Bryophytes from the southeastern Yukon Territory,Canada

One hundred and forty-five taxa of bryophytes are reported from the extreme southeastern Yukon. Eight species of Hepaticae and 13 of mosses represent new records for the Yukon, which are Calypogeia suecica (S. Arnell & Perss.) K. Müll., Chiloscyphus pallescens (Ehrh.) Dum., Conocephalum conicum (L.) Dum., Jamesoniella autumnalis (DC.) Steph., Pellia neesiana (Gott.) Limpr., Ptilidium pulcherrimum (Web.) Hampe, Riccardia palmata (Hedw.) Carruth., Tritomaria exsecta (Schmid.) Schiffn., Amblystegium varium (Hedw.) Lindb., Brachythecium rivulare B.S.G., B. rutabulum (Hedw.) B.S.G., Bryum blindii B.S.G., Didymodon rigidulus Hedw., D. tophaceus (Brid.) Lisa, Drepanocladus trichophyllus (Warnst.) Podp., Hygroamblystegium noterophilum (Sull. & Lesq. ex Sull.) Warnst., Hygrohypnum molle (Hedw.) Loeske, Plagiomnium ciliare (C. Müll.) Kop., P. cuspidatum (Hedw.) Kop., Platydictya minutissimum (Sull. & Lesq. ex Sull.) Crum, and Pylaisiella selwynii (Kindb.) Crum, Steere & Anderson. Many of the other collections represent wide extensions of range within the Yukon Territory.     

A contribution to the Sphagnum (Sphagnaceae) flora of Paraguay

The genusSphagnum is reviewed for Paraguay, with an emphasis on recent collections. Five species and one variety are recognized:S. perichaetiale Hampe,S. magellanicum Brid.,S. pulchricoma C. Müll.,S. itabense Crum & Buck, sp. nov.,S. flaccidum Besch., andS. flaccidum var.lindmanii (Warnst.) Warnst.Sphagnum flaccidum is redescribed; it is illustrated, as areS. perichaetiale andS. itabense.     

Phylogenetic relationships between Bryum and supposedly closely related genera

Abstract

The phylogeny of the genus Bryum was studied using cladistic analyses under the maximum parsimony criterion of evolution of anatomical and morphological characters. Three analyses were made with 32 Bryum species plus 20 species from genera supposedly closely related to Bryum, and with Amblyodon dealbatus (Sw. ex Hedw.) Bruch & Schimp., Meesia uliginosa Hedw., and Leptostomum macrocarpum (Hedw.) Bach. Pyl., as outgroups. It is here suggested that under earlier systematic concepts the genus Bryum is paraphyletic. A clade with Bryum billarderi Schwägr., B. capillare Hedw., B. donianum Grev., B. russulum Broth. & Geh., Rhodobryum giganteum (Schwägr.) Paris, and R. keniae (Müll. Hal.) Broth. are circumscribed by spathulate stem leaves that are crowded in the stem apex, suggesting that the rosulate species of Bryum are more closely related to Rhodobryum than to the rest of Bryum. Stem leaf costae without stereids and spores that mature in the winter are synapomorphies for a clade with Anomobryum julaceum (P. Gaertn. et al.) Schimp. and Bryum argenteum (Hedw.). The tropical species B. cellulare Hook. and B. flaccum Wilson ex Mitt. appear in a clade with Plagiobryum zieri (Dicks. ex Hedw.) Lindb. and Synthetodontium pringlei Cardot. In one analysis, B. limbatum Müll. Hal., Epipterygium tozeri (Grev.) Lindb., Leptobryum pyriforme (Hedw.) Wilson, and Roellia roellii (Broth. ex Röll) H.A. Crum came out in a clade with Mniobryum atropurpureum (Wahlenb.) I. Hagen, Mnium hornum Hedw., Pohlia cruda (Hedw.) Lindb., P. longicollis (Hedw.) Lindb., and Pseudopohlia didymodontia (Mitt.) A.L.Andrews. It is here suggested that gametophytic features, such as the orientation and anatomy of the stem leaves and the appearance of vegetative propagules, are important for the internal relationships within the studied ingroup, whereas characters related to the sporophyte, especially those of the peristome, may obscure phylogenetic relationships. Most of the subgenera and the sections of Bryum, as defined by earlier authors, appear to be paraphyletic. However, due to the low stability of most clades it is suggested that analyses including anatomical, morphological, and molecular data are needed.     

中国丝瓜藓属直蒴组的分类学研究

采用经典分类学研究方法,对河北师范大学植物标本馆(HBNU)等国内多个标本馆保存的中国丝瓜藓属直蒴组(Pohlia Hedw.section Cacodon Lindb.ex Broth.)及其近缘类群植物标本共计130余号进行了分类学分析鉴定,并对所确定的中国产直蒴组物种的芽胞大小、形状及叶原基数量和长度等形态学特征进行比较分析,以期为中国丝瓜藓属直蒴组的分类学研究提供理论依据。结果表明:(1)经鉴定中国丝瓜藓属直蒴组植物共有8种——夭命丝瓜藓[Pohlia annotina(Hedw.)Lindb.]、糙枝丝瓜藓[P.camptotrachela(Ren.Card.)Broth.]、林地丝瓜藓[P.drummondii(Müll.Hal.)Andr.]、疣齿丝瓜藓[P.flexuosa Harv.]、纤毛丝瓜藓[P.hisae T.J.Kop.et J.X.Luo]、异芽丝瓜藓[P.leucostoma(BoschSande Lac.)M.Fleisch.]、念珠丝瓜藓[P.lutescens(Limpr.)H.Lindb.]和卵蒴丝瓜藓[P.proligera(Kindb.ex Breidl.)Lindb.ex Arnell]。(2)中国丝瓜藓属直蒴组植物的重要鉴别特征是在假根或叶腋处生有芽胞,而且芽胞形状以及叶原基形状和数量在直蒴组中存在差异,如念珠丝瓜藓具有假根生芽胞,而疣齿丝瓜藓具有两型芽胞。(3)对所鉴定的中国丝瓜藓属直蒴组8种植物的识别特征、生境及其与相近种的关系进行了详细描述,并绘制了形态特征墨线图。     

On the identities of some neglected species of Grimmiaceae from Îles Kerguelen

Taxonomic status of five species of Grimmia Hedw. (G. minuta Müll.Hal., G. suborthotrichacea Müll.Hal., G. defoliata Müll.Hal., G. genuflexa Müll.Hal., and G. orthotrichacea Müll.Hal.) is appraised. All these species correctly belong within the genus Bucklandiella Roiv. They are actually inseparable from one another and represent a single species for which B. orthotrichacea (Müll.Hal.) Bednarek-Ochyra & Ochyra is chosen as the final name. The remaining four names are considered its synonyms, all but G. genuflexa being new. All the names are lectotypified and some details of the leaves of G. minuta and G. defoliata are illustrated.     

A tribute to Dr Royce E. Longton

Abstract

The relationships within the Bryaceae, with emphasis on the genus Bryum, were studied based on morphological and anatomical characters and using cladistic methods. The analysis was performed with thirty-six species representing the different parts of the family, and with Funaria hygrometrica Hedw., Mnium hornum (Dicks.) Lindb., and Tayloria lingulata Hedw. as outgroups. The Bryaceae, and the subfamilies Bryoideae, Mielichhoferioideae, and Pohlioideae, as defined by several earlier authors appear to be paraphyletic. The genus Bryum seems to be paraphyletic, because Leptobryum pyriforme (Hedw.) Wils., Osculatia columbica De Not., and Rhodobryum giganteum (Schwaegr.) Paris, appear as ingroups within this genus when the tree is rooted with Funaria. Mnium hornum came out as the sister taxon of a clade including Pohlia cruda (Hedw.) Lindb. and P. longicollis (Hedw.) Lindb., whereas P. drummondii (Müll. Hall.) A.L. Andrews, appears not to be closely related to the other two Pohlia species studied here, making this genus paraphyletic. Mielichhoferia mielichhoferiana (Funck.) Loeske, appears as the sister taxon of Schizymenium bryoides Harv., suggesting that both these genera are paraphyletic. Overall, the stabilities of the clades are low and it is suggested that combined analyses of morphological, anatomical, and molecular data are needed to get better resolved and more stable trees.     

Chromosome studies on some Turkish species of Grimmia Hedw.: G. laevigata (Brid.) Brid., G. ovalis (Hedw.) Lindb., G. pulvinata (Hedw.) Sm., G. trichophylla Grev.

In this research, mitotic chromosome numbers in four species of Grimmia Hedw. from Turkey were reported. Mitotic chromosome numbers of three species : Grimmia laevigata (Brid.) Brid. (n = 26), Grimmia ovalis (Hedw.) Lindb. (n = 10), Grimmia pulvinata (Hedw.) Sm. (n = 26) were reported for the first time. Grimmia trichophylla Grev. (n = 13) showed the same chromosome number as in a previous study.     

The identity of Racomitrium obtusifolium (Musci,Grimmiaceae) from North America

Trichostomum obtusifolium P. Beauv. is lectotypified and the type material is briefly described and illustrated. It is shown that the species correctly belongs within the genus Anomodon Hook. & Tayl., not Racomitrium Brid., and it is synonymous with A. minor (Hedw.) Lindb. A brief nomenclatural note on A. minor is presented and its homotypic synonyms are listed.     

Taxonomic notes on African Orthotrichaceae I. New synonymy in Macromitrium

New synonymy and distribution records for the genus Macromitrium Brid. in sub-Saharan Africa are presented. Seven names are placed in synonymy, reducing the number of accepted taxa for Macromitrium in sub-Saharan Africa to 33. Macromitrium anomodictyon Cardot was found to be conspecific with M. serpens (Bruch ex Hook. & Grev.) Brid.; M. mauritianum Schwägr. and M. sanctae-mariae Renauld & Cardot are placed in synonymy with M. subtortum (Hook. & Grev.) Schwägr. and M. belangeri Müll.Hal., M. seriatum Paris & Broth., M. bequaertii Thér. & Naveau and M. trollii Dixon in Herz. are all placed in synonymy with M. sulcatum (Hook.) Brid. subsp. sulcatum. The precise lectotypification is implemented for six species within the genus. Macromitrium chloromitrium (Besch.) Wilbraham and M. orthostichum Nees ex Schwägr. are reported new to Tanzania and M. chloromitrium new to continental Africa.     

Does the family Meesiaceae (Musci) occur in Africa?

Meesia kenyae P. Varde, a species originally described in 1955 from the altimontane elevation on Mt. Kenya in tropical East Africa, is briefly assessed taxonomically and some details of the type material are illustrated. This species is considered to be conspecific with Ceratodon purpureus (Hedw.) Brid., a weedy, cosmopolitan and highly protean species. Because M. kenyae is the only representative of the Meesiaceae in Africa, this taxonomic conclusion results in the exclusion of this small family from the bryoflora of this continent. Accordingly, the Meesiaceae are represented in the tropics only by three species, namely Meesia triquetra (Jolycl.) Ångstr. in Papua New Guinea, M. longiseta Hedw. in Colombia and M. ulei Müll. Hal. in Brazil.     

Contribution À L'Étude De Quelques Bryoassociations De L'Étage Subalpin Dans Le Sud-Est De La France

Resume Dans le présent travail, l'auteur effectue une étude bryosociologique de la strate muscinale des rochers et des marécages de l'étage subalpin dans le Sud-Est de la France. Il envisage en particulier les blocs siliceux ombragés occupés par le Grimmietum elatioris (avecGrimmia elatior Bruch,Orthotrichum rupestre Schleich,Grimmia funalis (Schwaegr.) Schimp.,G. doniana Sm.,Dicranum sauteri Schimp.) ainsi que les parois suintantes de gneiss (Blindio-Amphoridietum) colonisées parBlindia acuta (Huds) B.e.,Amphoridium mougeotti B.e.,Fissidens osmundoides Hedw.,Saccobasis polita (Nees) Bruch,Gymnostomum rupestre Schleich. Les rochers calcaires en exposition froide portent, sur le pourtour de l'Argentera-Mercantour l'association àEncalypta streptocarpa Hedw. etPlagiopus oederi (Schwaegr.) Schimp. (Encalypto-Plagiopetum) très largement répandue dans les Alpes du Sud, alors qu'au voisinage des cascades appara?t une formation tufogène montagnarde àHymenostylium recurvirostre (Hedw.) Dix etSouthbya nigrella (De Not) Spruce. Les tourbières àSphagnum acutifolium Warnst,S. teres Angst,S. platyphyllum Warnst,S. recurvum P.B.,Aulacomnium palustre (Hedw.) Schwaegr. etMeesea triquetra (Hedw.) Angst. semblent inféodées, dans les régions étudiées, au vieux massif cristallin (Lac Nègre, Boréon, Vallée des Merveilles) où elles se rencontrent à partir de 1900 mètres d'altitude. Enfin, les marécages sur calcaire ou sur flysch portent des peuplements à base deBryum pallens Sw.Philonotis fontana (Hedw.) Schwaegr.,Cratoneurum commutatum (Hedw.) Roth. avec par endroitsMeesea uliginosa Hedw. etAmblyodon dealbatus (Dicks) P.B.

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Summary In this work, the author carries out a bryosociological study concerning the vegetation on rocks and bogs in the subalpine district of the high mountains of southern France. He considers particularly the siliceous blocks covered with carpets ofGrimmia elatior Bruch,Orthotrichum rupestre Schleich,Grimmia funalis (Schwaegr.) Schimp.,G. doniana Sm.,Dicranum sauteri Schimp. (Grimmietum elatioris) and the oozing gneiss-walls (Blindio-Amphoridietum) colonized byBlindia acuta (Huds) B.e.,Amphoridium mougeotti B.e.,Fissidens osmumdoides Hedw.,Saccobasis polita (Nees) Bruch., andGymnostomum rupestre Schleich. On calcareous rocks, in cold places, surrounding the Argentera-Mercantour, he found an association withEncalypta streptocarpa Hedw. andPlagiopus oederi (Schwaegr.) Schimp., (Encalypto-Plagiopetum) which is very common in the southern alps, and near waterfalls occurs a travertinic mountain-formation withHymenostylium recurvirostre (Hedw.) Dix andSouthbya nigrella (De Not) Spruce. The peat-bogs containingSphagnum acutifolium Warnst.,S. teres Angst,S. platyphyllum Warnst.,S. recurvum P.B.,Aulacomnium palustre (Hedw.) Schwaegr. andMeesea triquetra (Hedw.) Angst. seem to be particular to the old crystalline chain where we found them at an altitude of 1900 meters. Finally, calcareous or flysch-bogs present a cryptogamic vegetation withBryum pallens Sew.,Philonotis fontana (Hedw.) Schwaegr.,Cratoneurum commutatum (Hedw.) Roth. and the rareMeesea uliginosa Hedw. andAmblyodon dealbatus (Dicks) P.B.

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Zusammenfassung In dieser Arbeit, führt der Verfasser ein bryosoziologisches Studien der Felsen und der Moore in der subalpinen Stufe Süd-Ost Frankreichs aus. Er beschreibt namentlich die Silikatbl?cke, die im Schatten der W?lder von dem Grimmietum elatioris (mitGrimmia elatior Bruch.,Orthotrichum rupestre Schleich.,Grimmia funalis (Schwaegr.) Schimp.,G. doniana Sm.,Dicranum sauteri Schimp.) bedeckt sind, und die feuchten Gneissw?nde (Blindio-Amphoridietum), die vonBlindia acuta (Huds.) B.e.,Amphoridium mougeotti B.e.,Fissidens osmundoides Hedw.,Saccobasis polita (Nees) Bruch.,Gymnostomum rupestre Schleich kolonisiert werden. Auf den kalkhaltigen Bl?cken, in der N?he des Argentera-Mercantour traf er eine Moosgesellschaft mitEncalypta streptocarpa Hedw. undPlagiopus oederi (Schwaegr.) Schimp. (Encalypto-Plagiopetum), die in den Süd-Alpen sehr h?ufig ist. An den Quellen und Wasserf?llen erscheint eine tufogene Bildung der montanen Stufe mitHymenostylium recurvirostre (Hedw.) Dix undSouthbya nigrella (De Not) Spruce. In unserer Gegend, sind die Torfmoore mitSphagnum acutifolium Warnst.,S. teres Angst,S. platyphyllum Warnst,S. recurvum P.B.,Aulacomnium palustre (Hedw.) Schwaegr., undMeesea triquetra (Hedw.) Angst. dem alten kristallartigen Gebirgsstock eng belehnt, wo wir sie immer ab 1900 meter gefunden haben. Endlich, tragen die Kalk oder Flysch-Sümpfe eine besondere Kryptogamenvegetation mitBryum pallens Sw.,Philonotis fontana (Hedw.) Schwaegr.,Cratoneurum commutatum (Hedw.) Roth. und manchmal die seltenenMeesea uliginosa Hedw. undAmblyodon dealbatus (Dicks) P.B.

     

柳叶藓科2种1变种在中国的新记录

笔者在编研(中国苔藓志》第七卷的过程中,发现柳叶藓科中的圆叶湿原藓Calliergon megalophyl- lum Mikut．、毛叶镰刀藓 Drepanocladus trichophyllus (Warnst．)Podp．和牛角藓宽肋变种Cratoneuron filicinum (Hedw．)Spruce var．atro-virens (Brid．)Ochyra为中国首次记录,而沼生湿柳藓 Hygroamblystegium noterophi-lum (Sull．et Lesq．)Warnst．属错误鉴定,所有以前被定为该种的中国标本应改为牛角藓宽肋变种。     

Two species and one variety of the Amblystegiaceae (Musci) new to China

During our recent study of the Chinese Amblystegiaceae, we found that Calliergon megalophyllum Mikut., Drepanocladus trichophyllus (Warnst.) Podp. and Cratoneuron filicinum (Hedw.) Spruce var. atro-virens (Brid.) Ochyra are new to China. The occurrance of Hygroumblystegium noterophilum (Sull. et Lesq. ) Warnst. in China as previously recorded represents only a misidentification. All the Chinese specimens referred to this species should be identified as Crato-neuron filicinum (Hedw.) Spruce var. atro-virens (Brid.) Ochyra.     

Entosthodon mouretii (Corb.) Jelenc new to Europe

Entosthodon mouretii (Corb.) Jelenc, a species related to E. fascicularis (Hedw.) C. Müll., is referred for the first time in Europe, from samples collected in Spain. The Spanish specimens are described and illustrated.     

Vittia salina L.Hedenäs & J.Muñoz,sp. nov., a new moss from Argentina

Abstract

A survey of the species of Campylopus Brid. reported from Sri Lanka (Ceylon) is presented. Of the 34 species that have been reported from this Island 16 are accepted at present. Campylopus nilghiriensis (Mitt.) Jaeg. is identical partly with C. goughii (Mitt.) Jaeg., partly with C. zollingerianus (C. Müll.) Bosch &; Lac., C. pseudogracilis Card. &; Dix. with C. goughii (Mitt.) Jaeg., C. caudatus (C. Müll.) Mont., C. reduncus (Reinw. &; Hornsch.) Bosch &; Lac. and C. trachythecius (C. Müll.) Jaeg. with C. comosus (Reinw. &; Hornsch.) Bosch &; Lac., C. herzogii Broth., C. subtricolor Lor. and probably also C. nodijlorus (C. Müll.) Jaeg. with C. aureus Bosch &; Lac., C nietneri (C. Müll.) Jaeg. with C. involutusz (C. Müll.) Jaeg., C. subgracilis Ren. &; Card. ex Gangulee and C. latinervis (Mitt.) Jaeg. with C. gracilis (Mitt.) Jaeg., and C. laetus (Mitt.) Jaeg. with C. savannarum (C. Müll.) Mitt. C. pterotoneuron (C. Müll.) Jaeg. is reduced to a variety of C. umbellatus (Arn.) Par. The occurrence of C. exasperatus Brid. on Sri Lanka could not be confirmed. Campylopus flagelliferus (C. Müll.) Jaeg. is reported as new to Sri Lanka.     

Observations of spore morphology of some Pottiaceae Schimp. species (Bryophyta) in Turkey

The spores of Syntrichia ruralis (Hedw.) F. Weber and D. Mohr., S. princeps (De Not.) Mitt., S. subulata (Hedw.) F.Weber and D.Mohr var. subulata, S. subulata (Hedw.) F.Weber and D.Mohr var. angustata (Schimp.) J.J. Amann and S. subulata (Hedw.) F.Weber and D.Mohr var. graeffii Warnst. were studied by light and scanning electron microscopy for the first time. The apertural region consists of a leptoma in all spores. All taxa of the family are uniform in their spore morphology. The spores of the five taxa are of granuloid type. The spore wall of the Pottiaceae family includes sclerine (the distinction between exine and perine may be difficult to define) and intine. The taxonomy of the genus Syntrichia is discussed on the basis of its spore morphology.     

甘肃省丛藓科植物新资料

通过对甘肃省金昌市永昌县境内龙首山及周边湿地苔藓植物进行调查鉴定,报道了甘肃省分布的丛藓科(Pottiaceae)1个新记录属——盐土藓属(Pterygoneurum Jur.)和4个新记录种——卵叶盐土藓[P.ovatum(Hedw.)Dix.]、盐土藓[P.subsessile(Brid.)Jur.]、卷叶墙藓[Tortula atrovirens(Smith)Lindb.]和狭叶链齿藓[Desmatodon cernuus(Hueb.)BruchSchimp.]。标本存放于内蒙古大学植物标本馆(HIMC)。     

The identity of the Pleistocene mosses Drepanocladus minnesotensis and Neocalliergon integrifolium (Amblystegiaceae)

The type specimens ofDrepanocladus minnesotensis Williams andNeocalliergon integrifolium Williams are shown to beD. aduncus var.kneiffii (B.S.G.) Monk. andScorpidium scorpioides (Hedw.) Limpr., respectively. These fossils, which were originally obtained from late glacial sediments in Minneapolis, Minnesota, together with other mosses identified from the deposit, indicate that deposition probably took place in a shallow depression filled with calcareous water. Pleistocene mosses from near Bronson, Minnesota, originally namedD. minnesotensis are reidentified asD. aduncus (Hedw.) Warnst. andCalliergon giganteum (Schimp.) Kindb.     

Observations on Chromosomes of Fourteen Moss Species from the Qinling Range,China

The chromosome numbers in fourteen moss species from the Qinling Range, and karyotype of Mnium lycopodioides are reported in this paper. Immature capsules were used as material. Bryum capillare L. ex Hedw was found. to have n = 10; Mnium lycopodioides Schwaegr. n = 6 with the karyotype formula n = 6 = 3m+3sm (1SAT); Plagiomnium maximoviczii (Lindl.)T. Kop. n=6; Bartramia halleriana var. elongate Turn. n=10+2m; Thuidium delicatulum (Hedw.) Mitt. n= 11; Thuidium philibertii Limpr. n= 10+m; Hygrohypnum ochraceum (Wils.) Loesk. n=10; Drepanocladus uncinatus (Hedw.) Warnst. n = 7+3m; Campylium chrysophyllus (Brid.) J. Lange. n = 10; Brachythecium starkei (Brid.) B. S. G. n=6; Brachythecium buchananii (Hook.) Jaeg. n=6; Brotherella yokohamae (Broth.)Broth. n= 9+2m; Gollamia neckerella (C. MÜll. )Broth. N=7; Pogonatum fastigiatum Mitt. n = 7. No report on the chromosome number of Brotherella yokohamae and Gollania neckerella has been found. The chromosome numbers in Brachythecium starkei andBrachythecium buchananii (Hook.) Jaeg. are different from previous reports, while the others are in accordance with the previous reports.