巴基斯坦盐岭地区二叠纪生物地层研究的新进展

近30年来,巴基斯坦盐岭地区的Wargal组和Chhidru组以及在喜马拉雅地区相当地层的地质时代被许多学者视为中二叠世(瓜达鲁普世)。根据华南与盐岭地区瓜达鲁普统和乐平统的牙形类化石带对比,可以确定Amb组和Wargal组底部的时代为瓜达鲁普世晚期,瓜达鲁普统与乐平统的界线位于Wargal组下部,吴家坪阶与长兴阶的界线位于Chhidru组的下部,而二叠与三叠系的界线位于Mianwali组下部的Kathwai段之内。我国西藏南部色龙群及相当地层含有与巴基斯坦盐岭地区Kalabagh段和Chhidru组可对比的乐平世腕足动物群以及二叠系—三叠系界线附近连续的牙形类化石带,由此可以推定色龙群的时代应为乐平世。     

Latest Guadalupian brachiopods from the Guadalupian/Lopingian boundary GSSP section at Penglaitan in Laibin, Guangxi, South China and implications for the timing of the pre-Lopingian crisis

A brachiopod fauna comprising nine species in eight genera from three closely spaced stratigraphic horizons of the same stratigraphic section is described for the first time from the Laibin Limestone in the uppermost part of the Maokou Formation in the Guadalupian/Lopingian (G/L) GSSP section at Penglaitan, Guangxi Autonomous Region, South China. The brachiopod assemblages are bracketed between two conodont zones: Jinogondolella xuanhanensis Zone below and Jinogondolella granti Zone above and, therefore, they can be safely assigned to the latest Capitanian in age. However, all but one of the nine brachiopod species from the Laibin Limestone carry strong early Lopingian (Wuchiapingian) aspect. Thus, the discovery of this brachiopod fauna not only suggests that some Lopingian brachiopod species had already appeared in the late Guadalupian (Capitanian); more importantly, it has also highlighted the fact that both the previously noted pre-Lopingian life crisis (or end-Guadalupian or Middle Permian mass extinction) and Lopingian recovery/radiation actually occurred in late Capitanian times, sometime before the G/L chronostratigraphic boundary. So far, the Penglaitan GSSP section provides the highest-resolution disappearance patterns of different fossil groups around the G/L boundary.     

Timing and progression of the end-Guadalupian crisis in the Fars province (Dalan Formation, Kuh-e Gakhum, Iran) constrained by foraminifers and other carbonate microfossils

The Middle-to-Upper Permian in the Kuh-e Gakhum anticline (southeastern Iran) has rarely been studied due to its structural complexity and the difficult access. Rich Permian fusulinid assemblages varying in age from Wordian to Changhsingian were found in a thick carbonate succession corresponding to the Dalan Formation. Three new species of foraminifers are described and a new biostratigraphic framework including five biozones is proposed. One of these, described and defined for the first time in the Dalan Formation, is based on the presence of Praedunbarula simplicissima n. gen. n. sp. When compared to the fossil content of existing bioprovinces, the floro-faunal biota of the Dalan Formation shows an affinity with Central and Western Tethys. A mass extinction of fusulinids and small foraminifers (70%) occurred concomitantly with the onset of a relative sea-level fall. This event led to a change in the carbonate factories from biologically induced carbonate production to ooid-rich chemically induced precipitation. The morphology of the platform at the Guadalupian/Lopingian transition evolves from a bioclastic ramp to a shelf. This transition is also characterized by a major sequence boundary and morphological anomalies in foraminifers. Therefore, as the regression and the changes in floro-faunal contents have been observed at the Guadalupian/Lopingian boundary, the extinction event is considered as end-Guadalupian. It is followed by a Lopingian transgression yielding renewed foraminiferal assemblages.     

The Guadalupian–Lopingian boundary (Permian) in a pelagic sequence from Panthalassa recognized by integrated conodont and radiolarian biostratigraphy

Guadalupian–Lopingian sedimentary rocks are widely distributed in accretionary complexes in Japan, but the Guadalupian–Lopingian boundary (G–LB) is not well documented from these pelagic sediments. To identify the G–LB and to better correlate an extinction event that occurred around the Guadalupian–Lopingian boundary, we examined the conodont biostratigraphy from a Permian pelagic chert sequence in the Gujo-hachiman section, Gifu, southwest Japan. Age-diagnostic conodonts, including Clarkina postbitteri postbitteri, were found in this section. The biostratigraphic occurrences of these age-diagnostic conodonts can pinpoint the “G–L transitional zone” in the Gujo-hachiman section by comparison with well-studied sections from south China, including the GSSP section. The transitional zone was recognized by the first occurrence horizons of both Clarkina postbitteri hongshuiensis and C. p. postbitteri. The G–LB has been placed at or above the first occurrence horizon of the radiolarians Albaillella yamakitai or Albaillella cavitata in previous studies from China and Japan. We detected the first occurrence horizon of A. yamakitai below the base of the “G–L transitional zone,” in the Upper Capitanian. The conodont biostratigraphy is consistent with the radiolarian biostratigraphy in this section, which can be correlated to relevant sections in China.     

Middle Permian smaller foraminifers from the Maokou Formation at the Tieqiao section,Guangxi, South China

This paper documents smaller foraminifers from the Maokou Formation (Guadalupian) at the Tieqiao section in Laibin area, South China. The assemblage of smaller foraminifers reported herein consists of 69 species of 42 genera. Although the Maokou Formation of this section was deposited in a relatively deep-water environment, smaller foraminifers are diverse and abundant, especially in debris flow deposits and thin turbidite beds. This fauna bears most similarity with coeval foraminiferal assemblages from the other Palaeotethyan regions. There is a sharp decrease in the diversity and abundance of smaller foraminifers at the lithological boundary between the Maokou and Heshan formations. An overwhelming majority of genera of smaller foraminifers that occur in the Maokou Formation at this section can be found in the Lopingian strata of South China. The study of smaller foraminiferal fauna of the Guadalupian at the Tieqiao section indicates a much less pronounced effect of the pre-Lopingian crisis on smaller foraminifers.     

广西来宾瓜达鲁普统—乐平统界线地层的牙形类化石——一个独立验证报告

王成源（Wang,2000）和金玉Gan（Jin,2000）提出了关于乐平统底界定义的不同论点。为了对此进行独立的验证,卡尔加里大学微体古生物实验室对采自华南广西来宾地区瓜达鲁普统－乐平统界线地层的牙形类化石大块样品进行了处理。结果如下：1.Jinogondolella和Clarkina在样品中没有共生;2.由Jinogondolella granti演化为Clarkina postbitteri的变化发生在层115－6i;3.C.postbitteri的首次出现层位（FAD）在蓬莱滩剖面位于层115－6i上部;4.这个变化标志着一个才的生物信号和一个重要的点断演化事件。Jinogondolella granti演化为Clarkina postbitteri的事件很可能是受瓜达鲁普统与乐平统之交的海平面降至最低水位所触发和控制。层115－6i下部为较浅水相颗粒灰岩,代表瓜达鲁普统末期层序高位域之顶或上覆层序低位域之底部。作为Clarkina属的第一个代表,Clarkina postbitteri是界线地层中最易识别的种之一,而Clarkina postbitterigkp带的识别较容易和稳定一致。Clarkina postbitteri和Clarkina dukouensis可以通过基于锯齿型式,居群和个体发育的分类体系来加以稳定地区分。Clarkina postbitteri和Clarkina dukouensis之间的过渡是渐变的,其间的分界点只能人为地确定,如果采用这样一个人为确定的点来定义全球界线层型（GSSP）,将难以稳定一致地对其加以识别。来自层114.6-6k的被Wang(2000)鉴定为Clarkina dukouensis的标本应当是C. postbitteri,或精确地可能是C. postbitteri的一个新亚种,没有争议的Clarkina dukouensis在蓬莱滩剖面最早出现于层114－7e。从生物演化和层序地层的角度看,C. postbitteri的首次出现层位（FAD）是瓜达鱼普统和乐平统界线在最清楚的侯选位置。     

Discovery of the Jehol Biota from the Celaomiao region and discussion of the Lower Cretaceous of the Bayingebi Basin,northwestern China

Some typical components of the Jehol Biota, including conchostracans Eosestheria sp., the mayfly Ephemeropsis trisetalis Eichwald, 1864, the aquatic beetle Coptoclava longipoda Ping, 1928, and a fragmentary dragonfly, are reported for the first time from the Bayingebi Formation in the Celaomiao region, western Inner Mongolia, China. This discovery indicates that the middle Upper Member of Bayingebi Formation can be correlated with the upper Yixian and the lower Jiufotang formations in western Liaoning Province. Combining it with the radio-isotopic dating result, we further believed that the Upper Member of Bayingebi Formation could be roughly correlated with the Yixian, Jiufotang, and Shahai formations, and the overlying Suhongtu Formation with the Fuxin Formation in western Liaoning Province.In the major Bayingebi Basin, palaeontological and radio-isotopic dating evidence shows that the Bayingebi Formation has a long depositional history of over 30 Ma: its Upper Member bearing the Jehol Biota and the early Fuxin Biota is probably coeval to the Yixian, Jiufotang and Shahai formations and has a Barremian–early Albian age; its Lower Member may be Berriasian–Hauterivian in age and could be correlated with the upper Tuchengzi, Zhangjiakou, and Dabeigou formations in northern Hebei Province. This suggests that the Bayingebi Formation should be promoted to the stratigraphic rank of group and subdivided into several secondary units (formations). Unlike the previous result, the Yingen Formation is considered across the Lower Cretaceous–Upper Cretaceous boundary and being late Albian–early Turonian in age.     

Changhsingian (Late Permian) foraminifers from the topmost part of the Xiala Formation in the Tsochen area,central Lhasa Block,Tibet and their geological implications

A foraminiferal fauna consisting of 15 genera and 19 species is documented for the first time from the topmost part of the Xiala Formation in the Aduogabu area in Tsochen County, central Lhasa Block. This foraminiferal fauna is dominated by Colaniella, indicating a broadly Changhsingian age. The presence of Colaniella, Reichelina and absence of the typical tropical fusuline genus Palaeofusulina in the fauna indicate that the Lhasa Block has not merged into equatorial regions during the Changhsingian. More importantly, the widespread Lopingian (Late Permian) marine carbonates with warm-water faunas and the stable platform carbonate sequence through the Guadalupian and Lopingian in the Lhasa Block present an obvious contrast against the contemporaneous sequences in the South Qiangtang Block and Gondwana margin. It suggests that both the Bangong-Nujiang Ocean and the Neotethys Ocean would have opened by the Guadalupian and Lopingian.     

Permian reefs re-examined: extrinsic control mechanisms of gradual and abrupt changes during 40 my of reef evolution

The evolution of Permian reefs is characterized by the following sequence of events: (1) Late Carboniferous–Cisuralian radiation, (2) early Late Cisuralian (Artinskian–Kungurian) turnover, (3) Guadalupian radiation, (4) end-Guadalupian crisis, (5) Lopingian radiation, (6) end-Lopingian crisis at the PTB (Permian–Triassic boundary), and (7) the at least 7 my (million years) metazoan reef gap during the Early Triassic. The early Late Cisuralian turnover and the end-Guadalupian reef crisis are gradual changes, while the end-Lopingian reef crisis represents an abrupt event. Lopingian reefs occur in a zone from 40 °N to 15 °S, Guadalupian reefs in an extended equatorial zone from 35 °N to 35 °S, and Lopingian reefs in a narrow equatorial zone of 20 °N and 20 °S. This pattern resulted from a network of global and regional control mechanisms including the assemblage of Pangea, the northward drift of continents, the opening of Neo-Tethys, and second-order sea level changes. The mechanism of the extinction has been intensely debated and a combination of the above mentioned long-term changes and abrupt ocean anoxia or hypercapnia (CO₂-poisoning) for the end-Guadalupian reef crisis is considered.     

Faunal migration into the Late Permian Zechstein Basin - Evidence from bryozoan palaeobiogeography

Late Permian bryozoans from the Wegener Halvø, Ravnefjeld and Schuchert Formations in East Greenland have been investigated. 14 genera are recognised.Integration of the new bryozoan data from the Upper Permian of East Greenland with data on the distribution of Permian bryozoans along the northern margin of Pangea is used to test hypotheses concerning Late Palaeozoic evolution of the North Atlantic region. During the Permian, the Atlantic rift system formed a seaway between Norway and Greenland from the boreal Barents Shelf to the warm and arid Zechstein Basin. This seaway is considered to be the only marine connection to the Zechstein Basin and therefore the only possible migration route for bryozoans to enter the basin. The distribution of Permian bryozoans is largely in keeping with such a connection from the cool Barents Shelf past the East Greenland Basin to the warm Zechstein Basin and also corroborates the change in temperature through this connection.     

Middle–Late Permian tetrapods from the Rio do Rasto Formation,Southern Brazil: a biostratigraphic reassessment

Dias‐da‐Silva, S. 2011: Middle–Late Permian tetrapods from the Rio do Rasto Formation, Southern Brazil: a biostratigraphic reassessment. Lethaia, Vol. 45, pp. 109–120. The Rio do Rasto Formation (Permian of Southern Brazil) was previously regarded as Guadalupian–early Lopingian age. Three tetrapod‐based localities are known: the Serra do Cadeado area, Aceguá and Posto Queimado. The latest tetrapod‐based biostratigraphic contribution considers that the Posto Queimado and Aceguá faunas are coeval and Wordian (middle Guadalupian) in age, correlated to the Isheevo faunas from Eastern Europe and to the Tapinocephalus Assemblage Zone of South Africa; whereas the Serra do Cadeado fauna is Capitanian (late Guadalupian), correlated to the Kotelnich fauna of Eastern Europe and, from bottom to top, to upper Pristerognathus, Tropidostoma and lower Cistecephalus assemblage zones of South Africa. A re‐evaluation of the tetrapods from the Rio do Rasto Formation and new fossil discoveries in the localities of Posto Queimado and Serra do Cadeado area (melosaurine and platyoposaurine temnospondyls, a basal anomodont, a dinocephalian and a basal dicynodont) supports a new tetrapod‐based biostratigraphic scheme for the Rio do Rasto Formation. Accordingly, the age of the fauna at Aceguá is late Roadian‐early Wordian, whereas the locality of Posto Queimado is late Wordian‐Capitanian. The Serra do Cadeado Area is correlated with both southernmost ones (Guadalupian) but also Wuchiapinghian (early Lopingian). □Paraná Basin, Passa Dois Group, tetrapod biostratigraphy, Western Gondwana.     

Flora of the Kazanian–Urzhumian boundary in the middle Permian of the Russian Platform

Rich fossil plant localities in the upper Guadalupian and the lowermost Lopingian of the Russian Platform are confined to several discrete “levels” or “horizons” divided by intervals that are almost devoid of plant remains. One such “horizon” is located at the boundary between the Upper Kazanian and the Urzhumian of the regional stratigraphic scale. Numerous localities of this level can be grouped into two geographical clusters, the northern one being confined to the Kama River Basin, and the southern one to the Orenburg Region and Southern Bashkortostan. Regarding the southern localities, three floras that are seemingly successive in time can be distinguished. Against a common background of articulates (Paracalamites and Equisetites), the lowermost flora is characterized by the dominance of leaves of Rufloria (Cordaitanthales) with very rare conifers. Conifers (Quadrocladus, Sashinia, Dvinostrobus) and leaves of Phylladoderma (Peltaspermales) are the most abundant elements of the middle assemblage, whereas Rufloria leaves are very seldom found together with the peltasperm Ginkgophyllum and the conifer-like Steirophyllum. The uppermost flora is dominated by Odontopteridium and Ustyugia, two closely related genera of peltasperms, whereas cordaitaleans are totally absent. Comparison with the northern localities, which can be linked to the type sections of the Upper Kazanian and the Urzhumian, enables the dating of these assemblages in terms of the regional stratigraphic scale. All three floras prove to be confined to the uppermost Kazanian, and only the youngest one could also occur in the lowermost Urzhumian. As the stratigraphic ranges of all observed plant taxa exceed the total stratigraphic interval under study, the sequence of the floras seems to be caused by ecological (most likely climatic) factors rather than the actual evolution of plants. In particular, the observed gradual elimination of cordaitaleans confirms the general view on the extinction of this group on the Russian Platform. The southwestern boundary of the occurrence of cordaitaleans on the Russian Platform stretched from the southwest to the northeast, approximately parallel to the palaeolatitudes reconstructed on the basis of palaeomagnetic data. During the Guadalupian it moved to the northeast, which is to the north considering the position of the North Pole of that time. Cordaitaleans were the main peat-forming plants in the Late Palaeozoic of northern Pangea (in the Kuznetsk, Tunguska and Pechora Basins). So their retreat to the north was most likely a consequence and a reflection of the warming and drying of the climate.     

Evolution of Permian conodont provincialism and its significance in global correlation and paleoclimate implication

The main components of Asselian through Artinskian conodont faunas found around the world are basically the same, and the provincialism is indicated only by less common endemic elements such as Gondolelloides and New Genus A Henderson in North Pangea, Sweetognathus bucaramangus around the equator and Vjalovognathus in eastern Gondwana. Provincialism is marked by differences at the species level of Mesogondolella, Neostreptognathodus and Sweetognathus during the Kungurian, and becomes very distinct with differences at the genus level during the Guadalupian and Lopingian. Three provinces of Permian conodonts, referred to as the North Cool Water Province (NCWP), the Equatorial Warm Water Province (EWWP) and the peri-Gondwana Cool Water Province (GCWP), are recognized. The NCWP is marked by Gondolelloides in the early Cisuralian, dominance of Neostreptognathodus and no or rare Sweetognathus in the late Cisuralian, dominance of Merrillina and Mesogondolella and absence of Sweetognathus in the Guadalupian, and dominance of Merrillina and Mesogondolella and absence of Iranognathus in the Lopingian. The EWWP is characterized by the absence of Gondolelloides and Vjalovognathus in the Cisuralian, abundance of Sweetognathus and Pseudosweetognathus in the Kungurian (late Cisuralian), Jinogondolella and Sweetognathus in the Guadalupian, and Clarkina and Iranognathus in the Lopingian. The GCWP is marked by Vjalovognathus, Merrillina in the Cisuralian, Vjalovognathus, Merrillina and Mesogondolella in the Guadalupian, and Vjalovognathus and Merrillina in the Lopingian. Mixed faunas are recognized in regions bordering between the EWWP and GCWP including Western Timor during the Artinskian, Pamirs during the Kungurian and the Salt Range during the Guadalupian and Lopingian.

Three different conodont zonations are proposed, one for each of the three conodont provinces. Four potential horizons for inter-provincial correlation of Permian conodonts are recognized. They are in ascending order: (1) the first appearance of Sweetognathus whitei, which is closely related to the last occurrence of Carboniferous-type conodonts such as Streptognathodus and Adetognathus; (2) the first appearance of Neostreptognathodus pequopensis; (3) the base of the Jinogondolella nankingensis Zone; and (4) the base of the Clarkina postbitteri–Iranognathus erwini Zone.

The spatial and temporal distribution pattern of Permian conodonts suggests that temperature is the primary controlling factor. Evolution of Permian conodont provincialism reveals a glaciation during the Asselian and Sakmarian, a global warming during the Artinskian, a climate cooling in North Pangea during the Kungurian, a continuation of Kungurian climate trends in the Guadalupian, a relatively minor warming during the Wuchiapingian, a returned cooling in the Changhsingian and Lower and Middle Griesbachian, and a global warming in the Late Griesbachian, which ended the Permian conodont lineage.     

Precocious sexual dimorphism and the Lilliput effect in Neo‐Tethyan Ostracoda (Crustacea) through the Permian–Triassic boundary

The Elikah River section spanning the Lopingian (Late Permian) to the Griesbachian (Early Triassic) time interval in the Central Alborz Mountains (north Iran) was sampled for ostracod analysis. We report 79 species distributed among 38 genera. Four new species are described: Acratia? pervagata Forel sp. nov., Microcheilinella alborzella Forel sp. nov., Basslerella superarella Crasquin sp. nov. and Cavellina nesenensis Crasquin sp. nov. The ontogeny of 13 species is described and sexual dimorphism in the genus Microcheilinella is here undoubtedly recognized for the first time. Six species show precocious sexual dimorphism of their carapace as early as A‐5 juvenile. The Lilliput effect is for the first time recorded and quantified for two species. Rare long‐time span Palaeocopida species, known throughout the entire Permian, document relatively long‐term evolution, including the size and growth rate modifications associated with the earlier appearance of carapace sexual dimorphism through time. These patterns might be related to the Guadalupian–Lopingian events and/or to climatic modifications occurring during the Permian interval.     

Carnian-Norian biomagnetostratigraphy at Silická Brezová (Slovakia): correlation to other Tethyan sections and to the Newark Basin

Correlations of Upper Triassic magnetic stratigraphies from Tethyan sections have been hampered by difficulties with conodont biostratigraphy and taxonomy, and discontinuous sedimentation, particularly in the ‘Hallstatt Limestones’ of Turkey and Austria. The magnetic stratigraphy and conodont biostratigraphy from the Upper Carnian to Upper Norian limestones exposed at Silická Brezová (Slovakia) can be correlated to other Tethyan sections and to the continental succession in the Newark Basin. The resulting correlations help to resolve some of the apparent discrepancies in existing conodont zonations, and result in a revised correlation to North American terrestrial vertebrate and palynological zones. The correlations imply that the Norian-Rhaetian boundary lies within Newark polarity zone E17r at ∼207 Ma. The Carnian-Norian boundary lies close to the base of Newark polarity zone E7r at ∼226 Ma. This implies durations for the Norian and Rhaetian stages of 19 Myr and 7 Myr, respectively.     

Late Triassic palynofloras in the Sichuan Basin,South China: Synthesis and perspective

Mesozoic strata are well developed and exposed continuously across the Sichuan Basin, South China. In particular, the Upper Triassic strata yield diverse and abundant spore-pollen fossils, providing a significant reference for the study of palaeoenvironmental variations across the Triassic–Jurassic transition where mass extinctions were occurring. In this paper, we summarize the major progress on Late Triassic palynological studies in this basin. To date, 151 genera (454 species) of sporomorph fossils have been reported from the Late Triassic strata in the Sichuan Basin. Three palynological assemblages are distinguished for the Late Triassic in the Sichuan Basin. Late Triassic vegetation in the Sichuan Basin shows a remarkable predominance of ferns, followed by conifers and cycads/ginkgophytes, and conifers show a distinct increase in abundance in the latest Triassic. In general, the Late Triassic palaeoclimate in the Sichuan Basin was tropical-subtropical, humid and warm. A synthesis of the data shows that the Late Triassic did not have a constant palaeoclimate in the Sichuan Basin, several climatic events are recognized: two warm and humid climate events in Norian-Rhaetian time, coupled with a cooler and drier condition in the latest Late Triassic. Further investigations in higher resolution at more continuous sections in the Sichuan Basin are needed to better understand the Late Triassic vegetation response, climate changes, as well as palaeoecosystem variations across the Triassic–Jurassic boundary.     

Discovery of the genus Crinoedischia in the Middle Permian of France (Insecta: Orthoptera)

New orthopteran Crinoedischia lapeyriei sp. nov. is described and illustrated from the Middle Permian (Guadalupian) of Lodève Basin in southern France. The new species is separated from the Lower Permian (Artinskian) Crinoedischia species by differences in wing venation pattern such as long ScP with anterior branches, RA with prominent distal branches, and MA ending with three branches. The significance and parallels of entomofaunas between Salagou and Wellington formations (Lodève, Elmo, Midco) are briefly discussed.     

Palaeogene marine stratigraphy in China

Palaeogene deposits are widespread in China and are potential sequences for locating stage boundaries. Most strata are non‐marine origin, but marine sediments are well exposed in Tibet, the Tarim Basin of Xinjiang, and the continental margin of East China Sea. Among them, the Tibetan Tethys can be recognized as a dominant marine area, including the Indian‐margin strata of the northern Tethys Himalaya and Asian‐margin strata of the Gangdese forearc basin. Continuous sequences are preserved in the Gamba–Tingri Basin of the north margin of the Indian Plate, where the Palaeogene sequence is divided into the Jidula, Zongpu, Zhepure and Zongpubei formations. Here, the marine sequence ranges from Danian to middle Priabonian (66–35 ma), and the stage boundaries are identified mostly by larger foraminiferal assemblages. The Paleocene/Eocene boundary is found between the Zongpu and Zhepure formations. The uppermost marine beds are from the top of the Zongpubei Formation (~35 ma), marking the end of Indian and Asian collision. In addition, the marine beds crop out along both sides of the Yarlong Zangbo Suture, where they show a deeper marine facies, yielding rich radiolarian fossils of Paleocene and Eocene. The Tarim Basin of Xinjiang is another important area of marine deposition. Here, marine Palaeogene strata are well exposed in the Southwest Tarim Depression and Kuqa Depression. They comprise mostly neritic and coastal lagoon facies of the Tethyan realm. Palaeontological evidence suggests that the Paleocene/Eocene boundary here is in middle of the Qimugen Formation. The Tarim Basin was largely drained by Late Oligocene. To the east, the marine offshore Palaeogene strata are widespread in the North Taiwan and East Zhejiang depressions of the continental shelf basin of East China Sea. Abundant fossils including foraminifera, calcareous nannofossils, ostracods, pollen and bivalves occur in the marine environment. Biostratigraphically, the sequence is well correlated with the international planktonic foraminiferal and nannofossil zonations.     

CALIBRATED DIVERSITY,TREE TOPOLOGY AND THE MOTHER OF MASS EXTINCTIONS: THE LESSON OF TEMNOSPONDYLS

Abstract: Three family‐level cladistic analyses of temnospondyl amphibians are used to evaluate the impact of taxonomic rank, tree topology, and sample size on diversity profiles, origination and extinction rates, and faunal turnover. Temnospondyls are used as a case study for investigating replacement of families across the Permo‐Triassic boundary and modality of recovery in the aftermath of the end‐Permian mass extinction. Both observed and inferred (i.e. tree topology‐dependent) values of family diversity have a negligible effect on the shape of the diversity curve. However, inferred values produce both a flattening of the curve throughout the Cisuralian and a less pronounced increase in family diversity from Tatarian through to Induan than do observed values. Diversity curves based upon counts of genera and species display a clearer distinction between peaks and troughs. We use rarefaction techniques (specifically, rarefaction of the number of genera and species within families) to evaluate the effect of sampling size on the curve of estimated family‐level diversity during five time bins (Carboniferous; Cisuralian; Guadalupian–Lopingian; Early Triassic; Middle Triassic–Cretaceous). After applying rarefaction, we note that Cisuralian and Early Triassic diversity values are closer to one another than they are when the observed number of families is used; both values are also slightly higher than the Carboniferous estimated diversity. The Guadalupian–Lopingian value is lower than raw data indicate, reflecting in part the depauperate land vertebrate diversity from the late Cisuralian to the middle Guadalupian (Olson’s gap). The time‐calibrated origination and extinction rate trajectories plot out close to one another and show a peak in the Induan, regardless of the tree used to construct them. Origination and extinction trajectories are disjunct in at least some Palaeozoic intervals, and background extinctions exert a significant role in shaping temnospondyl diversity in the lowermost Triassic. Finally, species‐, genus‐, and family trajectories consistently reveal a rapid increase in temnospondyl diversity from latest Permian to earliest Triassic as well as a decline near the end of the Cisuralian. However, during the rest of the Cisuralian family diversity increases slightly and there is no evidence for a steady decline, contrary to previous reports.     

准噶尔盆地侏罗纪介形类新知

准噶尔盆地侏罗纪介形类已由众多中外地层古生物工作者进行过数十年的研究并有系统成果问世。本文仅介绍两个侏罗系化石点,作为前人工作之补充,充实已有化石记录,也提出了进一步划分的建议和依据。