黔中贵阳乌当地区下-中奥陶统的牙形刺生物地层

奥陶纪时期紧邻黔中古陆南缘的贵阳乌当发育有富含牙形刺的碳酸盐岩沉积,但生物地层研究非常薄弱。本文对乌当地区小谷龙剖面和豹子窝剖面的奥陶系进行牙形刺生物地层研究,共识别出10个牙形刺带,自下而上为Chosonodina herfurthi带、Drepanodus arcuatus带、Triangulodus bifidus带、Serratognathus diversus带、Oepikodus evae带、Baltoniodustriangularis带、Baltoniodusnavis带、Baltoniodusnorrlandicus带、Lenodusantivariabilis带和Dzikodustablepointensis带,时代属于下奥陶统特马豆克阶至中奥陶统达瑞威尔阶,可与华南同期地层的化石带对比。根据牙形刺生物地层,本研究构建了乌当地区奥陶系的年代地层格架,并限定黔中古陆南缘开始暴露出水面的时间为不晚于中奥陶世达瑞威尔期早期。     

湖南益阳早奥陶世弗洛期的笔石

本文报道湖南益阳南坝剖面早奥陶世弗洛期笔石16属30种,描述其中具有重要地层意义的笔石10属16种:Acrograptus affinis,A.gracilis,Baltograptus geometricus,B.vacillans,Clonograptus rigidus,Corym-bograptus deflexus,Cymatograptus undulatus,Didymograptellus bifidus,"Didymograptus"demissus,"D."rigo-letto,Expansograptus hol mi,E.constrictus,Pendeograptus fruticosus,P.pendens,Tetragraptus akzharensis,T.approxi matus。南坝剖面弗洛期笔石动物群以均分笔石为主,同时含有少量反称笔石。该地区弗洛期笔石动物群的分异度、丰度均较高,展示了笔石演化辐射早期斜坡相沉积环境下的笔石动物群特征。本文研究为全面理解奥陶纪笔石的演化辐射积累了重要资料。     

湖南早奥陶世弗洛期的笔石新属——挪威笔石属(Norvegiograptus gen.nov.)

报道产自湖南益阳南坝剖面下奥陶统弗洛阶的笔石新属——挪威笔石属(Norvegiograptus gen.nov.),该属归属于均分笔石科(Dichograptidae)对笔石亚科(Didymograptinae),主要特征是第一个胞管(th11)自胎管中部生出,始端发育型式为artus型,笔石体两枝纤细、下斜至下曲伸展。属内包含两个种:Norvegiograptus liber(Monsen)和Norvegiograptus enshiensis(Ni)。     

湖北兴山古洞口下奥陶统几丁虫

系统分析兴山古洞口下奥陶统,包括南津关组、分乡组、红花园组及大湾组底部的几丁虫微体生物化石。鉴定几丁虫5属14种,含早奥陶世特征种Eremochitina baculata,早中奥陶世常见种Conochitina decipiens,以及华南上扬子区早奥陶世常见属种Euconochitina fenxiangensis和Lagenochitina chongqingensis。E.baculata是北冈瓦纳地区弗洛早中期的特征带化石,本文的材料为该种在华南的首次确切报道,且产出层位略低,为特马豆克期。在华南,常见于特马豆克晚期的带化石Euconochitina symmetrica在该剖面的同时段地层中暂未被发现,但形态上与之高度相似的Euconochitina fenxiangensis在该时段大量出现。比较以往发表的数据后发现,E.fenxiangensis在华南的分布广泛且产出层位较为稳定,较之E.symmetrica更易获得,或可将其作为特马豆克晚期至弗洛初期区域内地层对比的一个有效卡尺。     

志留纪笔石Oktavitesexcentricus(Bjerreskov,1975)的古生物学特征及地层意义*

陕西紫阳地区在古地理上位于扬子台地西北缘,区域内志留纪地层发育,其中志留系兰多维列统(Llandovery)特列奇阶(Telychian)笔石相地层出露较为完整,笔石带较为连续。Oktavites excentricus(Bjerreskov,1975)是特列奇阶Oktavites spiralis笔石带重要的伴生分子,在世界范围内广泛分布。文中通过对紫阳地区一系列特列奇阶剖面研究后认为,Oktavites excentricus较短的化石延限和广泛的古地理分布使其具备地层对比潜力,这将为Oktavites spiralis笔石带的进一步细分提供依据。同时,Oktavites excentricus胞管为典型的奥氏笔石式,其笔石体的盘旋方式又与稍晚的Cyrtograptus lapworthi(Tullberg,1883)(笔石带化石,弓笔石类的早期代表)十分接近,因此,对该种的演化过程的研究也为讨论弓笔石的起源过程提供了更多参考。     

奥陶-志留系界线地层生物带的全球对比

详述扬子区晚奥陶世晚期(阿什极期)至兰多维列世初期的生物带序列,修正部分的笔石带,与全球各重要地区进行对比.研究表明,全球奥陶系顶部赫南特亚阶及其上下地层在我国的扬子区发育最好,地层连续发育并广泛出露,生物带发育齐全,不但有连续的笔石带,而且有世界广布的赫南特贝腕足动物群,有望在我国建立赫南特亚阶的全球层型剖面和点(GSSP).     

黔南三都Etagraptus approximatus笔石带的疑源类化石

本文描述黔南三都ＥＷｔａｇｒａｐｔｕｓ ａｐｐｒｏｘｉｍａｔｕｓ笔石带中的疑源类化石２３属３８种,包括１新种及２未定神。这为建立阿化克期早期笔石带中的疑源类组合,提高疑源类在地层划分对比中的精度提供了依据。     

奥陶—志留系界线地层生物全球对比

详述扬子区晚奥陶世晚期（阿什极期）至兰多维列世初期的生物带序列,修正部分的笔石带,与全球各重要地区进行对比,研究表明,全球奥陶系顶部赫南特亚阶及其上下地层在我国的扬子区发育最好,地层连续发展育并广泛出露,生物带发育齐全,不但有连续的笔石带,而且有世界广布的赫南特贝腕足动物群,有望在我国建立赫南特亚阶的全球层型剖面和点（ＧＳＳＰ）。     

加拿大北极地区泥盆纪叶肢介修订及兼论早期李氏叶肢介类(leaiins)的形态特征

湘南地区在晚奥陶世发育深水坳陷盆地相沉积, 完整记录了晚奥陶世桑比–凯迪期笔石页岩相生物地层序列。本论文在湘南地区祁东双家口 B 剖面开展笔石化石系统采集和生物地层学研究, 针对特征的江西笔石 (Jiangxigraptus)开展研究, 根据最新的分类学方案鉴定出江西笔石 1 属 6 种, 包括 Jiangxigraptus alabamensis (Ruedemann)、 Jiangxigraptus divaricatus (Hall)、 Jiangxigraptus gurleyi (Lapworth)、 Jiangxigraptus intortus (Lapworth)、Jiangxigraptus sextans (Hall)和 Jiangxigraptus vagus (Hadding)。此外, 本文还讨论了该属在全球范围内的分布, 认为在华南等地, Jiangxigraptus 可作为全球上奥陶统底界笔石生物地层对比的辅助属, 对识别上奥陶统桑比–凯迪阶的地层具有重要意义。     

安徽石台下、中奥陶统紫台组的牙形刺

下、中奥陶统紫台组以紫红色泥质灰岩、瘤状泥质灰岩为特征,具有稳定的空间分布范围,沿扬子台地东南缘呈条带状分布,与扬子台地上发育的湄潭组、大湾组等大致同期。作者研究了安徽石台栗阳柳树亭剖面紫台组的牙形刺,共描述19属23种,识别出四个牙形刺带,从下至上分别是Oepikodus evae带、Baltoniodus triangularis带、Baltoniodus navis带和Paroistodus originalis带,并可与同期笔石带对比。除扬子区外,紫台组还可与欧洲、阿根廷前科迪勒拉等地区的同期地层对比。     

The messaoudensis-trifidum acritarch assemblage (Ordovician: late Tremadoc-early Arenig) of the Barriga Shale Formation, Sierra Morena (SW-Spain)

The "messaoudensis-trifidum acritarch assemblage" is currently considered to be characteristic of latest Tremadoc-earliest Arenig cold-water environments on the periphery of Gondwana, at high latitudes in the southern hemisphere. An integrated biostratigraphical study on both acritarchs and graptolites was until now only available for localities in northwest England. Reinvestigation of the "messaoudensis-trifidum acritarch assemblage" from the Barriga Formation (Sierra Morena, southwestern Spain), which contains some graptolite horizons that can be attributed to the latest Tremadoc (pre-phyllograptoides and pre-approximatus graptolite biozones), strengthens the biostratigraphical potential of the "messaoudensis-trifidum assemblage", and the importance of some of the acritarch taxa recovered from these levels. It is concluded that the acritarch genera Coryphidium Vavrdová, 1972, Peteinosphaeridium Staplin et al., 1965 emend. Playford et al., 1995, Striatotheca, Burmann, 1970, and the Veryhachium lairdii group (rectangular veryhachiids) appear in the latest Tremadoc, and should not be considered as indicators of the base of the Arenig, as previously suggested.     

Horizon of the oldest known bryozoans (Ordovician)

Among the six species of fossil bryozoans described by Xia et al. [Xia, F.S., Zhang, S.G., Wang, Z.Z., 2007. The oldest bryozoans: new evidence from the late Tremadocian (Early Ordovician) of East Yangtze Gorges. Journal of Paleontology 81 (6), 1308–1326] from the Fenghsiang Formation of the Chenjiahe section of Yichang and the Guanzhuangping section, Liujiachang Town of Songzi, the Yangtze Gorges, Nekhorosheviella nodulifera occurs in the pristinus Conodont Subzone (lower Subzone of deltifer Conodont Zone) and the five other species are associated with the conodont Paltodus deltifer deltifer and the graptolites Acanthograptus sinensis and A. erectoramus. Based on the conodonts and graptolites, this bryofauna stratigraphically corresponds to the A. sinensis Graptolite Zone or P. deltifer Conodont Zone. The bryofauna is of Tremadocian Age (Early Ordovician) and represents the oldest bryozoans so far as known.     

Tremadocian (Ordovician) biostratigraphy and graptolites at Dayangcha (Baishan,Jilin, NE China)

Based on a systematic study of Tremadocian graptolites from the Early Ordovician Yehli Formation at Dayangcha, Baishan of Jilin, NE China, a revised graptolite zonation is proposed for the Tremadocian rocks in the area. In ascending order, the graptolite zones include theRhabdinopora flabelliformis parabola Zone, theAnisograptus matanensis Zone, thePsigraptus jacksoni Zone and theAorograptus victoriae Zone. The first three zones are discussed with their definitions and durations clarified, whereas the last zone is introduced for the first time as a replacement of the formerAdelograptus-Clonograptus Zone. The entire Yehli Formation is considered to be older than the late Tremadocian Hunneberg Substage of Scandinavia, the latter being represented by the carbonate-dominated basal part of the overlying Liangchiashan Formation where no graptolites have so far been discovered. The Tremadocian succession in the Dayangcha area is regarded as being representative of the North China (Sino-Korea) Palaeoplate, and a correlation of the Tremadocian graptolite sequence with those of other continents is suggested. Nine species belonging toDendrograptus, Aspidograptus, Callograptus andAirograptus of the family Dendroidae, and eight species ofPsigraptus, Adelograptus, Aorograptus andKiaerograptus of the family Anisograptidae are described and some are revised in the present paper. Among the important revisions are the reconstructed proximal development ofPsigraptus jacksoni based on partially three-dimensional specimens from the area, and the complex thecal structures ofAspidograptus.      

江西崇义早奥陶世茅坪组的笔石

记述早奥陶世茅坪组笔石9属25种,其中2新种.根据一些新材料的发现和笔石群的详细研究,将茅坪组笔石划分为3个带,即上部的Adelograptus-Clonograptus 带,中部的Aletograptus-Trograptus带,下部的Staurograptus-Anisograptus带.     

Restudy of some Ordovician–Silurian boundary graptolites from Anticosti Island, Canada, and their biostratigraphic significance

The Ellis Bay Formation on Anticosti Island has long been recognized for the biostratigraphic importance of its latest Ordovician conodont, palynomorph, and shelly fossil assemblages. However, a sparse record of graptolites has made it difficult to correlate these assemblages with the graptolite biozonation. Restudy of some previously described and examination of newly collected normalograptid specimens from the Ellis Bay and lower Becscie formations, however, shows that biostratigraphically important taxa are present and these species provide important constraints on the age of the strata. Normalograptus parvulus and N. minor have been identified in the upper half of the Ellis Bay Formation, suggesting that these strata are late Hirnantian in age ( N. persculptus Biozone). Normalograptus imperfectus and Normalograptus sp. aff. N. acceptus occur in the basal beds of the Becscie Formation, indicating that these strata are earliest Silurian. These graptolite data support the hypothesis that the positive carbon isotope excursion seen in the uppermost strata of the Ellis Bay Formation is isochronous with that seen in the Hirnantian strata at Dob's Linn, Scotland, and does not span much of the lower to mid-Hirnantian, as is the case in Arctic Canada and Nevada, USA.     Anticosti Island , Hirnantian, graptolites, Ordovician, Silurian.     

Ecostratigraphy, zonation and global correlation of earliest Ordovician planktic graptolites

The succession of earliest Ordovician planktic graptolites (graptoloids) in 11 key sequences around the world is reviewed and used to assess their distribution in depth facies, in paleolatitude and in time. An ecostratigraphic model shows the pattern of distribution of species across the shore-to-ocean profile in time. Fluctuations in this pattern are related to eustatic change; the Acerocare, Black Mountain, Ceratopyge and possibly, the Peltocare Regressive events are recognized. Surface-water temperature is found to have had little significant effect on distribution of Early Tremadoc species, and once the effects of facies is allowed for, their biostratigraphic utility can be assessed. A global suite of four chronozones is proposed for the Early Tremadoc and a further five zones comprise a provisional suite for the Late Tremadoc. The nine zones enable precise correlation of Tremadoc graptolitic sequences. The zones replace the previously used graptolite assemblages, are correlated with key conodont horizons and are, in upward sequence, the Zones of Rhabdinopora praeparabola, R. flabelliformis parabola, Anisograptus matanensis, R. f. anglica, Adelograptus, Paradelograptus antiquus, Kiaerograptus, Araneograptus murrayi-Araneograptus pulchellus , and Hunnegraptus copiosus.     

The base of the Middle Ordovician in China with special reference to the succession at Hengtang near Jiangshan, Zhejiang Province, southern China

Extensive work during the last decade has led to the recent decision by the International Subcommission on Ordovician Stratigraphy that the base of the Middle Ordovician Series should be placed at the base of the Baltoniodus triangularis Conodont Biozone in the Huanghuachang GSSP (Global Stratotype Section and Point) section near Yichang, Hubei Province, China. A review of the biostratigraphy of successions in many parts of China shows that for various reasons, it is currently difficult to recognize the precise boundary level in many regions, and additional studies are clearly needed. A newly exposed, previously poorly known, condensed section in deeperwater facies at Hengtang near Jiangshan in the Chiangnan (Jiangnan) belt has yielded a substantial number of important graptolites and conodonts through the boundary interval. It provides a more informative illustration of the relations between the ranges of several key taxa of these groups than any other section in China, and the level of the base of the Middle Ordovician appears to be within an about 1.8 m thick interval of Ningkuo Formation, and lies in the lower Isograptus caduceus imitatus Biozone.     

重庆城口大塘口下奥陶统红花园组和湄潭组下部的几丁虫

文中图示和描述了重庆城口大塘口奥陶系红花园组和湄潭组下部营盘段的几丁虫计 6属, 17种。其中包括 7个未定种和 3个新种 (Conochitinadatangkouensissp. nov., C. wengxigouensissp. nov., Rhabdochitinachenk ouensesp. nov. )。讨论了几丁虫的地理和地层分布特点及其与笔石生物带的关系,阐明了红花园组和湄潭组下部营盘段不同笔石带(A. filiformis, D. bifidus和A. suecicus笔石带)中几丁虫的组合特征,并与国外同期地层几丁虫组合进行了对比。     

东秦岭下志留统的笔石

笔石采自河南淅川下志留统张湾组,其时代与华中区龙马溪组相当。包含Ｄ．ｔｒｉａｎｇｕｌａｔｕｓ,Ｏ．ｃｏｍｍｕｎｉｓ和Ｍ．ｓｅｄｇｗｉｃｋｉｉ３个笔石带,可以与国内外同时代笔石带相对比。共记述正笔石目２科９属１５种,其中含２新种和１新亚种。     

Revision of the Zone 13 graptolite biostratigraphy in the Marathon, Texas, standard succession and its bearing on Upper Ordovician graptolite biogeography

One of the long-standing problems in North American graptolite biostratigraphy is the distinct differences in assemblages of post-Climacograptus bicornis age between the classical graptolite sequences in the New York - Quebec and Marathon, west Texas, regions. These have been attributed either to faunal provincialism or to the presence of a major hiatus between the Woods Hollow and Maravillas formations in Texas. New collections from the key Marathon Picnic Grounds section contain diagnostic Late Ordovician graptolites that confirm the existence of a major stratigraphic gap below the Maravillas Formation. The lower Maravillas Formation (Zone 13) has a Late Ordovician, low-diversity Pacific Province graptolite fauna that includes the biostratigraphically diagnostic species Climacograptus nevadensis, C. tubuliferus, Orthograptus fastigatus and Dicellograptus ornatus. Zone 13 graptolite assemblages from the Marathon region correlate with the C. tubuliferus to D. ornatus zones in the Trail Creek, Idaho, succession, the Ea4-Bo2 interval in Victoria, the O. fastigatus Zone in the Canadian Arctic Islands, the O. quadrimucronatus to D. ornatus zones of the Canadian Cordillera, and the D. complanatus to D. anceps zones in Scotland. The hiatus between the Woods Hollow and Maravillas formations spans an interval corresponding to at least the Eal Ea3 interval in Australia, the C. americanus to upper A. manitoulinensis zones in the New York - Quebec succession, and the D. clingani and P. linearis zones in Scotland. These results agree with the magnitude of the hiatus previously indicated by conodont biostratigraphy. Late Ordovician graptolite distribution patterns in North America can be explained by an extension of Cooper, Fortey & Lindholm's (1991; Lethaia 24) Lower Ordovician graptolite biofacies model into the Upper Ordovician, which incorporates both lateral water-mass specificity and depth stratification. Using this model, we recognize in Laurentia two separate biofacies among tropical-zone Late Ordovician Pacific Province graptolite faunas, a cosmopolitan Oceanic biofacies, and a cratonic Laurentian biofacies. The lower Maravillas Formation graptolite fauna is clearly part of the Oceanic biofacies, whereas the coeval Appalachian faunas represent the Laurentian biofacies. □Graptolites, Ordovician, biostratigraphy, Texas, biofacies, biogeography.