恢复第四纪时期植被与气候,探讨人类活动与环境的关系提供理论依据和实践参考。     

喀斯特山地典型植被恢复过程中表土孢粉与植被的关系

喀斯特山地坡耕地、1a弃耕地、灌草地、15a疏林地、25a次生林群落表土孢粉分析和植被样方调查结果表明:(1)研究区植物孢粉共由46科属组成,以草本和蕨类植物占据优势,含量为45.77%-68.35%,其次是乔木(主要为雪松和马尾松),含量为(2.53%-21.31%),灌木植物花粉含量最低,为1.04%-8.88%,显示该区植被退化严重;(2)恢复过程中各阶段的花粉类型丰富度差别不明显,物种多样性主要表现为草本、蕨类和灌木植物种类的变化,乔木基本保持不变;(3)喀斯特山地典型植被恢复过程的表土孢粉组合差别明显,与其相应的实际植被情况大致相同,基本反映了石漠化地区现代植被的基本状况,表明孢粉可以为该区恢复和重建自然生态环境提供参考性依据。     

重庆喀斯特地区现代花粉组合与植被的关系

重庆喀斯特地区4种不同地貌单元的81份表土和苔藓样品的孢粉分析结果表明:(1)研究区植物孢粉组合共由109科属组成。除金佛山外,其他植被退化严重地区样点均以草本和蕨类植物占据优势(剔除石漠化治理大量引种的马尾松、柳杉和侧柏等乔木花粉),其次是乔木植物花粉(主要为马尾松),灌木植物花粉含量很低,显示该区植被退化严重;(2)孢粉组合的PCA分析能够很好地将研究区的现代孢粉组合区分开来,较好地反映了重庆喀斯特地区不同地貌单元的植被特征,特别是石漠化区植被退化严重的孢粉组合特征。生态保护较好、自然植被发育的金佛山地区具有106科属的乔木、灌木与草本孢粉类型,以及高含量的乔木花粉,而喀斯特石漠化地区的鸡公山(46科属)、青木关镇(56科属)和南平镇(50科属)则表现为乔灌木孢粉类型较少;(3)银杉作为金佛山特有的孑遗植物,其花粉含量明显低于松属花粉,说明其花粉不利于远距离传播;(4)随着土地利用强度的增加,现代孢粉组合中乔灌木花粉含量和种类越少,草本和蕨类植物孢粉含量越多,且以耕地杂草为主。因此,土地利用方式的变化是喀斯特石漠化区次生植被及其现代孢粉组合变化的主要影响因素。研究可为重庆喀斯特地区土地利用策略的制定,重庆及其周边地区第四纪古植被、古气候和古环境重建研究,以及中国孢粉数据库建设提供基本数据和科学依据。     

重庆金佛山国家自然保护区现代植被表土孢粉的初步研究

通过对重庆金佛山国家自然保护区10个不同植被类型的表土调查和采样,研究该区表土孢粉组合特征.结果表明,研究区植物孢粉共由106科属组成,其中以针叶乔木、蕨类和阔叶乔木孢粉占优势,含量分别为8.94%-56.30%、19.73%-46.77%和6.17%-39.28%,其次是灌木和草本花粉,分别为6.24%-29.77%...     

云南泸沽湖及周边山地表层孢粉与植被关系研究

本文对云南泸沽湖周边山地不同植被类型下采集的32个表土样品和36个泸沽湖湖泊表层样品进行了孢粉分析,探讨了不同植被类型内的表土孢粉散布特征,对比分析了山地表土与湖泊表层孢粉分布特征的差异。结果表明,不同山地植被类型的孢粉组合具有不同的代表性成分与变化特征,能够较好地反映相应的植被类型。湖泊表层孢粉研究表明,湖泊内不同位置的孢粉组合差异较小,说明湖泊表层孢粉反映的是区域植被的平均分布状况。对比分析泸沽湖周边山地表土与湖泊表层样品的孢粉组合,两者存在一定的差异,表现为:湖泊表层的孢粉组合中乔木花粉(主要是松属)平均含量比表土高,但波动范围要小;草本花粉平均含量要低,且波动范围也小。这可能是由于乔木植物植株较高,而且有些乔木植物花粉具有气囊,更易借助风力将花粉传播到比较远的湖泊中的原因造成的。冷杉和铁杉花粉在所有样品中含量都相对较低,分别不高于6.5%和3.6%;桦属、桤木属、常绿栎类花粉的最高值出现在有这些母体植物生长的表土样品中,在其他样品中则明显较低。在表土孢粉样品中经常可以见到冷杉的气孔器,但在湖泊表层样品中却没有见到,说明气孔器不易到达湖泊中。该研究是对中国西南表层孢粉研究的有益补充,同时为泸沽湖新生代特别是第四纪化石孢粉解释提供了可靠的参考和依据。     

西班牙西北部San Isidro和Leitariegos地区全新世孢粉学的比较研究

文中报道了西班牙西北部San Isidro和Leitariegos地区两口钻井全新世孢粉学比较研究的结果.孢粉分种析结果显示了两地孢粉组合成分上的不同.根据从San Isidro地区泥炭沼泽获得的孢粉组合中乔木和灌木花粉百分含量变化,建立了两个组合带.孢粉植物群代表了一个松属占统治地位的森林,桦属、栎属、榛属等花粉含量变化序列可能和气温升高,降雨量增加有关.根据乔木花粉相对丰度变化,Leitariegos地区钻井剖面建立了3个孢粉组合.桦木属花粉含量增加说明是桦树林,栎属和榛属花粉居次要地位,松属花粉含量下降.文中还总结了两个孢粉序列所反映的人类活动、植被演替以及地形景观改变等之间的关系.两地钻井剖面底部碳同位素年代测定表明二者的底部并不同时.     

云南省香格里拉地区现代孢粉与植被关系研究

本文对云南省香格里拉地区(海拔3 400～4 700 m)7个植物群系下采集的21个现代苔藓样品进行了孢粉研究,探讨了现代孢粉组合与样点周围植被之间的关系,以及主要孢粉类型的代表性与传播能力。结果表明,香格里拉地区不同植物群系的孢粉组合具有不同的代表性成分与变化特征,能较好地反映当地植被状况。桦木林和黄背栎林的孢粉组合中指示性成分分别是高含量的桦木属和常绿栎类花粉;冷杉林下的样品,除采自仅有稀疏冷杉的冷杉林下的样品外,其他样品的孢粉组合都较好地反映了高含量的冷杉属花粉;杜鹃灌丛和蔷薇灌丛的孢粉组合分别以高含量的杜鹃花科和蔷薇科花粉为特征;在花粉产量低且植被较开阔的流石滩疏生草甸中,孢粉组合以高含量的草本花粉和外来乔木花粉(如松、冷杉等)为主。松属花粉在本研究区各植物群系下所有样品的孢粉组合中均有较高含量,说明松属花粉产量高、传播能力最强,具有超代表性;铁杉属和桤木属花粉也具有较强的传播能力,它们是外来花粉的主要成分,在不同植物群系下其花粉含量的高低与当地植被的郁闭度及当地植被花粉产量有关;桦木属和常绿栎类花粉代表性强,花粉产量高,也具有较强的传播能力;冷杉属花粉具有明显的低代表性,但其传播能力较强;杜鹃花科花粉的代表性较好,但传播能力差,其花粉散布范围基本在有母体植物生长的地方;柳属花粉代表性低,其花粉含量高于1.0%可能指示柳树的存在;单裂缝孢子的蕨类植物在本研究区部分样品中有较高含量,说明单裂缝孢子的蕨类植物比三裂缝孢子的蕨类植物可能更能适应海拔3 400 m以上的高海拔环境。     

The Paleovegetation and Paleoclimate during Time of Homudu People

The Homudu site was discovered closed to the northeastern part of the village Homudu in the Lochiang Commune of Yu-yao zhejiang sheng. The excavations were conducted in 1973–1974 and 1977–1978. Four layers of cultural deposits are super-posed one upon the other. The first and topmost layer corresponds to the middle layer at Sun-tse in Ch’ing-p’n Country, while the second layer represents the Ma-jla-bin Culture of Chiahsing Country. The third and fourth layers belong to a hitherto unknown culture, which has been provisionally named as the Homudu Culture. A palynologicai and paleobotanical study is made for showing the paleovcgetation and palcoclimate of that time. 1. The first sporo-pollen assemblage, obtaind from the fourth and lower part of third layers is marked by predominence of Gramineae pollen (11%–71%). The majority of Gramineae pollen is of large size. Exine is thin and more psilate. Annulus of the single pore is about 2.5–3.7 μm in wide. This kind of pollen very much resembles that of the recent rice (Oryza sativa). Besides that, the herbaceous pollen of Artemisia, Caulophyllum, Actinostemma, Polygonum, Rubia and some elements of Cyperaceae,Chenopodiaceae, Umbelliferae are noted. Pollen of trees and shrubs belong to Quercus, Castanopsis, Liquidambar, Mallotus, etc. Fern spores are of Lygodium. Ophioderma, Lemmaphyllum, Microsorium and Pyrrosia. A lot of plant remains (leaves and seeds) have been observed in the fourth layer: Chloran- thus fortunei, Magnolia coco? Phoebe sheareri, Lindera glauca, Camptotheca acumninata Prunus davidiana, Choerospondias axillaris, Trapa bispinosa, Quercus myrsinaefolia, Cas- tanopsis tibetana, etc. Sporo-pollen assemblage and plant remains reflect a subtropical everygreen and deciduous broad-leaved mixed forests flourished on mountain slopes and some rice fields scattered in the plains around the site. The climate of that time was warm and humid and rather warmer than that of the present. Radiocarbon tests of an acorn and a wood fragment unearthed from the fourth layer reveal that they date from about seven thousand years ago, corresponding to the Atlan- tic period of Europe. 2. The second sporopollen assemblage, obtained from the middle and the upper parts of third layer is predominent by pollen of trees, such as Quercus spp., Liquidambar formosana and others. In herbs, the pollen of Gramineae, Cypcraceae and water plants deereased, but those of Artemisia and Chenopodiaceae increased. The climate than was warm but somewhat drier than that of previous period. 3. The sporo-pollen assemblages from the second and the first layers were mainly composed of temperate elements, such as Quercus, Ulmus, Salix, Where the subtropical elements Liquidambar and Altingia decreased or completely disappeared.. These indicate that the climate become cooler than that of the previous periods. Radiocarbon tests of a wood fragment obtained from the second layer reveal that its date was from 5600 years ago, then the age of the second layer would correspond to the Subboreal period of Europe. A lot of rice pollen grains, along with a large amount of the remains of rice busks were obtained from the fourth layer in the site, which indicats that the rice plants were cultivated in China at least seven thousand years ago.     

滦河流域及周边地区花粉与植被关系的研究

滦河流域及周边地区空气中的花粉组合基本上反映了当地的植被面貌, 与植物的花期相对应, 乔木植物的花期多在春季, 草本植物的花期多在夏秋季, 该地冬季基本上无植物开花, 冬季空气中的花粉应是当年或多年春、夏、秋季散落在地上又被风吹到空气中的表土花粉和一些外来花粉 ;表土花粉随海拔高程的降低依次出现山地草甸、针阔混交林或以针叶树为主的针阔混交林、山地灌草丛、滨海平原草甸和滨海草甸或滨海盐生草甸的等 5个花粉组合带, 分别为蒿 (Artemisia)唐松草 (Thalictrum)松(Pinus)桦 (Betula) 孢粉组合带、松桦栎 (Quercus) 蒿孢粉组合带、藜科 (Chenopodiaceae)蒿禾本科 (Gramineae)中华卷柏 (Selaginellasinensis) 孢粉组合带、藜科蒿菊科 (Com positae)香蒲 (Typha) 孢粉组合带和藜科蒿莎草科 (Cyperaceae)禾本科孢粉组合带 ;由于花粉在河水中是以悬移质颗粒被搬运, 因此在不同时期沉积物花粉组合存在着一定的差异, 其中洪水期间河水对孢粉的分选作用最为明显。     

Characteristics of the aeropollen dynamics at several localities in Spain

In the Iberian Peninsula and Balearic Islands there are several bioclimatic regions which result in many different and fragmentary landscapes. Results obtained from monitoring 18 localities in Spain allow an outline of the aeropollen dynamics of some of these environments. The major pollen types are: I. trees: Pinus, Qvercus, Olea, Cupressaceae, 2. shrubs: Corylus, Pislacia and 3. herbs: Poaccae, Urticaceae, Chenopodiaceae-Amaranthaceae, Plantago and Asteraceae. In some locations Alnus, Artemisia, Buxus, Betula, Castanea, Eucalyptus, Populus, Ulmus, and Ephedra also have to be considered. Quantitatively, pollen from trees predominates over that from herbs and shrubs, although at some localities this trend varies. The first pollen grains found in the weekly atmospheric analyses are from trees (winter). In spring tree pollen grains share the atmosphere with herb pollen. For this reason it is usual to find peak values of pollen content in the atmosphere during this part of the year, while winter and autumn are characterized by low pollen counts.     

贵州白鹇湖沉积物中孢粉记录的5.5kaB.P.以来的气候变化

通过对白鹇湖沉积物柱芯孢粉组合的剖面变化分析,在有机质14C定年基础上,探讨了白鹇湖地区过去5.5 ka calB.P.以来的植被演替和气候变化过程.研究结果表明,5500-4500 aB.P.期间,各类植被比较丰富,气候温暖湿润;4500-2750 aB.P.期间,干旱草本和蕨类植物开始出现,是气候转变过渡期;2750-1500 aB.P.期间,木本植物组合类型发生明显变化,喜湿草本减少,耐旱草本增加,气候向温凉干旱化发展;1500 aB.P.至今,木本植被和喜湿草本继续减少,中生耐旱草本和蕨类植物数量继续大幅增加,干旱化趋势明显,植被组合向典型石漠化植被组合类型发展.白鹇湖沉积物剖面孢粉组合变化表明,该地区近5000a来气候变化以温度下降、降水减少为主要趋势,并存在明显的陆地植被退化现象.研究还揭示了自然气候变化事件(如气候持续干旱)可导致喀斯特地区发生石漠化,证实了喀斯特地区生态环境具先天脆弱性.科学评估白鹇湖地区气候干旱化趋势及其生态环境影响对指导该地区科学应对气候变化具重要意义,亟待加强.     

Pollen analysis of modern tree bark samples from the Manendragarh Forest Range of the Koriya district,Chhattisgarh, India

Pollen from modern tree bark samples collected in the Manendragarh Forest Range, Koriya District, Chhattisgarh, India, was investigated with the objective to understand the pollen rain in and around the study area, using modern tree bark samples as a new tool. The palyno-assemblages revealed the dominance of non-arboreals (herbs) over arboreals (trees and shrubs). Trees constitute an average of 17.23% pollen in the total pollen rain, whereas the average contribution of shrubs is only 0.33%. The non-arboreals have an average of 82.44% pollen in the total pollen rain. This bias in the form of representation of trees and shrubs, despite their ample presence in the forest, could be due to the differences in pollen production, dispersal and preservation of taxa, which depends on plant species and climatic conditions.     

南京紫金山山麓湖泊表层沉积物的孢粉分析*

本研究对南京紫金山山麓玄武湖和前湖共17个湖泊表层沉积物样品进行孢粉分析,旨在获得花粉组合新数据,揭示湖泊表层花粉与现生植被的关系,为开展第四纪湖泊地层孢粉学研究提供现代过程的参考.研究结果表明,花粉组合在剔除非自然分布的种植木本植物干扰花粉外,主要以木本花粉的松属(Pinus)和栎属(Quercus)占优势,其他常见木本植物有枫香属(Liquidambar)、栗属/栲属/柯属(Castanea/Castanopsis/Lithocarpus)、漆树科(Anacardiaceae)等.表土孢粉谱反映了研究区周边山地丘陵针叶、常绿-落叶阔叶林为主的植被,与现生北亚热带森林植被面貌对应良好.此外,陆生草本植物花粉主要是禾本科(Poaceae)和蒿属(Artemisia),湖泊湿生植物如水鳖属(Hydrocharis)、莎草科(Cyperaceae)等含量丰富.玄武湖与前湖中不同样点的主要木本植物花粉表现出优势类型高度一致、含量稳定、受采样位置的影响较小、指示地域植被类型的代表性较强等特点.通过对比南京地区已发表的紫金山及周边丘陵林下表土花粉分析结果,发现湖泊表土花粉组合受局部生境影响小,百分比含量较稳定,故其花粉信息在反映区域地带性植被的稳定性和可靠性方面更具优势.因此,开展表土花粉研究时应优先选择湖泊表层沉积物,藉此可提高基于孢粉重建区域地带性植被的代表性与可靠性.本研究结果为长江下游地区,尤其是南京地区解释第四纪孢粉组合与植被的对应关系提供了新数据和必要的现代过程参考.     

Recovery of pristine boreal forest floor community after selective removal of understorey, ground and humus layers

In boreal spruce forests that rarely experience extensive disturbances, fine-scale vegetation gaps are important for succession dynamics and species diversity. We examined the community implications of fine-scale gap disturbances by selective removal of vegetation layers in a pristine boreal spruce forest in Northern Finland. The aim was to investigate how the speed of recovery depends on the type of disturbance and the species growth form. We also wanted to know if there appeared changes in species composition after disturbance. Five different treatments were applied in the study: Control, removal of the ground layer (bryophytes and lichens), removal of the understorey layer (dwarf shrubs, herbs and graminoids), removal of both the ground and understorey layers, and complete removal of the vegetation and humus layers above the mineral soil. The vegetation recovery was monitored in terms of cover and species numbers over a 5-year period. Understorey layer cover, composed mainly of clonal dwarf shrubs, recovered completely in 4 years in treatments where the humus layer remained intact, whereas ground layer cover did not reach the control level in plots from where bryophytes and lichens were removed. Recovery was faster in terms of species number than species cover. Bryophytes, graminoids and dominant dwarf shrubs appeared in all disturbed plots quickly after disturbance. Seedlings of trees appeared exclusively in disturbed plots. Graminoids dominated after the removal of humus layer. The results indicate that the regeneration of forest floor after small gap disturbance occurs mainly by re-establishment of the dominant species. Although destruction of the humus layer leaves a long-lasting scar to the forest floor, exposing of mineral soil may enhance the sexual reproduction of dominant species and the colonization of weaker competitors.