High-resolution sequence stratigraphy of a mixed carbonate-siliciclastic, cratonic ramp (Upper Ordovician; Kentucky–Ohio, USA): insights into the relative influence of eustasy and tectonics through analysis of facies gradients

Detailed facies analysis and event stratigraphy of an Upper Ordovician (Rocklandian–Edenian) cratonic ramp succession in eastern North America yields insights into eustatically driven sequence architecture and localized tectonic instability. Seven, predominantly subtidal, mixed carbonate-siliciclastic depositional sequences (3rd order) are identified and correlated across the length of a 275-km ramp–to–basin profile. Within the larger depositional sequences (3rd order) at least two smaller orders (4th and 5th) of cyclicity are recognizable. Three systems tracts occur within each sequence (transgressive, TST; highstand, HST; regressive, RST) and are considered in terms of their component parasequences (5th order). Generally, TSTs are composed of skeletal grainstone–rudstone facies, HSTs are dominated by shaly nodular wacke-packstone facies, and RSTs are mostly calcarenite facies. Systems tracts, sequence boundaries and their correlative conformities, maximum flooding surfaces, and forced regression surfaces were traced from shallow shelf to basinal settings. This high-resolution framework also provides insight into the timing of tectonic fluctuations on this cratonic ramp during the Taconic Orogeny and documents the relative influence of tectonism on lateral facies distributions and eustatically derived cyclicity.     

Cambrian (Series 2 to Miaolingian) platform facies from central Sonora,Mexico and the regional correlation

This study provides new insights about depositional paleoenvironments through siliciclastic microfacies, carbonate microfacies, and biofacies analysis from sedimentary formations of the lower and middle Cambrian (Stage 4–Wuliuan), exposed in central Sonora, northern Mexico. Results of the petrographic analysis of 48 samples revealed the following lithologies: quartzarenite, oncolytic rudstone, grainstone-packstone, wackestone, mudstone, and to a lesser extent sandy limestone. Two siliciclastic microfacies were identified: (A) quartzarenite with cross-bedded and horizontal stratification deposited in an intertidal and supratidal environment; and (B) massive quartzarenite with Skolithos ichnofacies deposited in subtidal and intertidal environments. Four carbonate microfacies were identified: microfacies 1 is a sandy limestone with trilobite fragments; microfacies 2 is a grainstone with intraclasts, salterellids, hyolithids, trilobites, and echinoderms plates; microfacies 3 is an oncolytic rudstone consisting of microbes and abundant echinoderms plates; and microfacies 4 is a packstone-grainstone with abundant ooids, trilobite fragments, and echinoderm plates. Two biofacies were identified: Agnostid-polymeroid biofacies with predominance of the trilobites Pentagnostus, Bathyuriscus, Oryctocephalites, and Elrathina; and Pagetia biofacies with abundant trilobites of the genera Pentagnostus, Pagetia and Elrathina. It is concluded by the sedimentation model that changes in sea level is the most important parameter in determining the siliciclastic microfacies, carbonate microfacies and biofacies; as well as the depositional environments that vary from the coastline (subtidal to supratidal) to shallow-water open circulation marine platform with low and high energy waters. The Cambrian deposits of northern Mexico are correlated with the deposits of California and Nevada (USA), as well as to the Precordillera (Argentina), where the species in common show a strong affinity.     

Facies of a Lower Ordovician carbonate shelf (Mungok Formation: Taebaeksan Basin,Korea)

Summary The lithologic associations within the Lower Ordovician Mungok Formation in Korea define four depositional facies that formed across a continental margin fringing the Sino-Korean block: these facies represent lagoonal/restricted marine, shoal, inner shelf, and outer shelf environments. The stacking pattern of these facies reveals two systems tracts composed of five depositional sequences. The lower highstand systems tract consists of the lagoonal/restricted marine and shoal facies, whereas the upper lowstand systems tract comprises, in ascending order, inner shelf, outer shelf, and inner shelf facies. Three trilobite biofacies are recognized in the Mungok Formation: i.e.,Yosimuraspis, Kainella, andShumardia biofacies in ascending order. TheYosimuraspis Biofacies is dominated byYosimuraspis but also containsJujuyaspis andElkanaspis. The predominance of the endemic eponymous taxon suggests a lagoonal/restricted marine environment. The nearly monotaxicKainella Biofacies, which comprises pandemic genera such asKainella and occasionallyLeiostegium, may represent a less restricted environment than theYosimuraspis Biofacies. TheShumardia Biofacies occurs in the marlstone/shale lithofacies through relatively thick stratigraphic interval and is dominated by cosmopolitan trilobite taxa with some endemic species. The lithofacies and cosmopolitan trilobite assemblage of theShumardia Biofacies indicate that it occupied an outer shelf environment. The vertical succession of lithofacies and trilobite biofacies in the Mungok Formation records in general a shift from a restricted, shallow water environment to deeper-water environment.     

Trilobite cuticle thickness in relation to palaeoenvironment

100 thickness measurements from thin sections of cephala or pygidia of early Ordovician trilobites occurring across an onshore to offshore environmental gradient show that progressively greater maximum cuticle thickness was characteristic of increasingly inshore sites. There is a 40-fold difference between the thinnest and thickest cuticles, and exclusively thin cuticles are confined to the offshore Olenid Biofacies. Variability in cuticle thickness increases offshore to onshore. Environmental control is shown to be more influential on cuticle thickness than is the overall length of the trilobite: some comparatively large trilobites having thin cuticles and small trilobites thick cuticles. The environmental factors which might be responsible for the pattern are briefly discussed. The thin cuticles dominating the offshore Olenid Biofacies were probably appropriate for dysaerobic conditions. Thick cuticles in the most inshore biofacies may have offered protection against predators and turbulence, but the additional presence there of trilobites with thinner cuticles is considered to reflect the greater heterogeneity of the epeiric habitat.     

Taphonomic versus ecologic controls on taxonomic relative abundance patterns in tempestites

Evaluation of taphonomic sample bias in tempestites is an important prerequisite for paleoecologic analysis in storm-dominated shelf settings. Data from tempestites in the Upper Cambrian Bison Creek Formation of southern Alberta indicate that size, sorting greatly influences trilobite relative abundance patterns. Shifts in relative abundances of taxa both within and between beds are matched by changes in trilobite size. That is, collections fall into two intergrading types of assemblage which are dominated by taxa which reach large and small adult sizes, respectively. These shifts in dominance are interpreted as taphonomic variability within a single trilobite biofacies. Biofacies differentiation is expressed as abrupt changes in relative abundance patterns that are independent of trilobite size. Qualitative general models for size-sorting in tempestites are developed. Size-sorting in graded tempestites produces vertical shifts in relative abundances of taxa that mimic those expected in short-term autogenic ecologic succession. Size-sorting between beds in a single section could lead to an overestimation of the number of biofacies present. Size-sorting between proximal and distal portions of a depth gradient might generate abundance patterns that mimic those expected for 'depth-related communities', even in the complete absence of any underlying ecologic differentiation. The models represent taphonomic null hypotheses which must be rejected prior to making paleoecologic inferences in storm-dominated shelf environments. 0 Taphonomy, tempestites, Trilobita, Upper Cambrian, paleoecology.     

Facies,depositional sequences,and biostratigraphy of the Oligo-Miocene Asmari Formation in Marun oilfield,North Dezful Embayment,Zagros Basin,SW Iran

The Asmari Formation in Marun oilfield (south-west Iran), is about 440 m-thick marine carbonate succession with subordinate siliciclastic rocks, characterized by abundant benthic foraminifera (perforate and imperforate). Foraminiferal biostratigraphy indicates that this unit is Oligocene–Miocene in age. The distribution of benthic foraminifera and other components have led to the recognition of three siliciclastic and ten carbonate facies that were deposited in inner ramp (shoreline, tidal flat, restricted and open lagoon and shoal), middle and outer ramp sub-environments. Based on vertical facies trends, three third-order sequences in the Oligocene and three third-order sequences in the Miocene sediments have been identified. These depositional sequences are bounded by both type 1 and type 2 sequence boundaries. The transgressive systems tracts (TST) of sequences show deepening-upward facies trend with a gradual upward increase in perforate foraminifera, whereas the highstand systems tracts (HST) have a shallowing-upward facies trend and contain predominantly imperforate foraminifera. Deposition of these depositional sequences (DS) were controlled by both eustasy and tectonic subsidence.     

Coral assemblages, biofacies, and ecological zones in the mid-Holocene reef deDosits of the Enriquillo Valley, Dominican Republic

Stemann, T. A. & Johnson, K. G. 1992 07 15: Coral assemblages, biofacies. and ecological zones in the mid-Holocene reef deposits of the Enriquillo Valley, Dominican Republic. A large, subaerially exposed mid-Holocene reef in the Enriquillo Valley (southwest Dominican Republic) provides an excellent opportunity to examine the relationship between reefal ecology and reefal deposits. Coral species richness and diversity in the Enriquillo reef are comparable to that found in the recent of the Caribbean, and ecological zonation comprised of a shallow-water branching coral zone and a deeper water mixed-coral zone is apparent. Similar zonation and diversity patterns have been recognized on living Caribbcan reefs with moderate wave exposure. Three statistically discrete biopdcies can be discriminated in the Enriquillo deposits using quadrat point-counting techniques commonly used to census modern reefs. They include a facies dominated by Acropora cervicornis, a low diversity assemblage with abundant, large colonies of Siderastrea siderea and Stephanocoenia intersepta, and a higher diversity assembbdge composed of various taxa including Montastraea spp., Colpophyllia spp., and Agaricia spp. Each facies can be recognized at scales of 1–3 m², though in some cases they extend for more than 20 m². In general, the A. cervicornis facies is spatially segregated from the other two biofacies. although neither the shallow nor the deep-water ecological zone is comprised of a single reef biofacies. Rather, the biofacies described here appear to represent distinct micro-environments resulting from ecological variation at a subzonal scale. Micro-environments of similar scale are most likely preserved in other reef deposits. Recognition of these subzonal biofacies may have important consequences for the stratigraphical and paleoccological interpretation of fossil reefs. Corals, biofacies, reef zonation, coral communities, fossil reefs.     

Baltoscandian conodont biofacies fluctuations and their link to Middle Ordovician (Darriwilian) global cooling

The Middle Ordovician conodont genera that are suitable for palaeoenvironmental interpretations from the epicontinental Baltoscandian platform have been identified and evaluated to establish and describe conodont biofacies and their relationship to global cooling. The construction of biofacies was based on multivariate statistical analyses of more than 375 700 conodont specimens from 520 samples and 21 localities across Baltica. Three distinct, recurrent and laterally extensive conodont biofacies existed across the Baltoscandian platform of the Baltica continent during the Dapingian and early to middle Darriwilian stages (Middle Ordovician). A relatively shallow water conodont assemblage named the Baltoniodus–Microzarkodina Biofacies characterized the inner shelf localities in central Sweden, Estonia, Russia and Ukraine. In the distal shelf areas, patterns are more complex. Here, genera of the Periodon Biofacies characterized the shelf margin areas of the Scandinavian Caledonides facing the relatively warm Iapetus Ocean towards the north, whereas the Protopanderodus Biofacies dominated the distal shelf areas facing the cooler Tornquist Sea towards the south‐west. Although these three main biofacies continued to dominate during the succeeding Darriwilian stage, distinct changes in the distribution of biofacies took place during the transition from the Dapingian Stage to the Darriwilian. We argue that the biofacies change was triggered by a regressive event related to early Darriwilian cooling, and that the palaeoclimatological changes influenced the Baltic conodont faunas near the Tornquist Sea margin before those of the Iapetus margin (early vs middle Darriwilian).     

The upper Moscovian and basal Kasimovian (Pennsylvanian) of Central European Russia: Facies,subaerial exposures and depositional model

Summary The type upper Moscovian-basal Kasimovian argillaceous-carbonate succession of central European Russia contains regionally traced cyclothem-bounded subaerial exposure horizons (geosols) represented mainly by rendzina-type palaeosols. Palaeokarst profiles occurrarely and grade laterally to palaeosols. Composite subaerial profiles divided by one or two thin marine beds are called ‘multiple geosols’. The biofaces structure of the studied succession is defined by brachiopod and fusulinoid biofaces. The heterogeneous Choristites biofacies characterizes openmarine intervals, which constitute the bulk of the succession, and is defined by presence of Choristites. The Meekella biofacies with monospecific concentrations of Meekella shells and extreme rarity of other brachiopods characterises restricted peritidal intervals which commonly constitute the terminal regressive parts of major cyclothems. Three fusulinoid biofacies defined by Baranova and Kabanov (2003) include restricted peritidal Biofacies 1 with only small fusulinoids Fusiella and Schubertella present, open shoal-to-subtidal Biofacies 2 with the richest fusulinoid assemblages, and the most offshore Biofacies 3 with less diverse, sometimes Hemifusulina-dominated, fusulinoid assemblages. Bioturbation patterns and ichnofossils allow recognition of deeper subtidal Zoophycos and shallower non-Zoophycos ichnofacies. Among the latter, shallowest subtidal facies are characterized by presence of thalassinoid burrows. Intertidal laminated lithofacies with suppressed bioturbation contain Skolithos burrows. Seventeen lithofacies are recognized. Terrestrial lithofacies include topclays (upper clayey palaeosol horizons) and aeolian grainstones. Restricted peritidal lithofacies include cross-stratified skeletal-peloidal grainstones, fine-grained laminated grainstones-mudstones, and lagoonal mudstones. Open shoal lithofacies include ooidal grainstones (rare, only in Podolskian) and coarse skeletal-peloidal grainstones. The open subtidal lithofacies include skeletal packstones-rudstones, shallow subtidal packstones-wackestones, deeper subtidal packstones-wackestones, Ivanovia boundstones (only in Podolskian), proximal tempestites, distal tempestites, and skeletal wackestones-mudstones. The fossiliferous shale lithofacies is a miscellaneous group of marine shales lacking distinct features of the above-listed lithofacies. Conglomerates of cyclothem bases that are regarded as early transgressive lithofacies are variable in their palaeoenvironmental position and are characterized by concentrated pebbles derived from palaeosol reworking. The shallowest subtidal lithofacies of fine packstones-grainstones is considered as transitional between open subtidal and restricted peritidal lithofacies. The origin of stratiform dolostones is shown to be early diagenetic in the subsurface. The depositional model involves a shallow and broad epicontinental ramp, where through water circulation prevented stratification of the water column and allowed large skeletal benthos to colonize the entire spectrum of depositional environments. Storms are thought to be the principal water-mixing agent. The anti-estuarine circulation carrying oxygenated waters down-ward may explain the lack of anoxic features in the deepest facies that may have formed below storm wave base.     

Palaeoecological analysis of a sclerobiont fauna on a single basibiont across the Valanginian of the Neuquén Basin,west‐central Argentina

Sclerobiont communities have proven their environmental and taphonomic value in extant and past settings; studies are beginning to quantify and evaluate their changes across time. Through the Valanginian of the Neuquén Basin, Argentina, trigonioids of the genus Steinmanella present an almost continuous record. Materials were collected from shales and shell beds across second‐order systems tracts (LST, TST and HST). Their sclerobiont communities were characterized and compared across systems tracts and facies. In addition, a link between them and coeval oyster mass occurrences (OMOs) was investigated, since studies on other local sclerobiont communities have consistently shown pronounced oyster dominance. Eleven sclerobiont taxa were found on Steinmanella (bryozoans, bivalves, foraminifers and polychaetes), of which oysters are usually strongly dominant but for the TST, in which their dominance is diminished. The sclerobiont community has a simple structure and interspecific relationships and competition for space seem to have had little importance. Few differences in richness, taxonomy and other parameters were found between facies. Across systems tracts, features of the sclerobiont communities are differentiated mostly from swings in relative abundance of taxa. The lowered oyster dominance in the TST may reflect a source–sink dynamics between OMOs (sources) and Steinmanella as sinks in shallower, soft substrate settings. Likely the sea‐level rise temporally drowned the oyster carbonate factory, resulting in decreased larvae emigration to the sinks and lowered oyster dominance during the TST.     

A sequence stratigraphic approach to a Middle Triassic shelf-slope complex of the Ligurian Alps (Ligurian Briançonnais,Monte Carmo-Rialto unit,Italy)

The Mesozoic sedimentary cover belonging to the Monte Carmo-Rialto unit of the Ligurian Briançonnais domain is composed of Scythian clastics and Anisian to Carnian carbonate rocks over 300 m thick. This paper focuses on the stratigraphy of this carbonate complex, its environmental significance, and its evolution in light of dynamic stratigraphy. Our facies analysis of limestones and dolomites of the Triassic complex allowed us to reconstruct an environmental model. Data support a distally steepened carbonate ramp of Anisian age evolving to a more diversified Ladinian platform with an oolitic sand-bar belt separating the lagoon from the slope. The Monte Carmo-Rialto slope facies are the only witnesses of deep sedimentation in the Triassic terrains of the Ligurian Briançonnais domain, otherwise represented by shallow-water carbonate deposits. On the basis of facies succession, we have identified nine medium-scale cycles (third-order sequences) in the study area, comparable to those evidenced in the Briançonnais s.s. domain by the French authors. Small-scale cycles analysis evidenced mainly shallowing-upward trends in the examined sequences; although a few evidences of transgression-related deposits (deepening upward cycles) have been found at the base three sequences, they have been mostly obliterated by dolomitization and masked by local tectonics. For this reason, we can undoubtedly distinguish only the part of each sequence belonging to HST, while the TST, though present, still remains a partition that cannot be precisely characterized. In the same way, LSTs are not present in the Monte Carmo-Rialto unit, due to the original relative landward position of the examined area. Sequence stratigraphy analysis indicates different long-term dynamics for the two evolutionary stages of the Triassic Ligurian platform: a general landward backstepping to moderate progradation during the Early Anisian and true progradation during the latest Anisian and Ladinian. In addition, a good fit with the sequences proposed by the SEPM chart has been found, indicating a correspondence for the third-order sequences of the Middle Triassic.     

An Upper Miocene siliciclastic-carbonate ramp: depositional architecture, facies distribution, and diagenetic history (Capo Vaticano area, southern Italy)

During the Late Miocene, the marginal areas of the Mediterranean Basin were characterized by the development of mixed siliciclastic-carbonate ramps. This paper deals with a temperate siliciclastic-carbonate ramp (late Tortonian–early Messinian in age) which crops out in the Capo Vaticano area, Southern Apennines (Italy). Carbonate components are mainly represented by calcitic skeletal fragments of coralline red algae, bryozoans, bivalves, and larger foraminifera, whereas corals, brachiopods, echinoderms, and planktonic foraminifera are subordinate. In the studied ramp, the depositional geometries of the main unit, the ‘Sabbie gialle ad Heterostegina’, show a gradual steepening from low/middle (dip about 2–5°) to steep slope settings (up to 25°). The microfacies observations, the quantitative analyses of the main biogenic components as well as the rhodolith shapes and growth forms allowed the differentiation between the middle and the outer ramp depositional setting and the refining of the stratigraphic framework. The middle ramp is characterized by coralline red algal debris packstone facies often associated with larger foraminiferal floatstone/packstone facies, while the outer ramp is characterized by rhodolith floatstone/rudstone facies. These facies pass basinward into typical open-marine deposits (planktonic foraminiferal facies). The taxonomic composition of the coralline red algal assemblage points to a temperate paleoclimate and emphasizes the Miocene Mediterranean phytogeographic patterns. The absence of non-skeletal grains (ooids and green algae), the paucity of Porites patch reefs, the rare occurrence of primary marine cementation, all confirm that the studied ramp was poorly lithified within a warm–temperate setting. The flat depositional profile of the ramp can be related to the absence or paucity of primary marine carbonate cements.     

Facies analysis and sequence stratigraphy of an Upper Jurassic carbonate ramp in the Eastern Alborz range and Binalud Mountains, NE Iran

Upper Jurassic (Oxfordian-Kimmeridgian-Tithonian?) strata of NE Iran (Lar Formation) are composed of medium- to thick-bedded, mostly grainy limestones with various skeletal (bivalves, foraminifera, algae, corals, echinoderms, brachiopods, and radiolaria) and nonskeletal (peloids, ooids, intraclasts, and oncoids) components. Facies analysis documents low- to high-energy environments, including tidal-flat, lagoonal, barrier, and open-marine facies. Because of the wide lateral distribution of facies and the apparent absence of distinct paleobathymetric changes, the depositional system likely represents a westward-deepening homoclinal ramp. Four third-order depositional sequences can be distinguished in each of five stratigraphic measured sections. Transgressive system tracts (TST) show deepening-upward trends, in which shallow-water (tidal flat and lagoonal) facies are overlain by deeper-water (barrier and open-marine) facies. Highstand systems tracts (HST) show shallowing-upward trends in which deep-water facies are overlain by shallow-water facies. All sequence boundaries in the study area (except at the top of the stratigraphic column) are of the nonerosional (SB2) type. Correlation of depositional sequences in the studied sections show that relatively shallow marine (tidal-flat, lagoonal, barrier, and shallow open-marine) conditions dominated in the area. These alternated with deep-water open-marine wackestone and mudstones representing zones of maximum flooding (MFZ).     

Applications of nummulitids and other larger benthic foraminifera in depositional environment and sequence stratigraphy: an example from the Eocene deposits in Zagros Basin,SW Iran

The Tale-Zang Formation in Zagros Mountains (south-west Iran) is a Lower to Middle Eocene carbonate sequence. Carbonate sequences of the Tale-Zang Formation consist mainly of large benthic foraminifera (e.g. Nummulites and Alveolina), along with other skeletal and non-skeletal components. Water depth during deposition of the formation was determined based on the variation and types of benthic foraminifera, and other components in different facies. Microfacies analysis led to the recognition of ten microfacies that are related to four facies belts such as tidal flat, lagoon, shoal and open marine. An absence of turbidite deposits, reefal facies, gradual facies changes and widespread tidal flat deposits indicate that the Tale-Zang Formation was deposited in a carbonate ramp environment. Due to the great diversity and abundance of larger benthic foraminifera, this carbonate ramp is referred to as a “foraminifera-dominated carbonate ramp system”. Based on the field observations, microfacies analysis and sequence stratigraphic studies, three third-order sequences in the Langar type section and one third-order sequence in the Kialo section were identified. These depositional sequences have been separated by both type-1 and type-2 sequence boundaries. The transgressive systems tracts of sequences show a gradual upward increase in perforate foraminifera, whereas the highstand systems tracts of sequences contain predominantly imperforate foraminifera.     

Anomalous facies and ancient faeces in the latest middle Cambrian of Sweden

The middle Cambrian–Furongian transitional interval was a time of significant biotic and environmental changes. Strata of this age in Scania, southern Sweden, contain two interlayered biofacies, a normal one dominated by trilobites and an anomalous one dominated by phosphatocopines (small bivalved arthropods). In places these biofacies are separated by intervals barren of fossils. In a phosphatocopine facies without trilobites in the upper Agnostus pisiformis Zone at Andrarum we recovered scattered fossil aggregates with a homogeneous composition of tightly packed and stacked phosphatocopines. These aggregates are interpreted as coprolites produced by an undetermined predator, possibly the chaetognath-like protoconodont animal or some other soft-bodied metazoan. The so-called barren intervals of Scania are not necessarily barren of fossils, but only trilobites, brachiopods, and other skeletal elements with a calcium-carbonate composition. The phosphatocopine facies with coprolites in the uppermost part of the A. pisiformis Zone correlates with the trilobite mass extinction at the top of the Marjumiid Biomere in Laurentia and immediately prior to the onset of the Steptoean Positive Carbon Isotope Excursion (SPICE), inferring a global shift in the oceanic chemistry that in Scania favoured phosphatocopines over the more common, trilobite-dominated faunas.     

Sequence stratigraphic analysis and sea-level history of the Upper Jurassic deposits (Mozduran Formation), south of Aghdarband,NE Iran

Abstract

The Kopet-Dagh Basin is a large sedimentary basin in northeastern Iran that host the giant Khangiran and Gonbadli gas fields. The Mozduran Formation with its various sedimentary facies is an important reservoir widely distributed in the basin. A sedimentological analysis of Upper Jurassic Mozduran Formation resulted in an accurate reconstruction of the sedimentary environments and the sequence stratigraphic framework south of Aghdarband. The strata consist of six different facies associations including 12 carbonate, one evaporate and two siliciclastic subfacies. On the basis of their various components, structural and textural characteristics, these facies were deposited on a homoclinal ramp in tidal flat to open marine environments ranging from supratidal to subtidal settings. Facies A1 and A2 represent open marine, B1-B4 Shoal, C1-C4 lagoonal and D1, D2, E, T1 and T2 tidal flat and Salina environments. In addition, based on detailed field and laboratory studies on the facies architecture, several large-scale (long-term) depositional sequences could be distinguished in the stratigraphic sections of the study area. These sequences are composed of LST, TST and HST that are separated by a SB1 and SB2 sequence boundaries. The paleogeography of the study area during the Late Jurassic time is reconstructed in five block diagrams.     

Taphonomic pathways and comparative biofacies and taphofacies in a Recent intertidal/shallow shelf environment

The development of taphonomic approaches to facies analysis requires a foundation in facies-based actualistic studies. Modern intertidal and shallow shelf environments at Provincetown Harbor. northern Cape Cod, Massachusetts (USA) provide an opportunity to compare pattcrns and controlling factors in molluscan biofacies and taphofacies distributions. Variation in faunal composition, ecologic variables, and taphonomic attributes of molluscan death assemblages produce distinct patterns of environmental zonation: (1) Faunal composition (biofacies) primarily tracks variation in substrate type among environments (sand, rock, peat, and Zostera marina beds). (2) Ecologic variables (equitability, infauna: epifauna ratio, gastropod:bivalve ratio, and predation on M. mercenaria) appear to reflect tidal exposure time. (3) Taphononic attributes (fragmentation, abrasionm, corrosion, bioerosion, and encrustation) of the common bivalve M. mercenaria track environmental energy, in particular its effects on the stability and reworking of hardparts at the sediment surface. Shells in different environments proceed along different taphonomic pathways - the order of acquisition of taphonomic features by hardparts. An encrustation/bioerosion-dominated pathway characterizes low energy environments; the upper intertidal and deeper subtidal. An abrasion-dominated pathway characterizes the high energy lower intertidal and shallow subtidal. Contrasting pathways produce distinct proportions of taphonomic attributes in time-averaged samples; proportions that delineate taphofacies. Integrated taxonomic, ecologic and taphonomic data provide a more complete picture of environmental processes than any approach alone. Taphonomic data not only furnish information not readily provided by other approaches, but free paleoecology from the constraints of taxonomic uniformitarianism. □Taphonomy, comparative taphonomy, taphofacies, biofacies, cluster analysis, multidimensional scaling, taphonomic pathways, Recent, actualism, intertidal, molluses.     

Filtration models, guilds, and biofacies: Crinoid paleoecology of the Stanton Formation (Upper Pennsylvanian), midcontinent, North America

Filtration models make several predictions concerning the distribution of crinoids among benthic habitats: (1) generally, higher energy shoreward settings should contain crinoid assemblages dominated by taxa with dense mesh filtration fans; (2) generally, lower energy offshore settings should be dominated by crinoids with open mesh filtration fans; (3) diversity should be highest in nearshore settings, which generally contain low energy microhabitats within high energy settings where both dense fan and open fan forms can co-occur, whereas offshore assemblages should be dominated by open fan taxa; thus, open fan forms should be more eurytopic than dense fan forms. Also, previous empirical models for crinoid distribution patterns note that nearshore assemblages are dominated by large forms whereas offshore assemblages are dominated by small forms. Filtration fan morphology and body size interact significantly to determine how and where crinoids feed and thus are used here as criteria for distinguishing late Paleozoic crinoid guilds. The distribution patterns of crinoid taxa and guilds among the benthic paleoenvironments of the Upper Pennsylvanian (Missourian) Stanton Formation were used to test the predictions of the filtration models.

The results of quantitative analyses show that large, dense fan crinoids are significantly more abundant in shoreward assemblages than smaller, open fan crinoids, which are more abundant in offshore assemblages; these results are consistent with the aforementioned models. However, diversity patterns are not consistent with the filtration models, indicating that mid-depth and aerobic offshore assemblages are more taxonomically and trophically diverse than nearshore and dysaerobic offshore assemblages, ostensibly due to the overlapping ranges of characteristically nearshore and offshore taxa. Analysis of guild distribution patterns also indicates some contradictions to predicted patterns of stenotopy and eurytopy, indicating additional complexities to late Paleozoic crinoid paleoecology.

Five recurrent crinoid assemblages, or biofacies, have been identified from the Stanton Formation of midcontinent North America. The guild structure and diversity of these biofacies are characterized here as: (1) a moderate diversity nearshore biofacies dominated by large calyx, dense fan guilds; (2) a high diversity midshelf biofacies characterized by equitable contributions of many taxa and guilds; (3) a moderate diversity offshore biofacies dominated by non-pinnulate and small calyx, open fan pinnulate guilds; (4) a low diversity offshore dysaerobic biofacies dominated by small calyx pinnulate guilds; (5) a depauperate offshore biofacies dominated by a unique stalkless species, Paragassizocrinus tarri. Except for Biofacies 5, the number of guilds appears to increase offshore; however, the equitability of taxa among and within the guilds generally decreases offshore. This pattern indicates that the heterogeneity of resources, or the recognition by taxa of different classes of resources, as reflected by the variety of functional types, increased from nearshore to offshore, but that the abundance or accessibility of these resources, and/or the relaxation of competition within guilds, increased from offshore to nearshore.     

Middle Ordovician cephalopod biofacies and palaeoenvironments of Baltoscandia

During the Middle Ordovician cephalopods became an important part of the macrofauna of the Baltoscandian carbonate platform. The earliest cephalopod abundance peak was reached during the early Darriwilian, within the Kunda Stage Yangtzeplacognathus crassus and Lenodus pseudoplanus conodont zones. In sediments of this time interval large orthoconic cephalopods often occur in masses with more than one specimen per square‐meter on bedding surfaces. The assemblages are characterized by the strong dominance of often large endocerids. In proximal depositional settings coiled tarphycerids and other cephalopod groups are an important additional component. In the most distal settings orthocerids are the most important secondary component. Correspondence Analysis of assemblages throughout Baltoscandia revealed three distinct biofacies, which here are termed Orthocerid, Proterovaginoceras and Anthoceras Biofacies, respectively. The biofacies reflect differences in depth and proximity to the shoreline and are consistent with the Baltoscandian Confacies Belts. Spatial changes in absolute abundance and taxonomic composition indicate increased original cephalopod population densities and habitat expansion within the Y. crassus and L. pseudoplanus conodont zones. A nearly coeval abundance peak in a similar facies in South China indicates supraregional causes of the mass occurrence, probably reflecting a globally increased nutrient availability in the water column during the Darriwilian.     

The western edge of the Mediterranean Pelagian Platform: A Messinian mixed siliciclastic–carbonate ramp in northern Tunisia

The marine Messinian deposits of Tunisia cover a narrow littoral strip some 300 km long between the northern Bizerte and Cap Bon areas and the central–eastern Sahel region. Litho- and biofacies analysis of six stratigraphic sections reveals the distinctive features of these deposits.The lower Messinian deposits are characterized by ubiquitous siliciclastics and abundant oolitic/bioclastic limestones organized in an eastward facing ramp. Westward (landward), this ramp changes into coastal lagoons, sometimes containing evaporites. Eastward, the ramp passes to the reefal Pelagian Platform extending as far as Lampedusa.Two main sedimentary cycles are distinguished: 1) an early Messinian siliciclastic retrogradational then oolitic/bioclastic progradational cycle (Beni Khiar Formation and lower Oued bel Khedim Formation); 2) a late Messinian brackish to continental cycle that probably accumulated in rapidly subsiding lagoons (Oued el Bir Formation and upper Oued bel Khedim Formation). The Tunisian early Messinian cycle is partly eustatically controlled, but the late Messinian cycle cannot be confidently correlated to other well-known Messinian series because of tectonic movements.The lower Messinian deposits of Tunisia are also characterized by abundant suspension-feeding organisms (molluscs and bryozoans) and rare corals, calcareous algae, echinoids, and larger benthic foraminifers. The proposed palaeoenvironmental model shows that the lower Messinian ramp of Tunisia was located on a current-protected margin and subjected to continent-derived sediment and nutrient supply. Eastward, nutrient influx diminished and a shallow-water isolated carbonate platform with coralgal facies developed between the western and the eastern Mediterranean basins. The main hydrological connection between these two basins occurred through a narrow seaway situated to the northeast of the Pelagian Platform, south of Sicily and Malta.