Late Ordovician–Early Silurian facies development and environmental changes in the Subpolar Urals

The continuous Upper Ashgill–Sheinwoodian carbonate succession in the most eastern Kozhym River area in the Subpolar Urals comprises the Yaptikshor (Rawtheyan), Kamennaya baba (Hirnantian), Ruchej and Manyuku (Llandovery–?Sheinwoodian) formations. The facies of the deep subtidal Yaptikshor Fm. mark an abrupt sea‐level rise following emergence of the Bad’ya reef (Rawtheyan). Carbonate breccias at the base of the Kamennaya baba Fm. correlate with the beginning of the Hirnantian glaciation and change upwards towards the Ordovician–Silurian boundary with the development of light‐grey massive boundstone/packstone shoal deposits. An abrupt change in facies to the Rhuddanian–Aeronian Ruchej Fm. continental slope environment marked the start of a long‐term sea‐level rise. The uppermost Aeronian–?Sheinwoodian is represented by submarine canyon carbonate conglobreccias of the Manyuku Fm. unconformably underlying the Balban’yu reef. The rapid facies changes at the base of the Hirnantian and at the Ordovician–Silurian boundary were of global eustatic origin. In contrast, the abrupt changes in the Rawtheyan and the formation of the Manyuku Fm. conglobreccias were of local or regional origin associated with tectonics. They were followed by the start of a regional transgression (Yaptikshor Fm.) and a global transgression marked by the initiation of the Balban’yu Reef in the Sheinwoodian.     

New bryozoan species from the stratotype section of the Upper Kazanian Substage

Representatives of nine bryozoan genera of four orders are identified for the first time from the carbonate deposits of the stratotype section of the Upper Kazanian Substage of the Middle Permian. Among them four new species are established and described: Permopora volgensis sp. nov., Stellahexaformis burovi sp. nov., Alternifenestella esaulovae sp. nov., and Wjatkella definita sp. nov. A brief lithological characterization is provided of the carbonate deposits that contain the remains of these bryozoans and were formed under the conditions of transgression-regression cycles of the Late Kazanian marine basin. Some questions concerning the taphonomy, ecology, and morphology of the bryozoans studied are considered.     

新疆柯坪和甘肃平凉上奥陶统底部附近的牙形刺

新疆柯坪大湾沟和甘肃平凉官司庄剖面是Nemagraptusb gracilis带及上奥陶统底界的重要剖面,研究这两剖面的牙形刺具有十分重要的意义。本文描述了采自新疆柯坪大湾沟和甘肃平凉官司庄剖面上奥陶统底界附近的牙形刺共13属18种。     

New conodont species from the Kasimovian Stage (Upper Carboniferous) of Moscow and Moscow Basin

Three new conodont species, Streptognathodus neverovensis, S. isakovae, and Idiognathodus mestsherensis, which are widespread in the Kasimovian Stage of central Russia, are described.     

Preliminary palynological investigation of Saudi Arabian Upper Ordovician glacial sediments

An Ordovician stratigraphically admixed palynomorph assemblage that contains palynomorphs eroded from Middle through Upper Ordovician strata characterizes the Hawban Member (restricted) of the Sarah Formation in central Saudi Arabia. This distinctive assemblage, combined with detailed sedimentology, helps identify the presence of Hirnantian Gondwanan glacial sediments on the Arabian Plate. Similar Ordovician admixed assemblages have been recognized from Upper Ordovician glacial sediments elsewhere along the Gondwanan margin. Within Saudi Arabia the composition of reworked assemblages depends upon the stratigraphic succession exposed to glacial erosion. Sylvanidium? hawbanense, which is one of the acritarchs found in glacial sediments, is newly described from Arabian Upper Ordovician strata.     

New Rugosa taxa from the Middle Devonian Subpolar Urals

New coelenterate taxa of the subclass Rugosa from the Middle Devonian of the western slope of the Subpolar Urals, the genus Syvjuphyllum (with the type species S. sokolovi gen. et sp. nov.) and the species Cosjuvia yushkini sp. nov., are described.     

The graptolite correlation of the North American Upper Ordovician Standard

The North American Upper Ordovician reference standard, the Cincinnatian Series, contains rich shelly microfossil faunas in its type area in the Cincinnati Region but graptolites are uncommon in most of its shallow-water calcareous sediments. Consequently, the graptolite correlation of this key sequence has remained uncertain, in part, even controversial. A review of both previously published recently discovered graptolite Occurrences in the type Cincinnatian, combined with data from the important graptolite successions in Oklahoma New York-Quebec, has not only clarified the graptolite correlation of the Cincinnatian but also added new data on the morphology taxonomy, the vertical horizontal distribution, of several taxa. The information now at hand indicates that the Edenian Stage correlates with the C. spiniferus Zone, the Maysvillian Stage with the C. pygmaeus lower middle P. manitoulinensis Zone, the Richmondian Stage with the upper P. manitoulinensis , the D. complanatus , possibly part of the C. inuiti Zone. Comparison between graptolite conodont biostratigraphic evidence reveals no apparent conflict. Correlations are proposed between Upper Ordovician North American stages, graptolite conodont zones, successions in Texas, Oklahoma, Sweden, European graptolite zones, British series.     

New gastropod species from the Pleistocene of the Upper Don basin

Two new species of fossil freshwater gastropods, Borysthenia intermedia sp. nov. and Tanousia krasnenkovi sp. nov., from the Tiraspol deposits of the Middle Pleistocene of the Oka-Don Plain are described. Parafossarulus priscillae (Girotti, 1972), Viviparus diluvianus (Künth, 1865), and Lithoglyphus jahni Urbasnski, 1975 are reported from Russia for the first time. The status of some problematic species of freshwater gastropods from the Middle Pleistocene of the region studied is discussed. The chronological and environmental significance of all gastropods studied is given.     

Ordovician reefs of the Ural Mountains,Russia: A review

Summary The Upper Ordovician reefs of the Urals were formed at a subsiding shelf-margin during an early Late Ashgillian (Sur’ya time interval) regressive phase. Reefs of this age were studied in detail from the western slope of the Northern, the Subpolar and the Polar Urals with respect to lithofacies, biotic composition and paleogeographical patterns. The thickness of the reefs varies between 100 and 500 m. The backreef areas are characterized by lagoons with increased salinity and sabkha development. Microbial associations and a diverse algal flora (Cyanophyta, green and red algae and alga incertae sedis) are the main constituents of reefal boundstones. Tabulate and rugose corals, heliolitids, calcareous sponge-like fossils, bryozoans and problematic hydroids were also part of the reef communities. Each reef exhibits a characteristic framework-building association. Reef development was terminated by a rapid and abrupt sea-level rise at the end of the middle Upper Ashgillian connected with the global Late Ordovician glaciation.     

A new coral genus (Tabulata) from the Upper Devonian of the Subpolar Urals

A new tabulate genus, Sokolovia gen. nov., with the type species B. pershinae sp. nov., from the Upper Devonian (Lower Famennian Substage) of the western slope of the Subpolar Urals is described.     

New early Ordovician calcareous foraminifers of the middle Tien Shan

Twelve new species of calcareous foraminifers of the genera Parathurammina, Parathuramminites, Bisphaera, Archaesphaera, and Vicinesphaera from the Lower Ordovician of the Middle Tien Shan are described.     

Chitinozoan biostratigraphy of the Upper Ordovician Fågelsång GSSP, Scania, southern Sweden

In the Fågelsång section, the new Global Stratotype Section and Point (GSSP) for the base of the Upper Ordovician Series, 22 samples from the E14 (a, b and c) and E15 outcrops have been studied for chitinozoans. They yield rich and rather diverse species assemblages of this microfossil group. The approximately 16-m-thick sampled part of the section can be subdivided, from bottom to top, into two biozones and a subbiozone: the Laufeldochitina stentor zone, the Eisenackitina rhenana subzone and the Lagenochitina dalbyensis zone. The boundary between the lowest biozone and the subzone is situated 1.7 m below the marker “Fågelsång Phosphorite” bed, thus only slightly lower (0.3 m) than the base of the Nemagraptus gracilis graptolite biozone, which defines the base of the Upper Ordovician Series. The base of the L. dalbyensis zone is located just above the Fågelsång Phosphorite, remarkably low compared to the graptolite biostratigraphy. It is evident that the chitinozoan biozonation provides an additional tool to recognise the base of the Upper Ordovician in this section. Furthermore, a distinct faunal change is shown across the phosphorite bed, possibly indicating a hiatus.     

An opportunistic Upper Ordovician trilobite assemblage from Missouri

The trilobite Ampyxina bellatula (Savage) occurs in monospecific accumulations within dolomitic strata of the shaly Upper Ordovician Maquoketa Group of northeastern Missouri. A dolomitic stratum containing 97 complete individuals and considerable amounts of pyrite exhibited no associated benthic megafauna. The strata appear to be equivalent to the Scales Formation of the Maquoketa Group of northern Illinois and eastern Iowa which have been interpreted as having been deposited in a muddy environment with severe circulation restrictions. The presence of the monospecific trilobite association in what otherwise would be an unfossiliferous stratigraphic sequence is interpreted as being the result of opportunistic strategy by this trilobite in a resource-rich unstable environmental setting. The association was apparently cat-astrophically buried, thus preserving the age structure of the holaspid population. A size-distribution histogram of the holaspid population indicates that reproduction may have occurred in periodic bursts with lesser reproductive events occurring during the intervening periods. Holaspid survivorship information indicates that mortality was low during the early holaspid stage and increased with age. Anomalous single bed trilobite accumulations of various age rocks have been documented by numerous authors. These may in part be analogous to the Ampyxina occurrence in the Maquoketa Group of Missouri, and be attributable to opportunistic strategies.     

Establishment and development of patch reefs in the intracratonic Ordovician sequence near Chicoutimi, Quebec

Patch reefs occur near the top of the transgressive sequence of Ordovician Trenton Group limestones in the Chicoutimi area of Quebec, eastern Canada. Despite their small sue, these reefs comprise diverse assemblages dominated by bryozoans, corals, stromatoporoids and receptaculitid algae. Pelmatozoans and gastropods are also conspicuous. The reefs were initiated and grew in a fully marine, open shelf setting. Available substrates varied from loose skeletal lenses to soft, firm or hardened bioturbated wackestones, and the earliest stages of reef growth reflect this heterogeneity. Loose or less firm substrates were colonised by bryozoans and pelmatozoans and/or by receptaculitids, which, together with accessory organisms, stabilised the sediments and provided the basis for further reef development. The resultant firmer, slightly elevated substrates provided sites for attachment of stromatoporoids and colonial corals which spread over earlier reef organisms and sediments and dominated the later stages of reef growth. On hardened areas of sediment, stromatoporoids and corals colonised the surface directly and the early stabilising stage of reef growth is absent. The compositions and developmental stages of these Trenton Group reefs are comparable with those seen in broadly contemporaneous and often larger reefs elsewhere, and are among the earliest in which corals played an important role.     

Discovery of Lower Famennian Stromatoporoidea in the Kuznetsk Basin

Stromatoporoidea recorded for the first time from the Lower Famennian of the Kuznetsk Basin are described. The assemblage studied includes three new species, Actinostroma quasifenestratum sp. nov., Kyklopora tomiensis sp. nov., and Coenostroma rarum sp. nov., and three previously known species of different genera.     

Methodological advances in the use of faunal gradient analysis for regional high-resolution correlation in the type Cincinnatian Series (Upper Ordovician)

Historically, strata of the type Cincinnatian Series have not been correlated with precision because of an assumption that individual beds are not regionally persistent. In an effort to establish a regional chronostratigraphic framework for the Cincinnatian, a technique has been developed recently that uses gradient analysis of fossil abundance data to compare vertical trends in biotic composition among five localities that span a distance of 65 km. Using faunal data to establish correlations below the meter-scale has been problematic, because variability in the lateral expression of many beds has made it difficult to sample equivalent fossiliferous strata at all localities.

Here, this correlation technique is modified in an attempt to draw meter-scale correlations of the Kope and lower Fairview Formations. This modification involves grouping fossil census samples into stratigraphic intervals, such that the stratigraphic spacing of samples is identical at all localities. Samples are binned using two protocols: one that groups samples into 0.5 m thick intervals, and another that groups samples by lithologically delineated meter-scale cycles. Gradient analysis of binned samples using both protocols produces curves in which patterns in faunal composition below the meter-scale can be correlated with success only for limited intervals at select localities. The inability to consistently correlate at high resolution might indicate that correcting for regional variations in the expression of strata is not enough. Alternatively, the ability to correlate some intervals below the meter scale suggests that other problems, such as fine-scale lateral variations in biotic composition, might be impeding correlation. In addition, gradient analysis techniques are employed at a new locality to extend the lateral breadth of the existing, broader-scale chronostratigraphic framework by approximately 40 km. Although interpreted to be in an up-ramp direction from previously measured localities, this new locality does not have an overall shallower water faunal assemblage until just below the Kope/Fairview contact.     

Probing the deep shelf: a Lagerstätte from the Upper Ordovician of Girvan, southwestern Scotland

A deep‐water Konservat Lagerstätte from the lower Caradoc (Sandbian) at Girvan is dominated by the trilobite Diacanthaspis trippi, the carpoids Anatifopsis n. sp.? and a new genus of ctenocystoid together with the polyplacophoran Solenocaris solenoides and the brachiopod Onniella williamsi. Most of these are multi‐element organisms, with many specimens preserved in an articulated state in finely laminated rocks, indicating minimal disturbance and suggesting that the fauna is largely an in situ association. It contains few of the species known from other deep‐water sites of similar age at Girvan which contain diverse assemblages of trilobites and brachiopods absent from the Lagerstätte. The taphonomy of the site indicates preservation by rapid burial followed by early diagenesis under dysaerobic conditions. It provides a ‘taphonomic window’ on otherwise unknown faunas from distal shelf facies on the Ordovician Laurentian margin, and, moreover, is an important reminder of the hidden biodiversity that resided in thin‐shelled, multi‐element organisms.     

A Curious Rusophycus (Arthropod Ichnofossil) Assemblage from the Upper Ordovician of Ontario,Canada

The ichnogenus Rusophycus includes a wide range of short bilobate excavations generally attributed to variable feeding behaviors of arthropods, especially trilobites. An unusual Rusophycus assemblage from Upper Ordovician Georgian Bay Formation in Ontario departs radically from previously described examples and presents new challenges for understanding the behavior represented by these traces. This specimen is unique in the arrangement of multiple Rusophycus burrows in a circular, lens-shaped array (as opposed to a linear or random arrangement typical of other Rusophycus assemblages). The size and shape of the individual Rusophycus components are consistent with traces attributed to the coeval trilobite Flexicalymene. Multiple Rusophycus assemblages likely reflect aggregations of trilobites in response to a local concentration of food. The topology of this particular Rusophycus assemblage suggests that the trilobites opportunistically exploited a rich and narrowly restricted food source, perhaps the decaying remains of a buried organism.     

New species of syringoporids from Eifelian deposits of the Subpolar Ural Mountains

A high diversity of tabulate corals in Eifelian deposits of the Syv“yu River basin (Subpolar Urals) is reported for the first time. Most of the corals are represented by colonies of syringoporids. Five new species Syringopora parva sp. nov., S. insueta sp. nov., S. indecora sp. nov., Tetraporinus syvjuensis sp. nov., and Armalites serotinus sp. nov. are described.