Revision of Late Permian tetrapod tracks from the Dolomites (Trentino-Alto Adige,Italy)

The Val Gardena Formation of the Dolomites region in northern Italy preserves the most significant assemblage of Late Permian tetrapod footprints in the world. More than 120 years of collecting resulted in about 900 publicly accessible specimens from the study area. This huge amount of data is comprehensively revised in the light of recent advances in the study of Late Palaeozoic – Early Mesozoic tetrapod ichnofossils. According to our analyses, the Val Gardena Sandstone Formation includes tracks that can be assigned to cf. Batrachichnus isp. (temnospondyl amphibian), Capitosauroides isp. (amphibian), Dicynodontipus isp. (cynodont therapsid), Dolomitipes accordii n. igen. n. comb. (dicynodont therapsid), cf. Dromopus isp. (neodiapsid), Pachypes dolomiticus (pareiasaurian parareptile), Paradoxichnium problematicum (archosauromorph neodiapsid), Procolophonichnium tirolensis n. comb. (procolophonoid parareptile), cf. Protochirotherium isp. (archosauriform neodiapsid) and Rhynchosauroides pallinii (neodiapsid). The ichnoassociation is dominated by tracks of neodiapsid and parareptilian tetrapods, whereas synapsid and anamniote tracks are rather minor components. It includes 10 out of 12 tetrapod ichnogenera known from Lopingian deposits and thus it constitutes a reference for the Paradoxichnium biochron. It shows striking similarities with other low-latitude non-aeolian contemporaneous ichnoassociations of Europe and North Africa, differences may be linked to the palaeoenvironment. Moreover, it shows a clear Triassic affinity.

The new ichnogenus Dolomitipes was registered in Zoobank.org. urn:lsid:zoobank.org:act:5B4D871C-D16A-4E93-8211-CEE08019BA60     

The first Mexican record of peccary footprints (Artiodactyla,Tayassuidae) from the late Cenozoic of Puebla: ichnotaxonomy and palaeobiological considerations

Abstract

A set of artiodactyl footprints preserved in the Pie de Vaca Formation from the Plio-Pleistocene of south-central Puebla in central Mexico is formally described. The sample consists of five footprints that represent a trackway produced by one individual. A comparative study and a taxonomic assessment were performed in order to identify the track-maker. The ichnites are characterized by having a rounded shape, length of about 5 cm, two hoof imprints that are longer than wide, and tips of the hooves directed forwards. Their size and morphology are comparable to those produced by extant peccaries. The spatial distribution of the footprints has been related to an individual in fast walking or even in a slow trot. Ichnotaxonomically, the record represents a new ichnospecies formally referred as to Tayassuichnum felixarangutii. A member of Platygonus, the most common peccary for the late Cenozoic of North America is proposed as the potential track-maker. The record contributes to the (indirect) presence of peccaries in central Mexico during the late Cenozoic, representing the first documented ichnites referable to that group of artiodactyls in the Mexican territory and the second mention in the North American subcontinent.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:41D44DD9-49F0-475D-B210-84DE5E9CBFAA     

Eolacertidae: a new extinct clade of lizards from the Palaeogene; with comments on the origin of the dominant European reptile group – Lacertidae

We describe a new lizard taxon, Stefanikia siderea gen. et sp. nov., from the early-middle Eocene locality of Messel in Germany based on a nearly complete skeleton, which we studied using μCT methods. It shares many characters with the Eocene taxon Eolacerta, which is broadly distributed in the Eocene of central and Western Europe, but is much smaller and shows several important anatomical differences. The new discovery sheds light on the paleodiversity of these lizards in the Eocene of Europe, and the new family name Eolacertidae is proposed to encompass Eolacerta and Stefanikia. The relationships of Eolacerta have been intractable. Our phylogenetic analyses confirm that Eolacertidae is a member of the clade Lacertiformes and provide strong support for a sister-group relationship to Lacertidae. In some places, skin impressions are preserved, displaying the body scalation. As such, the exquisitely preserved specimens of Eolacertidae from Messel provide new insight into the morphology and ecology of lizards on the stem of Lacertidae, Europe’s dominant group of living reptiles.

http://zoobank.org/urn:lsid:zoobank.org:pub:DFD01563-A913-4286-B64B-E0912474FD08     

Eocene ctenodactyloid rodent assemblages and diversification from Erden Obo,Nei Mongol,China

New ctenodactyloid occurrences (Yuomys magnus sp. nov., Gobiomys neimongolensis and Gobiomys exiguus) from the ‘Lower Red’ to the ‘Upper Red’ beds of the Erden Obo section in Nei Mongol, China are reported. Here, we present an updated review of the ctenodactyloid rodents from the Erden Obo section. We have recognised eight species and one indeterminate species belonging to six genera: Advenimus, Chenomys, Gobiomys, Tamquammys, Simplicimys, and Yuomys. The ctenodactyloids show an initial burst in diversification in the upper part of the ‘Basal Red’ of the early Eocene. A maximum richness peak is present at the lower part of the ‘Basal White’ which falls near the boundary of early and middle Eocene. Gobiomys, the only surviving genus from the ‘Middle Red’ to the ‘Upper Red’ beds, shows that the diversity of ctenodactyloids declined rapidly after late middle Eocene. Based on the study of the abundant ctenodactyloid rodents, we recognise some evolutionary tendencies among Eocene ctenodactyloids.

http://www.zoobank.org/urn:lsid:zoobank.org:act:33D2442E-DA27-494A-80CB-2DB9DA9E4E1A     

Unrecognized species diversity and new insights into colour pattern polymorphism within the widespread Malagasy snake Mimophis (Serpentes: Lamprophiidae)

Although many wide-ranging taxa occur in Madagascar, phylogeographic studies for most of these species are still lacking. This is especially the case for snakes, where of more than 100 endemic species, the population structure of only two species has so far been examined. Here, we examine genetic population structure of one of the most common snakes of Madagascar, Mimophis mahfalensis (Grandidier, 1867). This taxon is the only representative of Psammophiinae in Madagascar, where the majority of species in this subfamily is distributed throughout mainland Africa. Applying an integrative approach, where both morphological data and genetic results from coalescent species delimitation analyses are considered, we determine that Mimophis mahfalensis is composed of two distinct taxa: M. mahfalensis in the central montane and southern parts of Madagascar, and a second new species restricted to the north and north-west, which we describe here. We also examine the colour pattern polymorphism exhibited in Mimophis, which has been previously hypothesized as sexually dimorphic and/or geographically correlated. However, we find all three colour morphs in both sexes, and both species of Mimophis. Our work highlights the importance of phylogeographic studies that examine wide-ranging taxa to detect cryptic species diversity, even amongst species that are common, or have been previously considered to be well known.

www.zoobank.org/lsid:zoobank.org:pub:9791DC0B-49E5-4571-884C-4AA85EAF2472     

Re-evaluation of Chelichnus tazelwürmi,a non mammalian therapsid-grade track from the Upper Permian Arenaria di Val Gardena

In this paper, a revision of tracks referred to as Chelichnus tazelwürmi is reported. The performed analysis, consisting of a holistic approach by means of a mainly morphological analysis, and a secondarily functional one, led to the proposal of a new ichnogenus, named as Contiichnus tazelwurmi. The three dimensional morphology of the tracks allows for the inference of a complex cycle of locomotion by the trackmakers. The tracks were formed in the main phases (i.e. touch-down, weight-bearing and kick-off) by different axes of body load and transference, indicating that the whole fore autopod was involved in the cycle of locomotion and actively contacted the substrate, while for the hind autopod the functional prevalence was markedly centro-medial. Some track features suggest a therapsid-grade synapsid as potential trackmaker. However, the reconstructed autopodial structure does not correlate with known autopods from the Late Permian body fossil record. These observations stress the importance of tetrapod ichnology studies in improving knowledge in the field of vertebrate palaeontology. http://www.zoobank.org/urn:lsid:zoobank.org:pub:B4EB4D42-1A3B-48EC-B83F-6942F741AF30

http://www.zoobank.org/urn:lsid:zoobank.org:pub:B4EB4D42-1A3B-48EC-B83F-6942F741AF30     

New species and new records of Apataniidae (Insecta: Trichoptera) from east central China

Three new species of Apataniidae from China, Apatania protracta sp. n., Apatidelia paramartynovi sp. n., and Moropsyche dawuensis sp. n. are described and illustrated. We also report Apatania semicircularis Leng and Yang, 1998 for the first time from Hubei Province.

Apatania protracta: http://zoobank.org/urn:zoobank.org:pub:8317EB90-9316-4F7B-94C6-65938A6F2873

Apatidelia paramartynovi: http://zoobank.org/urn:zoobank.org:pub:8741CE6F-B339-4842-BD8C-04BE8B21A222

Moropsyche dawuensis: http://zoobank.org/urn:zoobank.org:pub:B5CB815D-59B5-401F-9308-9EE24A9BBAF7     

New information and establishment of a new genus for the Egyptian Paleogene turtle ‘Stereogenys’ libyca (Podocnemididae,Erymnochelyinae)

The poorly known pleurodiran turtle ‘Stereogenys’ libyca (Podocnemididae, Erymnochelyinae) is studied in this paper. It was defined in lower Oligocene levels of the Fayum Depression (Egypt), its presence in the upper Eocene levels of the same area being confirmed here. Its holotype corresponds to an almost complete and well preserved shell, that was until recently considered to be lost. No photograph of material attributable to this species had hitherto been published. Thus, the current knowledge about it was exclusively based on the limited information published more than a century ago. The recently rediscovered holotype, and also more than fifteen additional historical but unpublished specimens recognized here as attributable to this species, are studied in this paper. This species is attributed to a new genus, Andrewsemys, and the new combination Andrewsemys libyca is proposed.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:6D87831A-4510-4A3E-A624-C40FD644F448     

Bivalve borings in Lower Jurassic Lithiotis fauna from northeastern Italy and its palaeoecological interpretation

Random shell sections of the Pliensbachian (Early Jurassic) larger bivalve Opisoma from columns within the Main Post Office building of Ferrara, northern Italy, have been discovered to bear neat clavate-shape boreholes. These boreholes belong to the ichnogenus Gastrochaenolites Leymerie and represent bivalve borings. Opisoma is a subordinate component of the Lithiotis fauna characterised by aberrant shells thriving in tropical lagoonal settings which were widespread throughout the Tethyan and Panthalassa coasts. Although the Lithiotis fauna is well known in the palaeontological literature, no bivalve boring have been so far been formally described. The uniqueness of the morphology, size and substrate of these borings merits the designation of the new species Gastrochaenolites messisbugi ichnosp. nov. which thereby represents the first ichnospecies described from this fauna. The morphology of the boreholes and the included bivalves allows the boring activity to be ascribed to a mytilid bivalve. Palaeoecological and taphonomic analyses allowed the presence of the boreholes to be correlated to the Opisoma mode of life (epifaunal, free-living form) as well as to generally low sedimentation rates and seasonal mesotrophic conditions during an overall oligotrophic regime.

http://zoobank.org/urn:lsid:zoobank.org:act:CCF77B32-D459-4305-BC86-93F228852E50     

Succinapion telnovi n. gen. et n. sp. of the tribe Kalcapiini (Coleoptera: Brentidae: Apioninae) in Baltic amber

A new genus and species, Succinapion telnovi n. gen. et n. sp. (Coleoptera: Curculionoidea: Brentidae: Apioninae: Kalcapiini) is described and illustrated from Upper Eocene Baltic amber. The new genus is similar to the genus Melanapion Wagner, 1930 but differs from it in having femora ventrally with spine at distal 1/3, simple claws, a longer rostrum, elytra weakly widened towards apex, longer antennae and slightly narrower elytral striae.http://www.zoobank.org/urn:lsid:zoobank.org:pub:2A95C49D-5589-4ACA-8A87-0DDF635BA25E     

Early Miocene rodents of Gökler (Kazan Basin,Central Anatolia,Turkey)

Abstract

The rich and relatively diverse fossil mammalian assemblage from Gökler is of special importance for understanding of faunal evolution in Central Anatolia. Large mammals were not recovered, but insectivores and rodents are abundant. The assemblage of rodents is studied in detail and comprises mainly diversified cricetids. Dormice are abundant, but are represented by only one species. Squirrels are represented only by few specimens and also beaver remains were identified. Spanocricetodon sinuosus is referred to a new genus Latocricetodon nov. gen that is tentatively assigned to the Pseudocricetodontinae. Newly named species are Cricetodon goklerensis sp. nov., Democricetodon haltmari sp. nov., Eumyarion lukasi sp. nov. and Glirudinus matusi sp. nov. The rodent assemblage is assigned to local zone C which is correlated to the European biounit MN2 (early Miocene). Our biochronological assessment is supported by radiometric dating from two volcanic ash layers.

Latocricetodon

LSID http://zoobank.org/urn:lsid:zoobank.org:act:3414DB1E-0C5E-4154-BE5E-02A9ED183B1A

Cricetodon goklerensis

LSID http://zoobank.org/urn:lsid:zoobank.org:act:1B658872-6C10-4355-B87C-3E6277AF4EDA

Democricetodon haltmari

LSID http://zoobank.org/urn:lsid:zoobank.org:act:9B13F956-7F8C-406A-9970-1F5E999E54C6

Eumyarion lukasi

LSID http://zoobank.org/urn:lsid:zoobank.org:act:34DED87E-855F-4969-AB84-10BED5C572BF

Glirudinus matusi

LSID http://zoobank.org/urn:lsid:zoobank.org:act:798ECB9A-E3B5-4C38-B5B3-FEB17AF734FE     

Contribution to the knowledge of the click beetle genus Zorochros (Coleoptera: Elateridae) from Saudi Arabia

Examination of specimens of Elateridae revealed new generic and species records for Saudi Arabia. Zorochros amalec (Peyerimhoff) is recorded from Al Majardah, and Zorochros yosrae sp. n. is described as a new species from Thalouth Almanazar.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:E4B9EB99-BE55-4884-B220-BF9928926312     

A new species of Hercostomus Loew, 1857 (Diptera: Dolichopodidae) from Turkey

A new species of Hercostomus is described from Turkey: Hercostomus anatoliensis sp. n. The new species is close to Hercostomus parvilamellatus (Macquart, 1827), but is distinguished by its genitalia.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:38B210E3-610D-4BAB-91AC-A212A8054474     

A new sawshark,Pristiophorus laevis,from the Eocene of Antarctica with comments on Pristiophorus lanceolatus

The highly fossiliferous Eocene deposits of the Antarctic Peninsula are among the most productive sites for fossil remains in the Southern Hemisphere and offer rare insights into high-latitude faunas during the Palaeogene. Chondrichthyans, which are represented by abundant isolated remains, seemingly dominate the marine assemblages. Eocene Antarctic sawsharks have only been known from few isolated rostral spines up to now, that were assigned to Pristiophorus lanceolatus. Here, we present the first oral teeth of a sawshark from the Eocene of Seymour Island and a re-evaluation of previously described Pristiophorus remains from Gondwana consisting exclusively of rostral spines. The holotype of Pristiophorus lanceolatus represents a single, abraded and insufficiently illustrated spine from the Oligocene of New Zealand. All other Cenozoic rostral spines assigned to this species are morphologically very indistinct and closely resemble those of living taxa. Consequently, we regard this species as dubious and introduce a new species, Pristiophorus laevis, based on oral teeth. The combination of dental characteristics of the new species makes it unique compared to all other described species based on oral teeth. Rostral spines from the Eocene of Seymour Island are assigned to this new species whereas those from other Cenozoic Gondwana localities remain ambiguous.

LSID urn:lsid:zoobank.org:pub:7177A373-527B-4315-85F6-25180DB5E087     

A new species of Maladera from Jordan (Coleoptera: Scarabaeidae: Sericini)

A new Maladera species is described from Jordan: M. schnitteri sp. n. A key to the species of the subgenus Eusericula is given, and the habitus and genitalia are illustrated.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:D41D75B5-D700-4C2B-9058-04D14006C810     

Description of a new pseudoscorpion species of the genus Amblyolpium Simon (Pseudoscorpiones: Garypinidae) from north-western Iran

A new species of pseudoscorpion, Amblyolpium atropatesi n. sp. is described and illustrated on the basis of the adult specimens extracted from soil by Berlese funnel.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:0150B850-E80D-46F5-983C-7E958FE69B87     

Three new species of Zerconidae (Acari: Mesostigmata) from Turkey

Three new species, Zercon afyonensis sp. n., Zercon karacamehmeti sp. n. and Zercon soguticus sp. n., are described and illustrated based on samples collected in the Inner Aegean Region. The similarities and differences between the related species are discussed.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:3FA1701A-A243-4E45-8AE9-FCB733CC56A6     

A new species of ant-loving cricket Myrmecophilus Berthold, 1827 from Cyprus (Orthoptera: Myrmecophilidae)

A new species of ant-loving cricket, Myrmecophilus cyprius sp. n., collected in the Republic of Cyprus from the nests of Messor structor (Latreille, 1798) (Hymenoptera: Formicidae), is described and illustrated. The species belongs to the subgenus Myrmophilina Silvestri, 1912.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:B588546E-FC72-4996-A0C2-583BCE46905E