New Eocene dinoflagellate cysts from the southern edge of the Junggar Basin,Xinjiang, China

The Anjihaihe Formation in the southern edge of the Junggar Basin was previously considered a series of freshwater lacustrine depositions. However, abundant marine dinoflagellate cysts were recently recovered from the middle to upper part of the middle member of the formation. Two new genera, six new species and one new subspecies among the abundant dinoflagellate cysts are described and illustrated, i.e. Circulodinium? laeve sp. nov., Kaiwaradinium abbreviatum sp. nov., Spiniferites adnatus subsp. latispinus subsp. nov., Oligosphaeropsis accreta gen. et. sp. nov., Oligosphaeropsis complex gen. et. sp. nov., Oligosphaeropsis megaprocessa gen. et. sp. nov. and Tianshandinium biconicum gen. et. sp. nov. They are rare to common constituents of the dinoflagellate assemblage in the Anjihaihe Formation and may prove useful for regional biostratigraphic correlation and palaeoenvironment re-establishment.     

Discovery of Lower Famennian Stromatoporoidea in the Kuznetsk Basin

Stromatoporoidea recorded for the first time from the Lower Famennian of the Kuznetsk Basin are described. The assemblage studied includes three new species, Actinostroma quasifenestratum sp. nov., Kyklopora tomiensis sp. nov., and Coenostroma rarum sp. nov., and three previously known species of different genera.     

Neoselachians (Chondrichthyes,Elasmobranchii) from the Lower and lower Upper Cretaceous of north‐eastern Spain

Extensive sampling of several Barremian and Albian–Cenomanian levels across the Aguilón, Oliete and Aliaga subbasins of the Iberian Basin, north‐east Spain, yielded abundant material of new or so far poorly known neoselachians. The faunas consist of 16 different species, five of which represent new species and two new genera: Cantioscyllium brachyplicatum sp. nov. , Platypterix venustulus gen. et sp. nov. , Ptychotrygon pustulata sp. nov. , Ptychotrygon striata sp. nov. and Iberotrygon plagiolophus gen. et sp. nov. In addition, teeth of Heterodontus cf. H. carerens, Lamniformes indet., Pteroscyllium sp., Scyliorhinidae indet., Rhinobatos sp., Spathobatis sp., Belemnobatis sp., Ptychotrygon geyeri, Ptychotrygon sp. and Celtipristis herreroi are described. The new family Ptychotrygonidae is defined. The localities comprise palaeoenvironments ranging from lacustrine and shallow lake to open marine settings. Neoselachians are almost completely absent from continental settings in the Barremian, as a result of prevailing freshwater conditions, but became more abundant in marine strata. The Albian–Cenomanian selachian assemblage is the most profuse and diverse of the three assemblages studied. It is dominated by small, benthic and near‐coastal taxa, for instance Cantioscyllium and Ptychotrygon, and contains several new species, including an endemic batoid, Iberotrygon plagiolophus gen. et sp. nov. © 2009 The Linnean Society of London, Zoological Journal of the Linnean Society, 2009, 155 , 316–347.     

PERMIAN OSTRACODS FROM THE LERCARA FORMATION (MIDDLE TRIASSIC–CARNIAN?), SICILY,ITALY

Abstract: A rich, diverse Permian ostracod fauna has been recovered from the red and grey, laminated shales and quartz‐rich shales of the Triassic Lercara Formation. Forty‐seven species have been identified, 13 of which are newly described here; they belong to 26 genera of which three are new: Anahuacia lercaraensis sp. nov., Aurigerites siciliaensis sp. nov., Bairdia portellaensis sp. nov., Cristanaria? katyae sp. nov., Fabalicypris gruendeli sp. nov., Lethiersa salomonensis gen. et sp. nov., Lethiersia sinusoventralis gen. et sp. nov., Portella trapezoida gen. et sp. nov., Siciliella elongata gen. et sp. nov., Siciliella infernespinosa gen. et sp. nov., Siciliella prima gen. et sp. nov., Siciliella quadrata gen. et sp. nov., and Siciliella spinorobusta gen. et sp. nov. The assemblages contain or are composed of palaeopsychrospheric forms, which are regarded as index fossils for deep environments. The bathymetry of the different associations in life is evaluated.     

A Middle Miocene endemic freshwater mollusc assemblage from an intramontane Alpine lake (Aflenz Basin,Eastern Alps,Austria)

The mollusc fauna of the early Middle Miocene (Langhian) intramontane Alpine Lake Groisenbach is described for the first time. The shells derive from the Feistring Formation in the Aflenz Basin in Austria, which was covered by Lake Groisenbach. The assemblage is moderately diverse with 12 gastropod and 2 bivalve species, suggesting shallow lacustrine and fluvial settings. Among the gastropods, only Theodoxus crenulatus (Klein, 1853) is known from other Miocene localities, whilst all other species are documented so far only from Lake Groisenbach. None of the Early and Middle Miocene lake systems of the Alpine-Carpathian Foredeep and the Balkan Peninsula displays any faunistic resemblance with this new fauna. Even coeval lake faunas from the close-by Graz Basin have no species in common with Lake Groisenbach. This pattern points to a surprising endemicity and biogeographic fragmentation in the Central European freshwater systems during the Early and Middle Miocene. The uniqueness of the newly described fauna is also indicated by the completely erratic occurrence of the otherwise African-Mediterranean genus Bulinus, which is unknown from all other central European Miocene freshwater systems. Emmericia roetzeli Harzhauser and Neubauer nov. sp., Nematurella zuschini Neubauer and Harzhauser nov. sp., Romania fastigata Neubauer and Harzhauser nov. sp., Odontohydrobia groisenbachensis Neubauer and Harzhauser nov. sp., Odontohydrobia pompatica Neubauer and Harzhauser nov. sp., Odontohydrobia styriaca Harzhauser and Neubauer nov. sp., Planorbis austroalpinus Harzhauser and Neubauer nov. sp., Gyraulus sachsenhoferi Harzhauser and Neubauer nov. sp., Bulinus corici Harzhauser and Neubauer nov. sp., Ferrissia crenellata Harzhauser and Neubauer nov. sp. and Stagnicola reinholdkunzi Harzhauser and Neubauer nov. sp. are introduced as new species.     

A new coral‐associated decapod assemblage from the Upper Miocene (Messinian) Upper Coralline Limestone of Malta (Central Mediterranean)

Abstract: A rich coral‐associated decapod assemblage is recorded from the ‘Depiru Beds’ of the upper part of the Upper Coralline Limestone (Messinian, Upper Miocene), from the island of Malta. Nineteen species within 17 genera have been discovered, where 14 genera are new for Malta. Four new species are described, namely Micippa annamariae sp. nov., Pilumnus scaber sp. nov., Panopeus muelleri sp. nov. and Herbstia melitense sp. nov. Herbstia melitense sp. nov. constitutes the first record of the genus from the fossil record in the Mediterranean region. This discovery more than doubles the number of known fossil decapod species from Malta. The fossil bivalve Jouannetia (J.) semicaudata Des Moulins, 1830 and the extant decapod Maja goltziana D’Oliveira, 1888, are also recorded for the first time from Malta. Other Neogene coral‐associated decapod assemblages are investigated and correlated with the new assemblage from Malta. The migration of taxa between the Mediterranean region and the Paratethys, particularly during the Lower Badenian (Langhian), is evidenced by the strong affinity of the Maltese decapod assemblage with that of the Middle Miocene Badenian assemblages from Hungary, Poland and Ukraine. Upper Miocene, Messinian assemblages from Spain, Algeria and Morocco are also similar to that from Malta.     

New data on molluscs and their shell microstructures from the Middle Cambrian Gowers Formation, Australia

Abstract: Numerous new cases of preserved shell microstructure were discovered in molluscs from the Middle Cambrian Gowers Formation (Ptychagnostus atavus/Peronopsis opimus Zone, Floran Stage) in the Georgina Basin, Australia. The new data provide further evidence that, by the Middle Cambrian, molluscan shell microstructures were diverse, and many molluscs had a complex shell with multiple types of shell microstructure. In addition, many new occurrences of laminar microstructures are described herein. For many, the nature of these laminar microstructures is not known, but in three species the microstructure is foliated calcite, and in at least two the microstructure is more likely to have been calcitic semi‐nacre, a type of microstructure known in brachiopods and bryozoans but unknown in modern molluscs. This commonality among these three closely related lophotrochozoans underscores a similar mechanism of biomineralization. Moreover, these observations suggest a prevalence of calcite‐shelled lineages among molluscs from the Middle Cambrian, a time of calcite seas. In addition, the broad occurrence of laminar, nacre‐like microstructures in many of these fossils reveals how widespread these strong (fracture‐resistant) microstructures were in Middle Cambrian molluscs. Additionally, a few specimens of Yochelcionella preserve imprints of a bilaterally symmetrical pair of muscle scars. New taxa described here include Corystos thorntoniensis gen. et sp. nov., Yochelcionella snorkorum sp. nov., Yochelcionella saginata sp. nov., and Anhuiconus? agrenon sp. nov.     

Review of grylloblattid insects (Insecta: Grylloblattida) from the Solikamsk deposits of the Perm Region

The Solikamsk faunas of the Vorkuta series of the Pechora Basin and the Solikamsk Formation of the Perm Region of Russia are reviewed. New taxa are described from Tyul’kino locality (Solikamsk District): Artinska ufimica sp. nov., Sylvaprisca tyulkinensis sp. nov., Ufimoprisca gen. nov. (the type species Sylvaprisca alba Aristov, 2005), U. mica sp. nov., U. kamensis sp. nov. (family Lemmatophoridae), Sylvaella ovalis sp. nov. (family Liomopteridae), Ivakhosara prima sp. nov., and Megakhosarella prisca sp. nov. (family Megakhosaridae). Additionally, Uralioma sp. is described from Shchugor locality (Cherdyn’ District). Two species, Paraprisca solikamskensis Aristov, 2009 (family Lemmatophoridae) from Tyul’kino and Kolvidelia curta G. Zalessky, 1950 (family Ideliidae) from Pokcha locality (Cherdyn’ District)), are redescribed.     

A new clupeid genus (pisces, clupeiformes, clupeidae) from the Sarmatian of the Eastern Paratethys, Krasnodar Region

Two collections of clupeid fishes, which have previously been assigned to the species Sardinella sardinites (Heckel, 1850), from the Lower and Middle Sarmatian of the Pshekha River Basin (Western Ciscaucasia, Krasnodar Region) were studied. A new genus, Illusionella gen. nov., with two species, I. tsurevica sp. nov. and I. pshekhensis sp. nov., was described based on morphological characteristics. They are the most abundant species in Sarmatian fish assemblages of the Psekups-Belaya interfluve.     

Spore assemblages from the Lower Devonian Xujiachong Formation from Qujing,Yunnan, China

Abstract: The Lower Devonian Xujiachong Formation from the vicinity of Qujing City, Yunnan, China is interpreted as a terrestrial‐fluviatile‐lacustrine sequence. It contains important nonmarine biotas including plants, fish and invertebrates. The plants are particularly interesting as they include many endemic taxa. Dispersed spore assemblages have been recovered from the upper part of this formation. The spores are well preserved and of moderate thermal maturity. They are systematically described and four new species erected: Aneurospora xujiachongensis sp. nov., Chelinospora ouyangii sp. nov., Camptozonotriletes? luii sp. nov. and Leiozonospora xichongensis sp. nov. One new combination is proposed: Aneurospora conica (Ouyang and Lu) comb. nov. This is a rare report of a Lower Devonian dispersed spore assemblage from the South China Plate. Indeed, few dispersed spore assemblages of this age are known outside of Euramerica and Northern Gondwana. It is suggested that the Xujiachong Formation spore assemblages can all be equated to the polygonalis‐emsiensis Spore Assemblages Biozone (PE SAB) of Richardson and McGregor (1986) indicating an early (but not earliest) Pragian to ?earliest Emsian age. However, caution is urged, because biostratigraphical interpretation is difficult owing to distinct differences between dispersed spore assemblages from South China and Euramerica/Northern Gondwana. This almost certainly reflects palaeophytogeographical variation and regional endemism among early land plant floras on widely separated land masses. Palynofacies analysis supports a nonmarine origin for the deposits of the Xujiachong Formation, with the very rare marine palynomorphs that were encountered interpreted as reworked.     

Late Emsian Rutoceratoidea (Nautiloidea) from the Prague Basin,Czech Republic: morphology,diversity and palaeoecology

Abstract: Nautiloids of the superfamily Rutoceratoidea from the late Emsian (late Early Devonian) of the Prague Basin (Czech Republic) are commented upon. Species recognized include the hercoceratids Hercoceras mirum, H.? transiens, Ptenoceras proximum, P. nudum, P. minusculum and Anomaloceras anomalum, as well as the rutoceratids Adelphoceras bohemicum, Homoadelphoceras devonicans, Pseudorutoceras bolli and Goldringia? devonicans. In addition, four new species are described: Parauloceras regulare sp. nov., Roussanoffoceras chlupaci sp. nov., Otomaroceras sp. nov. and Goldringia sp. nov. Morphology and distribution patterns of Pragian and late Emsian rutoceratoid faunas from the Prague Basin are compared. They show that an increased diversity was accompanied by a higher level of specialization of rutoceratoids, which manifested itself in low abundance, increased facies dependence and greater variation in shell size during the Early Devonian. The evolution of sculpture and a contracted aperture, both regarded as protective adaptive features, was also examined, but no adaptive trend towards more pronounced sculpture and constriction of the aperture was found to have occurred in the Early Devonian. A more distinctive sculpture was, however, observed in shallow‐water assemblages of P. proximum in comparison with deeper‐water faunules, and two additional cephalopod species were examined in order to obtain comparative data. The presence of distinct sculpture patterns in coeval shallow‐ and deeper‐water assemblages suggests limited migration between them and consequently reflects some degree of territoriality in Devonian nautiloids. New data on early shell development in P. proximum are presented. During the Chote? Event, rutoceratoid generic diversity dropped dramatically, one family became extinct and the Early Devonian diversification of the group came to an end. The recovery of nautiloid faunas was slower than that of other cephalopods and associated, unrelated invertebrates. The absence of change in abundance patterns between Pragian and late Emsian rutoceratoid faunas, i.e. prior to and subsequent to ammonoid radiation, suggests that the appearance and radiation of the latter group in the early Emsian did not affect the structure of nautiloid assemblages, i.e. these two clades did not occupy the same niches.     

Early Cretaceous non‐marine Ostracoda from the North Falkland Basin,South Atlantic

Twenty‐seven species of well‐preserved and abundant Early Cretaceous non‐marine ostracod crustaceans were recovered from the North Falkland Basin. The assemblage is unusually diverse for a non‐marine palaeoenvironment and is sourced from cuttings samples collected during 2011 drilling of wells by Desire and Rockhopper Exploration, in the northern and southern areas of the basin. Ostracoda are entirely undocumented in published accounts from this basin, and all but one species appear to be new to science. For the new taxa, one new family (Alloiocyprideidae; type genus Hourcqia) is proposed and includes Hourcqia woodi sp. nov. Four new genera are erected: Falklandicypris gen. nov.; type species F. petrasaltata sp. nov., Gangamoncythere gen. nov.; type species G. colini sp. nov., Paraplesiocypridea gen. nov.; type species P. alloios sp. nov., and Musacchiocythere gen nov.; type species M. sarunata (Musacchio, 1978 ). Nine other species are described as new and are Falklandicypris desiderata, Clinocypris epacrus, Cypria poietes, Ilhasina? leiodermatus, Looneyellopsis tuberculatus, Theriosynoecum petasmathylacus, Theriosynoecum ballentae, Timiriasevia fluitans, and Vecticypris samesi. The remainder are left in open nomenclature due to paucity of material. The ostracod assemblage is largely restricted to the southern part of the basin (wells 25/5‐1 and 26/6‐1). In the northern part of the basin only four species, dominated by Vecticypris samesi, are present and with one possible exception are restricted there. There appears to have been little or no interchange of species suggesting that a barrier probably existed between the northern and southern regions. The more diverse southern assemblage indicates that more favourable conditions existed to the south. Stratigraphically, a distinct change in faunal composition recorded in both southern wells is likely to be an isochronous event correlatable across the southern area, and of an age no younger than Hauterivian.     

Diversity of Menispermaceae from the Paleocene and Eocene of South China

We present here the earliest known Asian fossil records of the Menispermaceae based on fossil fruits from Paleocene and Eocene localities in South China. A new genus and species, Paleoorbicarpum parvum sp. nov., and two new species of Stephania Loureiro, S. ornamenta sp. nov. and S. geniculata sp. nov., are recognized from Paleocene deposits of the Sanshui Basin, Guangdong, and a new occurrence of the widespread Eocene species Stephania auriformis (Hollick) Han & Manchester is recognized from the Maoming Basin, Guangdong. The Paleocene Stephania specimens described here represent the earliest fossil endocarp record of the Menispermaceae in eastern Asia. This discovery shows that the moonseed family had arrived in tropical and humid South China by at least the middle Paleocene, which provides important evidence for the origin and phytogeographic history of the family.     

Early to Middle Devonian miospores from northern Saudi Arabia

Well-preserved palynomorph assemblages are recovered from the Devonian Jauf and Jubah formations in five shallow boreholes in the northern part of Saudi Arabia. These fully cored boreholes overlap stratigraphically to form a 1640 ft composite sequence. Miospores dominate the palynological assemblages of most samples. The assemblages are mainly composed of trilete spores but also contain cryptospores and monolete spores. One new genus, sixteen new species and two new varieties of miospores are described from the studied assemblages: Artemopyrainconspicua nov. sp., Artemopyrarecticosta nov. sp., Camarozonotriletesfilatoffii nov. sp., Camarozonotriletesrugulosus nov. sp., Cymbohilatesbaqaensis nov. sp., Cymbohilatescomptulus nov. sp., Cymbohilatesheteroverrucosus nov. sp., Cymbosporitesasymmetricus nov. sp., Dibolisporitespilatus nov. sp., Dictyotriletesbiornatus nov. sp., Gneudnasporadivellomedia (Chibrikova) Balme, 1988 var. divellomedia, Gneudnasporadivellomedia (Chibrikova) Balme, 1988 var. minor nov. var., Latosporitesovalis nov. sp., Scylasporacostulosa nov. sp., Squamisporaarabica nov. gen. and sp., Stellatisporamulticostata nov. sp., Zonotriletesarmillatus nov. sp. and Zonotriletessimplicissimus nov. sp. Their stratigraphic distribution is compared to the well-established Devonian West European zonation of Streel et al. (1987) (Streel, M., Higgs, K.T., Loboziak, S., Riegel, W., Steemans, P., 1987. Spore stratigraphy and correlation with faunas and floras in the type marine Devonian of the Ardenne-Rhenish region. Review of Palaeobotany and Palynology 50, 211-219). A late Pragian-Givetian age is suggested for this sequence. No characteristic Eifelian taxa are recorded, but this could be explained by a gap in palyniferous samples.     

Rare bryozoans (Stenolaemata) with bilateral colonies from the Jurassic and Cretaceous of the East European Platform

New bryozoans characterized by a bilateral vertical colonies colonies—Cardioecia refuga sp. nov. (Tubuliporida), Elea lyapini sp. nov., E. troshkovensis sp. nov., and E. taylori sp. nov. (Melicerititida) from Middle Jurassic (Middle Callovian) of the Moscow region (Russia), and Biforicula legitima sp. nov. (Melicerititida) from the Upper Cretaceous (Lower Campanian) of the Southern Donets Basin (Ukraine)—are described. All species described in this paper belong to genera that have been recorded for the first time in the East European Platform. Some morphological structures in the colonies of these bryozoans and some distinctive features of the order Melicerititida that emphasize the difference of this order from bryozoans of other orders of the class Stenolaemata are examined.     

NEOSELACHIANS FROM THE UPPER CAMPANIAN AND LOWER MAASTRICHTIAN (UPPER CRETACEOUS) OF THE SOUTHERN PYRENEES, NORTHERN SPAIN

Abstract: Bulk sampling of upper Campanian to lower–middle Maastrichtian coastal and lagoonal deposits in five sections of the Tremp Formation in the south‐central Pyrenees yielded numerous neoselachian teeth. The fauna comprises nine taxa of which three species and one genus are new: Hemiscyllium sp., Lamniformes indet., Paratrygonorrhina amblysoda gen. et sp. nov., Coupatezia trempina sp. nov., Coupatezia sp., Coupatezia? sp., Rhombodus ibericus sp. nov. and Igdabatis indicus. The neoselachian fauna is dominated by small nectobenthic rays. This composition resembles assemblages known from the marine Upper Cretaceous, but differs from nearby localities of the Basque‐Cantabrian region and continental selachian associations of the French Pyrenees. The results indicate that Rhombodus might not be a reliable biostratigraphic marker for the Maastrichtian. The faunal composition suggests a shallow trans‐Tethyan connection between Eurasia and India at the end of the Cretaceous Period.     

Lower Carboniferous (middle Visean) foraminifers and algae from an interior sea,Southern Carnarvon Basin,Australia

A moderate diversity biota of foraminifers, algae and calcareous microproblematica is recorded from the Lower Carboniferous Yindagindy Formation within the intracratonic Southern Carnarvon Basin in Western Australia. Very shallow metahaline to hypersaline conditions are suggested. The palaeobotanical components are sporadic calcitarcha, tuberitinaceans, codiaceans Orthriosiphon? sp. 1, dasycladaleans? Koninckopora tenuiramosa, and algosponges Issinella devonica, Serrisinella cf. serrensis, Issinella? sp. 1, Kamaena cf. awirsi, Proninella? sp. and Stacheoides spp. The foraminiferal assemblage is dominated by a new koktjubinid taxon: Praekoktjubina yindagindyensis nov. gen., nov. sp. Subordinate associated foraminifers are more sporadic Salpingothurammina? sp. 1, Plectinopsis michelseni nov. gen., nov. sp., Rectopravina multifida nov. gen., nov. sp. (with 4 morphotypes), and scarce Endothyra spp. Despite many of these taxa being endemic, a middle Visean age is probable, based on: (1) the maximum global geographic extent of K. tenuiramosa reached during the middle Visean, (2) the disappearance of Plectinopsis nov. gen. in Palaeotethys and the Urals, and (3) the first occurrence of Praekoktjubina nov. gen. in Palaeotethys. An age no older than the middle Visean is supported by sparse conodonts from the Formation and comparison to episodes of carbonate deposition in the Bonaparte Basin, further north in the East Gondwana rift system. Absence of middle Visean archaediscoids may be due to high salinity. The following algal and foraminiferal families and superfamilies are emended: Scribroporellaceae (a lectotype is selected for Orthriosiphon saskatchewanense); Palaeospiroplectamminidae (due to the creation of Plectinopsis nov. gen.); Haplophragminoidea (due to the creation of Rectopravina nov. gen.); and Globivalvulinoidea and Koktjubinidae (due to the creation of Praekoktjubina nov. gen.).     

Ostracodes from the Upper Permian Khivach formation in Kolyma-Omolon Basin

Ostracodes Healdianella splendida Belousova, Paraparchites ganelini sp. nov., Paraparchites? hivachensis sp. nov. and Arcibairdia vodopadniensis sp. nov. are described from the Upper Permian deposits of the Khivach Formation (Kolyma-Omolon Basin, Eastern Siberia). Paraparchites species most closely resemble ancient Paleozoic ostracodes of the Boreal province. Valve morphology of Healdianella and Acribairdia points to their similarity to ostracodes from the upper levels of the Upper Permian of the Tethyan realm.     

Patterns of evolution and extinction in proetid trilobites during the late Devonian mass extinction event,Canning Basin,Western Australia

Abstract: In the early Late Devonian, terminal Frasnian proetid trilobites have previously only been known from Europe and North Africa. For the first time, a rich fauna of late Frasnian proetids is described from the Virgin Hills Formation, Canning Basin, Western Australia. Seventeen species in six genera are described, of which three are new: Rudybole gen. nov., Palpebralina gen. nov. and Canningbole gen. nov. A new subgenus, Chlupaciparia (Australoparia) subgen. nov. is also described. Fourteen of the species are new: Palpebralia initialis sp. nov., P. pustulata sp. nov., ?P. sp. nov. A, Rudybole depressa sp. nov., Palpebralina pseudopalpebralis sp. nov. (comprising the subspecies P. pseudopalpebralis pseudopalpebralis subsp. nov. and P. pseudopalpebralis ultima subsp. nov.), P. minor sp. nov., P. ocellifer sp. nov., Canningbole latimargo sp. nov., C. henwoodorum sp. nov., C. macromma sp. nov., Pteroparia extrema sp. nov., Chlupaciparia (Chlupaciparia) planiops sp. nov., Chlupaciparia (Australoparia) australis sp. nov. and C. (Australoparia) lata sp. nov. The subspecies Rudybole adorfensis angusta subsp. nov. is also described. The proetids range through conodont Zones 11–13b and terminate at the Upper Kellwasser Event, which marks the terminal Frasnian mass extinction event. Three of the six proetid lineages, Palpebralia, Canningbole and Pteroparia, show evolutionary trends of eye reduction. Two of the remaining lineages, Rudybole and Palpebralina, consist exclusively of blind taxa. The last, Chlupaciparia, also comprises forms with reduced eyes. The proetids show a stepped pattern of extinction during the late Frasnian, which correlate with two Kellwasser biocrises documented in European/North African Frasnian sections. The highest diversity preceded the Lower Kellwasser event that occurred at the end of conodont Zone 12 and saw the extinction of all species present in that zone. However, only one genus, Pteroparia, locally became extinct. A major higher‐level taxonomic mass extinction at the top of Zone 13b initiated the Upper Kellwasser extinction event. This included extinction at the generic level, with all five remaining genera becoming extinct, and at the family level, with the loss of the Tropidocoryphidae.     

Revision of the genus <Emphasis Type="Italic">Entactinosphaera</Emphasis> Foreman, 1963 and the new genus <Emphasis Type="Italic">Retientactinosphaera</Emphasis> gen. nov. (Paleozoic radiolaria)

The genus Entactinosphaera Foreman, 1963 is revised; it is shown to combine seven different genera, which are distinguished by morphological characters and belong to three orders of two radiolarian classes: the class Sphaerellaria, order Entactiniata (Bientactinosphaera, Entactinosphaera, Helioentactinia, Radiobisphaera), and class Spumellaria, orders Cancelliata (Retientactinosphaera, Magnisphaera) and Spongiata (Spongentactinia). New radiolarian taxa from the Middle Frasnian-Lower Famennian of the Timan-Pechora Basin and from the Lower Famennian of the Polar Ural Mountains, Russia, are described: Retientactinosphaera gen. nov., R. magnifica sp. nov., and R. clavata sp. nov.