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热带森林植物多样性及其维持机制 总被引:20,自引:0,他引:20
热带森林具有地球上最丰富的植物多样性。关于热带森林植物多样性的维持机制,虽有众多假说,但均未形成完善的理论体系。不同的学者从不同的角度出发得出了许多结论,但也引起了不少争议。本文主要简述了4种经常被用来解释热带森林高植物多样性的机制:生态位分化、取食压力、生活史负相关和随机竞争,每一种机制都有大量的证据支持。热带森林植物沿微环境梯度的非随机空间分布表明其生态位分化很明显,并对其多样性起重要作用。动物的取食降低母树周围同种幼苗的生长率和存活率,为其他物种存活提供了机会,这就是取食压力假说,这是一个极有前景但仍需大量实验验证的假说。生活史负相关使得热带森林的许多植物能够共存。冠层植物的抑制使得随机性在林下植物的建立过程中起决定作用。 相似文献
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木质藤本是热带森林的一个重要组分, 直接或间接地影响着森林中树木的生长和更新, 改变森林树木的种类组成, 并且可以通过改变森林碳固定量等方式在生态系统水平上发挥作用。全球气候的变化, 以及热带森林片断化程度的加剧, 将很大程度上影响着木质藤本的多样性和丰富度, 其特殊的生物学特性将在森林动态中发挥更加重要的作用。本文结合国内外目前木质藤本研究现状, 概述了木质藤本的一般知识(包括木质藤本的定义和生物学特性等), 介绍了木质藤本全球分布格局、其多样性维持机理以及木质藤本在森林生态系统中的功能与作用, 并就存在的一些问题以及需进一步开展的工作展开了讨论。 相似文献
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木质藤本及其在热带森林中的生态学功能 总被引:5,自引:0,他引:5
木质藤本是热带森林的一个重要组分,直接或间接地影响着森林中树木的生长和更新,改变森林树木的种类组成,并且可以通过改变森林碳固定量等方式在生态系统水平上发挥作用。全球气候的变化,以及热带森林片断化程度的加剧,将很大程度上影响着木质藤本的多样性和丰富度,其特殊的生物学特性将在森林动态中发挥更加重要的作用。本文结合国内外目前木质藤本研究现状,概述了木质藤本的一般知识(包括木质藤本的定义和生物学特性等),介绍了木质藤本全球分布格局、其多样性维持机理以及木质藤本在森林生态系统中的功能与作用,并就存在的一些问题以及需进一步开展的工作展开了讨论。 相似文献
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从植物功能群角度探讨陆地生态系统功能与稳定性维持机理、植物对环境变化的适应与响应以及水分、养分利用效率等成为当前生态系统生态学研究的主要技术路线.植物功能群的提出和研究,为研究复杂的生态系统提供了一个良好的方法和途径.综述了陆地生态系统功能群方面的最新研究进展,介绍了植物功能群定义的发展历程,详细比较了植物功能群划分的依据及方法,对于植物功能群与群落稳定性之间的关系、植物功能群对群落生产力的影响以及植物功能群与环境因子的动态关系等进行了深入讨论.这些研究资料表明,植物功能群整合了功能及对环境响应相似的一类植物,但植物功能特征不是绝对的、单一的,所以对植物功能群就会有不同的理解,会有不同的定义及划分方法.许多研究者从不同的角度、尺度来对植物功能群进行研究,这些研究结果有不同的针对方向和目的,使人们可以从不同的角度更全面的理解复杂的陆地生态系统.学者们在研究生态系统时,或多或少地总要与植物功能群相联系,这大大拓宽了植物功能群的应用范围.所有前人的研究使植物功能群的概念、划分、方向、应用等诸方面越来越清晰.这要求应有一个规范、统一、明确的植物功能群研究方案,这样能使对植物功能群的研究更加深入,能整合全球所有植物功能群的相关研究. 相似文献
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中国热带森林水文生态功能 总被引:15,自引:2,他引:15
中国热带森林水文生态功能周光益(中国林科院热带林业研究所,广州510520)Eco_hydrologicalFunctionsofTropicalForest,China.ZhouGuangyi(TheResearchInstituteofTropi... 相似文献
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根据在鼎湖山季风常绿阔叶林集水区3年的观测结果,对这一南亚热带地带性植被的水文功能进行了较深入的探讨。由于林分结构复杂,季风常绿阔叶林对大气降水具有较高的林冠截留率,截留率的季节变化动态与降水量的季节分配有关。该森林土壤结构良好,其渗水速度快,加上有较厚的枯落物层,对于减缓地面径流,减少水土流失,功效甚大。但由于土层较薄,其蓄水功能有限。该森林的径流量占总降水量的比例特别是地表径流所占比例较小,分别为50.22%和14.19%。其季节分配有利于保持水土和植物生长。由于土壤蓄水量变化出现负值,用水量平衡法计算得出鼎湖山季风常绿阔叶林年蒸发散量为1323.98mm,占大气降水量的51.81%。该森林的降水中主要营养元素浓度较高,降水带来大量的养分,特别是氮素的年输入量高达35.29kg·hm-2。淋溶也给该森林带来较高的养分归还量,其中钾的淋溶量高达35.65kg·hm-2;镁的淋溶量高达8.9kg·hm-2,为林外雨输入的2.37倍。径流中的氮输出为28.35kg·hm-2,降水中的氮素净输入量为6.94kg·hm-2;但镁的净输入呈负值。 相似文献
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植物功能群(PFG)把一系列植物归为一个功能团体,可以作为植被随环境动态变化的基本单元,简化相关研究。植物功能群内不同物种的生理特性差异是功能群存在的基础,也影响着功能群整体的功能。采用种间联结法并结合优势种在海拔梯度上的变化,将伏牛山自然保护区森林生态系统各层优势种划分为4组植物功能群,研究每组植物功能群内不同物种的光合特性。对光合-光响应、最大净光合速率(Pn)、暗呼吸速率(Rd)、光补偿点(LCP)、光饱合点(LSP)和表观量子效率(AQY)等指标进行分析和比较,结果表明:植物功能群内不同物种的光补偿点和暗呼吸速率的变异系数最高分别达70.47%和74.29%,功能性状存在着显著的互补性,这是物种能长期共存的主要原因。乔木的最大净光合速率在7—8μmol CO_2m~(-2)s~(-1)之间,对高光强的适应能力较高。同一功能群内各层植物光饱合点在1200—1500μmol m~(-2)s~(-1)之间,差别不大。林下灌木、草本植物对弱光的利用效率较乔木高。同一功能群内物种功能性状的差异及不同植物功能群间整体性状的差异,影响着森林生态系统的结构、功能、过程和生态系统服务。 相似文献
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中国北亚热带次生森林植被研究述评 总被引:1,自引:0,他引:1
对我国北亚热带植被的研究,可追溯到本世纪二三十年代对南京森林植物的调查工作\"'和钟山森林之变迁的研究\"'。受战争的影响,约有20年的停动期。50年代后,研究工作得以继续,而且研究内容日益广泛,研究程度日益深入,这些工作大体可分为以下4个方面。1植被特征的研究曲仲湘'-\"利用LUtZ树木结构图解首次对北亚热带的南京灵谷寺、琅郎山、栖霞山森林和林木进行了研究,并依据树种的耐阴程度探讨了其可能的变化过程。文中所用的划分树木生态年龄为5个等级的方法,确立了我国森林树种\"年龄\"划分的标准。虽然,这种划分方法比较简化,但因… 相似文献
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Manfredo di Porcia e Brugnera Flicien Meunier Marcos Longo Sruthi M. Krishna Moorthy Hannes De Deurwaerder Stefan A. Schnitzer Damien Bonal Boris Faybishenko Hans Verbeeck 《Global Change Biology》2019,25(11):3767-3780
There is mounting empirical evidence that lianas affect the carbon cycle of tropical forests. However, no single vegetation model takes into account this growth form, although such efforts could greatly improve the predictions of carbon dynamics in tropical forests. In this study, we incorporated a novel mechanistic representation of lianas in a dynamic global vegetation model (the Ecosystem Demography Model). We developed a liana‐specific plant functional type and mechanisms representing liana–tree interactions (such as light competition, liana‐specific allometries, and attachment to host trees) and parameterized them according to a comprehensive literature meta‐analysis. We tested the model for an old‐growth forest (Paracou, French Guiana) and a secondary forest (Gigante Peninsula, Panama). The resulting model simulations captured many features of the two forests characterized by different levels of liana infestation as revealed by a systematic comparison of the model outputs with empirical data, including local census data from forest inventories, eddy flux tower data, and terrestrial laser scanner‐derived forest vertical structure. The inclusion of lianas in the simulations reduced the secondary forest net productivity by up to 0.46 tC ha?1 year?1, which corresponds to a limited relative reduction of 2.6% in comparison with a reference simulation without lianas. However, this resulted in significantly reduced accumulated above‐ground biomass after 70 years of regrowth by up to 20 tC/ha (19% of the reference simulation). Ultimately, the simulated negative impact of lianas on the total biomass was almost completely cancelled out when the forest reached an old‐growth successional stage. Our findings suggest that lianas negatively influence the forest potential carbon sink strength, especially for young, disturbed, liana‐rich sites. In light of the critical role that lianas play in the profound changes currently experienced by tropical forests, this new model provides a robust numerical tool to forecast the impact of lianas on tropical forest carbon sinks. 相似文献
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Aim To explore potential shifts in vegetation and fire regime in some dominant forest types in the north-eastern part of the Mediterranean basin under climate change.
Location Two altitudinal gradients in the continental part of Greece.
Methods We developed a forest gap dynamics simulator that provides feedback from the stand to its water balance and flammability status. The model is used to simulate vegetation dynamics in two mountainous areas, currently found under different aridity conditions. Two climatic change scenarios (Intergovernmental Panel on Climate Change A1 and B2) were applied to explore differences in the response of the established forest types. In addition we explicitly accounted for the role of fire, under both current and altered climate patterns.
Results Fire was identified to play a significant role in low-altitude sites. Its significance increased with the severity of the climate change scenario. Elevational shifts of the dominant species were simulated for each site, while in some cases these changes were associated with a shorter fire cycle and a frequent resetting of processes of vegetation change.
Main conclusions Our simulations suggest a greater vulnerability of mountainous Mediterranean drier areas regarding compositional alteration and flammability trends. Changes in vegetation could take place through both a discrete and synergistic realization of changes in the drought stress and fire frequency. 相似文献
Location Two altitudinal gradients in the continental part of Greece.
Methods We developed a forest gap dynamics simulator that provides feedback from the stand to its water balance and flammability status. The model is used to simulate vegetation dynamics in two mountainous areas, currently found under different aridity conditions. Two climatic change scenarios (Intergovernmental Panel on Climate Change A1 and B2) were applied to explore differences in the response of the established forest types. In addition we explicitly accounted for the role of fire, under both current and altered climate patterns.
Results Fire was identified to play a significant role in low-altitude sites. Its significance increased with the severity of the climate change scenario. Elevational shifts of the dominant species were simulated for each site, while in some cases these changes were associated with a shorter fire cycle and a frequent resetting of processes of vegetation change.
Main conclusions Our simulations suggest a greater vulnerability of mountainous Mediterranean drier areas regarding compositional alteration and flammability trends. Changes in vegetation could take place through both a discrete and synergistic realization of changes in the drought stress and fire frequency. 相似文献
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Lianas are an important growthform in tropical forests, and liana abundance and biomass may be increasing in some regions. Explanations for liana proliferation hinge upon physiological responses to changing resource conditions that would favour them over trees. Testing a chemical basis for such responses, we assessed 22 foliar traits in 778 lianas and 6496 trees at 48 tropical forest sites. Growthform differences in chemical allocation occurred on a leaf mass and area basis. Light capture-growth and maintenance-metabolism chemicals averaged 14.5 and 16.7% higher mass-based concentration in lianas than in trees globally, whereas structure and defence chemicals averaged 9.0% lower in lianas. Relative differences in chemical allocation by lianas and trees were mediated by climate with peak differences at about 2500 mm year(-1) and 25 °C. Differences in chemical traits suggest that liana expansion could be greatest in forests undergoing increased canopy-level irradiance via disturbance and climate change. 相似文献
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Florian Hofhansl Eduardo ChacnMadrigal ke Brnnstrm Ulf Dieckmann Oskar Franklin 《Ecology and evolution》2021,11(9):3856
Plant functional trait variation in tropical forests results from taxonomic differences in phylogeny and associated genetic differences, as well as, phenotypic plastic responses to the environment. Accounting for the underlying mechanisms driving plant functional trait variation is important for understanding the potential rate of change of ecosystems since trait acclimation via phenotypic plasticity is very fast compared to shifts in community composition and genetic adaptation. We here applied a statistical technique to decompose the relative roles of phenotypic plasticity, genetic adaptation, and phylogenetic constraints. We examined typically obtained plant functional traits, such as wood density, plant height, specific leaf area, leaf area, leaf thickness, leaf dry mass content, leaf nitrogen content, and leaf phosphorus content. We assumed that genetic differences in plant functional traits between species and genotypes increase with environmental heterogeneity and geographic distance, whereas trait variation due to plastic acclimation to the local environment is independent of spatial distance between sampling sites. Results suggest that most of the observed trait variation could not be explained by the measured environmental variables, thus indicating a limited potential to predict individual plant traits from commonly assessed parameters. However, we found a difference in the response of plant functional traits, such that leaf traits varied in response to canopy‐light regime and nutrient availability, whereas wood traits were related to topoedaphic factors and water availability. Our analysis furthermore revealed differences in the functional response of coexisting neotropical tree species, which suggests that endemic species with conservative ecological strategies might be especially prone to competitive exclusion under projected climate change. 相似文献
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Pedro H. S. Brancalion Ricardo A. G. Viani James Aronson Ricardo R. Rodrigues André G. Nave 《Restoration Ecology》2012,20(6):704-711
High‐diversity reforestation can help jumpstart tropical forest restoration, but obtaining viable seedlings is a major constraint: if nurseries do not offer them, it is hard to plant all the species one would like. From 2007 to 2009, we investigated five different seed acquisition strategies employed by a well‐established tree nursery in southeastern Brazil, namely (1) in‐house seed harvesters; (2) hiring a professional harvester; (3) amateur seed harvesters; or (4) a seed production cooperative, as well as (5) participating in a seed exchange program. In addition, we evaluated two strategies not dependent on seeds: harvesting seedlings from native tree species found regenerating under Eucalyptus plantations, and in a native forest remnant. A total of 344 native tree and shrub species were collected as seeds or seedlings, including 2,465 seed lots. Among these, a subset of 120 species was obtained through seed harvesting in each year. Overall, combining several strategies for obtaining planting stocks was an effective way to increase species richness, representation of some functional groups (dispersal syndromes, planting group, and shade tolerance), and genetic diversity of seedlings produced in forest tree nurseries. Such outcomes are greatly desirable to support high‐diversity reforestation as part of tropical forest restoration. In addition, community‐based seed harvesting strategies fostered greater socioeconomic integration of traditional communities in restoration projects and programs, which is an important bottleneck for the advance of ecological restoration, especially in developing countries. Finally, we discuss some of the limitations of the various strategies for obtaining planting stocks and the way forward for their improvement. 相似文献
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Tropical forests are among the most species-diverse ecosystems on Earth. Their structures and ecological functions are complex to understand. Functional group is defined as a group of species that play similar roles in an ecosystem. The functional group approach has been regarded as an effective way of linking the compositions of complex ecosystems with their ecological functions. To understand the variation of functional groups in species-rich ecosystems after disturbance, the present study investigated the spatial pattern and temporal dynamics of woody plants in a typically fragmented natural forest landscape of Hainan Island in South China. The study area was classified into eight landscape types based on vegetation type, disturbance manner and the time of recovery. The woody plant species were aggregated into seven functional groups based on the growth form, successional status and plant size. The results gained from the present study showed that all functional groups, except for the emergent and canopy tree species, were present in all eight landscape types. Each landscape type had different numbers of dominant functional groups. There are similar species richness and stem abundance structure among functional groups between mid-successional clear cut lowland rainforest and old growth tropical coniferous forest. This similarity exists in selective logged lowland rainforest and old-growth lowland rainforest, as well as among landscape types of montane rainforest. The functional groups with the same successional status had similar patterns of species richness and stem abundance ratios among different landscape types. The variation patterns of functional groups along the successional stages in terms of species richness and stem abundance among the tropical lowland rainforest landscape types were more similar to each other than those in the tropical montane reinforest landscape types. This study provides further support for the competition-colonization tradeoff and successional niche theory as opposed to models of neutrality and ecological equivalence. 相似文献
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热带次生林具有重要的物种保育和固碳功能, 然而高强度的干扰会导致次生林早期出现类似季雨林的阶段, 因而群落恢复速度和方向是当前热带生态学研究中最为关注的议题之一。该文以海南岛在刀耕火种弃耕地形成的不同演替阶段的次生林为研究对象, 比较森林不同恢复时间(12年、25年、55年)群落中的不同年龄(幼树、小树、成年树)个体与低地雨林老龄林的物种组成、多样性和群落结构差异, 探讨刀耕火种弃耕地恢复过程中的群落组配过程。首先, 在海南岛霸王岭林区内建立7个1 hm2(100 m × 100 m)的样地, 并调查样地内所有胸径≥ 1 cm的木本植物个体(不包括木质藤本)的种类、胸径大小和树高。无度量多维标定法(NMS)排序结果表明, 刀耕火种弃耕地恢复群落与老龄林的物种组成存在明显差异, 并且其物种组成差异随着径级增加而逐渐减小。刀耕火种弃耕地群落物种累积速度缓慢, 25年和55年恢复群落的种面积、种个体和种多度曲线无差异, 存在一个明显的停滞阶段。与物种组成相比, 群落结构恢复相对较迅速, 但仍没有形成老龄林阶段中的复杂结构。萌生个体在早期恢复群落中占有较高比例, 其个体密度和胸高断面积分别占总数的39.9%和55.9%, 但在恢复中后期迅速降低。刀耕火种弃耕地恢复群落中以先锋种和非先锋喜光种为主。虽然耐阴种随演替而逐渐增加, 但恢复中后期群落中的耐阴种重要值仅为老龄林的27.7%。这些结果表明, 虽然刀耕火种弃耕地恢复群落缓慢地逐渐接近最终恢复目标, 但仍然存在 很大的不确定性。刀耕火种弃耕地恢复过程中的异速恢复和停滞阶段需要纳入今后群落演替模型构建和森林固碳效益核甘共苦算中。 相似文献
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Abstract We expected that in clonal plants with the phalanx growth form, prereproductive ramets function as a meristem bank that helps buffer ramet demography against environmental fluctuations. Specifically, we evaluated four expectations for the forest understorey clonal palm Geonoma brevispatha in a swamp forest in south‐eastern Brazil. Our expectations were are follows: (i) the vital rates of prereproductive ramets are more responsive to genet‐related characteristics (genet size, distance and size of the nearest genet neighbour, and reproductive ramet survivorship) than to environmental variables (canopy openness, basal area of small trees, flooding, and distance and size of the nearest large tree neighbour), whose effects are expected to affect ramet dynamics indirectly through physiological integration; (ii) the progress of prereproductive ramets to later stages is dependent on the death of reproductive ramets; (iii) the population dynamics of G. brevispatha is resistant to significant interannual changes in the vital rates of ramets, (i.e. the occurrence of good and bad years); and (iv) there are large negative covariances among demographic transitions within a given year, as well as between years, reflecting the flexibility of G. brevispatha to adjust to environmental variability. Expectations 1, 2 and 4 were confirmed, while expectation 3 was only partially confirmed. Matrix models predicted that the growth rate of the overall ramet population was significantly greater than unity in the first study year, but it was significantly smaller than 1.0 in the second study year. Ramet population growth rates at the level of individual genets were greater than unity in the majority of genets in the first year, but in the second year genets with growth rates smaller than unity prevailed. Ramet population dynamics seems to be resistant to the succession of good and bad years; in effect, the presence of a meristem bank of ramets helps stabilize λ. 相似文献