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The use of biomimetic tandem flapping foils for ships and underwater vehicles is considered as a unique and interesting concept in the area of marine propulsion.The flapping wings can be used as a thrust producing,stabilizer and control devices which has both propulsion and maneuvering applications for marine vehicles.In the present study,the hydrodynamic performance of a pair of flexible flapping foils resembling penguin flippers is studied.A ship model of 3 m in length is fitted with a pair of counter flapping foils at its bottom mid-ship region.Model tests are carried out in a towing tank to estimate the propulsive performance of flapping foils in bollard and self propulsion modes.The same tests are performed in a numerical environment using a Computational Fluid Dynamics (CFD) software.The numerical and experimental results show reasonably good agreement in both bollard pull and self propulsion trials.The numerical studies are carried out on flexible flapping hydrofoil in unsteady conditions using moving unstructured grids.The efficiency and force coefficients of the flexible flapping foils are determined and presented as a function of Strouhal number.  相似文献   

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The effect of the false caudal eyespots (ocelli) of fish predators on guppy inspection behaviour was tested by use of predator ( Crenicichla ) models with and without a caudal ocellus. When a false eyespot was present, inspecting guppies spent significantly less time near the tail compared to a control model without a tail eyespot.  相似文献   

4.
A new species of Akanthepsilonema and the first-stage juvenile of Glochinema trispinatum are described. Furthermore, additional morphological information is provided for Triepsilonema tripapillata. Animals originate from a cold-water coral degradation zone in the Porcupine Seabight area (North-East Atlantic Ocean). Akanthepsilonema sinecornibus sp. n. differs from A. heUeouetae in number of body annules, sexual dimorphism in amphid size, absence of copulatory thorns in males, absence of large spines and horns, shape of the copulatory apparatus, and position of ambulatory setae relative to vulva in females. The genus diagnosis for Akanthepsilonema is adjusted to incorporate the new species. Akanthepsilonema mainly differs from every other genus in the family by the combination of six rows of ambulatory setae situated around the vulva in females and eight subcephalic setae not displaced toward the anterior part of the head capsule. Small differences between the Papua New Guinea and the Porcupine Seabight populations of T. tripapillata indicate minimal intraspecific variability. Second-stage juveniles from Papua New Guinea have two rows of three ambulatory setae, whereas Porcupine Seabight specimens have two rows of four ambulatory setae. First- and fourth-stage juveniles of T. tripapillata are described for the first time. Literature data and personal observations showed that the molting of first-stage juveniles into second-stage juveniles and of third-stage juveniles into fourth-stage juveniles involves a decrease in the number of body rings, resulting in a loss of flexibility which is possibly compensated for by the development (I-II) or the doubling of the number of rows (III-IV) of ambulatory setae. This decrease is also linked with the formation of the head capsule and the smooth tail tip, although intergeneric variability is evident. The molting of second-stage juveniles into third-stage juveniles and of fourth-stage juveniles into adults is also subject to intergeneric variability. The variability in the number and orientation of caudal gland outlets among different nematode taxa is discussed. The presence of separate outlets for the caudal glands seems to be widespread within the family Epsilonematidae and has also been observed in various other, unrelated taxa of free-living aquatic nematodes, although their arrangement in Epsilonematidae is opposite. This aberrant arrangement is probably related to the aberrant locomotory pattern in this family.  相似文献   

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IS-Tlk/Kyo, a mutant derived from IS/Kyo strain, exhibits a kinked and/or short tail, in addition to the congenital lumbar vertebral anomaly. Homozygotes of Tlk dominant gene are known to die during embryonic development. We previously reported the morphological features of the skeleton in IS/Kyo and IS-Tlk/Kyo fetuses and of the heart in IS/Kyo fetuses [19]. This study was conducted to clarify the morphological features of the skeleton in both adult rats and of the heart in adult IS/Kyo rats. Ventricular septal defect (VSD) was observed in 3 out of 10 IS/Kyo rats. Neither splitting of lumbar vertebra and supernumerary rib (in both strains) nor fused or absent caudal cartilage (in IS-Tlk/Kyo strain) was detected in adult rats. Fusion of lumbar vertebrae was observed in almost all specimens together with lumbarization of sacral vertebrae in a few specimens in both adult rats as well as fusion of sacral and caudal vertebrae only in adult IS-Tlk/Kyo rats. In addition, a severe reduction in the ossified sacral and caudal vertebrae was noted in adult IS-Tlk/Kyo rats (mean number: 20.6) and IS/Kyo rats (31.8), and the difference was similar to that in the length of sacral and caudal vertebrae. These results suggest that the Tlk gene may be involved in both the congenital and acquired abnormal formation of the lower vertebral centra as well as the persistent occurrence of VSD by the background gene in IS/Kyo strain.  相似文献   

6.
The relative viscosities of acetic acid solutions of chitosan at varied temperatures were measured in a common glass capillary Ubbelohde viscometer in the range from dilute down to extremely dilute concentration. With the aid of the newly proposed theory related to the effect of solute adsorption on the relative viscosity measurement, the experimental data was analyzed, and some interesting parameters describing the interfacial properties of viscous capillary flow were deduced. The change of the conformation of the chitosan chains both free dissolved in solution and adsorbed on the glass capillary surface were discussed in detail. Furthermore, the morphologies of the adsorbed chitosan was also observed by AFM.  相似文献   

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Vertebrate cardiogenesis is believed to be partially regulated by fluid forces imposed by blood flow in addition to myocardial activity and other epigenetic factors. To understand the flow field within the embryonic heart, numerical simulations using the immersed boundary method were performed on a series of models that represent simplified versions of some of the early morphological stages of heart development. The results of the numerical study were validated using flow visualization experiments conducted on equivalent dynamically scaled physical models. The chamber and cardiac cushion (or valve) depths in the models were varied, and Reynolds numbers ranging from 0.01 to 1000 corresponding to the scale of the early heart tube to the adult heart were considered. The observed results showed that vortex formation within the chambers occurred for Reynolds numbers on the order of 1-10. This transition to vortical flow appears to be highly sensitive to the chamber and cushion depths within the model. These fluid dynamic events could be important to induce shear sensing at the endothelial surface layer which is thought to be a part of regulating the proper morphological development and functionality of the valves.  相似文献   

9.
Fishes suspended in water are subject to the complex nature of three-dimensional flows. Often, these flows are the result of abiotic and biotic sources that alter otherwise uniform flows, which then have the potential to perturb the swimming motions of fishes. The goal of this review is to highlight key studies that have contributed to a mechanistic and behavioural understanding of how perturbing flows affect fish. Most of our understanding of fish behaviour in turbulence comes from observations of natural conditions in the field and laboratory studies employing controlled perturbations, such as vortices generated in the wake behind simple geometric objects. Laboratory studies have employed motion analysis, flow visualization, electromyography, respirometry and sensory deprecation techniques to evaluate the mechanisms and physiological costs of swimming in altered flows. Studies show that flows which display chaotic and wide fluctuations in velocity can repel fishes, while flows that have a component of predictability can attract fishes. The ability to maintain stability in three-dimensional flows, either actively with powered movements or passively using the posture and intrinsic compliance of the body and fins, plays a large role in whether fish seek out or avoid turbulence. Fish in schools or current-swept habitats can benefit from altered flows using two distinct though not mutually exclusive mechanisms: flow refuging (exploiting regions of reduced flow relative to the earth frame of reference) and vortex capture (harnessing the energy of environmental vortices). Integrating how the physical environment affects organismal biomechanics with the more complex issue of behavioural choice requires consideration beyond simple body motions or metabolic costs. A fundamental link between these two ways of thinking about animal behaviour is how organisms sense and process information from the environment, which determines when locomotor behaviour is initiated and modulated. New data are presented here which show that behaviour changes in altered flows when either the lateral line or vision is blocked, showing that fish rely on multi-modal sensory inputs to negotiate complex flow environments. Integrating biomechanics and sensory biology to understand how fish swim in turbulent flow at the organismal level is necessary to better address population-level questions in the fields of fisheries management and ecology.  相似文献   

10.
Genetic variation and gene flow in the scleractinian coral Goniastrea aspera (Verrill), found around the north–south Okinawa Islands, were studied using allozyme and starch gel electrophoresis. The relative contribution of sexual and asexual reproduction to recruitment was determined. Analysis of multilocus genotypes of samples, collected at least 3 m apart, identified a high number of unique genotypes (NG) relative to the number of individuals sampled (N) (mean NG: N=0.97±0.03 [SD]), and also highly observed genotypic diversity (GO) relative to expected genotypic diversity under Hardy–Weinberg equilibrium (GE) (mean GO: GE=0.95±0.06 [SD]). These results suggest that the collected G. aspera propagated predominantly by sexual reproduction in Okinawan populations. The UPGMA grouping of five populations in the Okinawa, based on Neis unbiased genetic distance, showed two clusters that were south and north Okinawa populations. AMOVA analyses that incorporated the data of populations from the Kerama and the Ishigaki Islands detected significant FSC values among the populations within groups in analyses distinguishing north–south Okinawa, Okinawa–Kerama, and Okinawa–Kerama–Ishigaki. Significant FCT values were neither detected between north–south Okinawa nor between the Okinawa–Kerama groups. The local-recruits hypothesis, which assumes that a substantial proportion of recruits is produced locally, appears to be applicable to coral populations in the Ryukyu Archipelago. On the other hand, detection of nonsignificant FCT values indicated the presence of genetic connectivity between north–south Okinawa and Kerama–Okinawa groups.  相似文献   

11.
目的:通过对血管中血液流动对血管的影响及血液内低密度脂蛋白(sLDL)微粒行为的模拟分析,研究动脉粥样硬化产生的血流动力学原因。方法:第一步,使用流体力学软件CFD,建立动脉血管弯曲分叉仿真模型;第二步,分析血液流动特性,跟踪血液中15—25纳米尺度范围类的sLDL粒子在动脉分叉模型中的流体力学行为,研究sLDL在血液流速稳定下在血管中的空间分布及流场特征分布。结果:血管起始段出现压强很高的区域。在动脉血管弯曲内侧处及分岔处的分支外侧血液流动较慢,并且在这些部位出现压强较高的区域。在血管弯曲外侧处及分岔点处,sLDL与血管壁发生碰撞的几率较其它位置较高,粒子在血管上沉积高发区域在这些部位呈斑块状分布。讨论及结论:在血管起始段的高压,可能是导致这一部分血管损伤,并进而引起动脉硬化形成的主要原因;在动脉血管弯曲外侧处及分岔点处出现的高压低速血流分布,一方面增大了血液中包括sLDL粒子在内的致病因子与血管壁的接触时间,另一方面则引起这些部位血清的侧漏加强,出现所谓的’浓度极化’现象,从而导致这些部位出现高浓度的sLDL分布,增大sLDL粒子与血管壁的接触几率;粒子在血管上沉积高发区域往往存在于动脉血管分岔点处,而在血管弯曲外侧处也有较高几率沉淀,呈斑状分布;长期性轻微性振动的剪切压力的作用使多数血管内皮细胞性质改变,促进动脉硬化形成。在动脉血管起始段、弯曲处及分岔点处血液的高压低速分布、sLDL粒子的高沉积率及低剪切应力等是动脉硬化产生及演化的重要因素.  相似文献   

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Collenchyma fibres from celery ( Apium graveolens L.) were extracted with detergent and phenol-acetic acid-water to leave the intact cell walls, free from active enzymes. Under a small, constant stress the cell wall fibres showed elastic and plastic extension and viscoelastic deformation, but viscous flow was observed only at high stresses close to the breaking stress. After complete removal of calcium ions with cyclohex-anediamine tetraacetic acid (CDTA) and incubation for 18 h, comparable levels of these extensibility components were observed at much lower stresses. However, partial removal of calcium ions with citrate did not increase the plastic, elastic or viscoelastic components even when the residual calcium was reduced to 3.5% of the exchange capacity. The breaking stress of the fibres was rather more sensitive to calcium removal, being reduced by 50% at 7% calcium saturation. CDTA-extracted fibres broke by cell separation at very low stress. These characteristics did not appear compatible with removal of calcium ions, or their displacement by protons, as a mechanism for auxin-induced growth in this material: however such mechanisms are not excluded in other tissues or under other conditions. Strong chelating agents which remove enough calcium to weaken cell walls should be avoided in experiments on other mechanisms of auxin-induced growth.  相似文献   

14.
The tail of the earliest known articulated fully skeletonized vertebrate, the arandaspid Sacabambaspis from the Ordovician of Bolivia, is redescribed on the basis of further preparation of the only specimen in which it is most extensively preserved. The first, but soon discarded, reconstruction, which assumed the presence of a long horizontal notochordal lobe separating equal sized dorsal and ventral fin webs, appears to have considerable merit. Although the ventral web is significantly smaller than the dorsal one, the presence of a very long notochordal lobe bearing a small terminal web is confirmed. The discrepancy in the size of the ventral and dorsal webs rather suggests that the tail was hypocercal, a condition that would better accord with the caudal morphology of the living agnathans and the other jawless stem gnathostomes.  相似文献   

15.
Segregation patterns observed in the progenies from 13 different crosses of the spike-tailed paradisefish Pseudosphromenus cupanus support the hypothesis that caudal peduncle banding is controlled by a major effect gene exhibiting the classical monohybrid Mendelian pattern of inheritance.  相似文献   

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The pitching-down flapping is a new type of bionic flapping,which was invented by the author based on previous studieson the aerodynamic mechanisms of fruit fly(pitching-up)flapping.The motivation of this invention is to improve the aerodynamiccharacteristics of flapping Micro Air Vehicles(MAVs).In this paper the pitching-down flapping is briefly introduced.Themajor works include:(1)Computing the power requirements of pitching-down flapping in three modes(advanced,symmetrical,delayed),which were compared with those of pitching-up flapping;(2)Investigating the effects of translational accelerationtime,Δτ_t,and rotational time,Δτ_r,at the end of a stroke,and the angle of attack,α,in the middle of a stroke on the aerodynamiccharacteristics in symmetrical mode;(3)Investigating the effect of camber on pitching-down flapping.From the above works,conclusions can be drawn that:(1)Compared with the pitching-up flapping,the pitching-down flapping can greatly reduce thetime-averaged power requirements;(2)The increase in Δτt and the decrease in Δτ_r can increase both the lift and drag coefficients,but the time-averaged ratio of lift to drag changes a little.And α has significant effect on the aerodynamic characteristicsof the pitching-down flapping;(3)The positive camber can effectively increase the lift coefficient and the ratio of lift to drag.  相似文献   

18.
In Dutch coastal dunes the relation was studied between the composition of the vegetation on the banks of pools and the nutrients supplied by the groundwater. Areas infiltrated with eutrophic river water were compared with non-affected areas. The floristic composition was interpreted according to classification systems of ruderal, nitrophilous tall hemicryptophytes and the conservation parameters richness and rarity. Of the three main macro-nutrients (orthophosphate, nitrate and potassium), the orthophosphate load shows the highest correlation with the vegetation variables. It is suggested that a reduction of orthophosphate during pre-purification of the infiltration water would reduce the dominance of the ruderals to the extent that the original dune slack species could return locally.Nomenclature of species follows Arnolds & Van der Meijden (1975).We thank G. J. de Bruyn, W. H. van Dobben, W. T. de Groot, W. J. ter Keurs, F. van der Meulen, W. H. van der Molen, M. A. W. Noordervliet, and I. S. Zonneveld for their comments on the study; J. Braak, J. Brakkee and R. van Nood for their assistance with the chemical analyses; A. R. Kaal for the translation, and G. P. G. Hock for the drawing of the figures.  相似文献   

19.
Most insects and hummingbirds can generate lift during both upstroke and downstroke with a nearly horizontal flapping stroke plane,and perform precise hovering flight.Further,most birds can utilize tails and muscles in wings to actively control the flight performance,while insects control their flight with muscles based on wing root along with wing's passive deformation.Based on the above flight principles of birds and insects,Flapping Wing Micro Air Vehicles(FWMAVs)are classified as either bird-inspired or insect-inspired FWMAVs.In this review,the research achievements on mechanisms of insect-inspired,hoverable FWMAVs over the last ten years(2011-2020)are provided.We also provide the definition,func-tion,research status and development prospect of hoverable FWMAVs.Then discuss it from three aspects:bio-inspiration,motor-driving mechanisms and intelligent actuator-driving mechanisms.Following this,research groups involved in insect-inspired,hoverable FWMAV research and their major achievements are summarized and classified in tables.Problems,trends and challenges about the mechanism are compiled and presented.Finally,this paper presents conclusions about research on mechanical structure,and the future is discussed to enable further research interests.  相似文献   

20.
doi: 10.1111/j.1741‐2358.2011.00525.x Effect of electromagnetic field on bone regeneration around dental implants after immediate placement in the dog mandible: a pilot study Background: Accelerating bone healing around dental implants can reduce the long‐term period between the insertion of implants and functional rehabilitation. Objective: This in vivo study evaluated the effect of a constant electromagnetic field (CEF) on bone healing around dental implants in dogs. Materials and methods: Eight dental implants were placed immediately after extraction of the first pre‐molar and molar teeth on the mandible of two male dogs and divided into experimental (CEF) and control groups. A CEF at magnetic intensity of 0.8 mT with a pulse width of 25 μs and frequency of 1.5 MHz was applied on the implants for 20 min per day for 2 weeks. Result and conclusion: After qualitative histological analysis, a small quantity of newly formed bone was observed in the gap between the implant surface and alveolar bone in both groups.  相似文献   

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