Phylogenetic reassessment and biogeography of the Ectateus generic group (Coleoptera: Tenebrionidae: Platynotina)

Owing to the reinterpretation of its morphological synapomorphies, the taxonomic composition of the Ectateus generic group had been ambiguous. The present study scrutinized all existing taxonomic concepts of the group based on a cladistic analysis of the adult morphology of all of the Afrotropical platynotoid Platynotina genera. The phylogenetic relationships were reconstructed using parsimony and Bayesian inference. The results show that all previous taxonomic concepts of the Ectateus generic group concerned paraphyletic entities. The cladistic analysis revealed the following synapomorphies for the taxon: (1) presence of basal indentations of the pronotal disc, (2) ratio of prothorax width to its maximal height > 6.0, and (3) ratio of maximal height of the prothorax to total height < 0.3. Moreover, phylogenetic studies revealed the existence of the Upembarus generic group, a sister‐taxon group to the Ectateus generic group, within the Afrotropical platynotoid Platynotina. Autapomorphic and synapomorphic character mapping show that several taxonomic and nomenclatural changes are needed to consider the particular generic‐level entities traditionally assigned to Afrotropical platynotoid Platynotina as monophyletic lineages. The following taxonomic and nomenclatural adjustments are made in this paper: P teroselinus gen. nov. is erected to accommodate a single species that was previously assigned to Zidalus: Pteroselinus insularis comb. nov. Additionally, the following synonymies are proposed: Anchophthalmops (= Platykochius syn. nov. ), Angolositus (= Aberlencus syn. nov. , = Platymedvedevia syn. nov. ), Glyptopteryx (= Microselinus syn. nov. , = Quadrideres syn. nov. , = Synquadrideres syn. nov. ). In addition, Kochogaster is lowered in rank and is treated as one of the subgenera of Anchophthalmus. Moreover, Pseudoselinus is treated as a subgenus of Upembarus. An identification key to all Afrotropical platynotoid Platynotina genera and subgenera is presented. Zoogeographical analyses revealed the following dispersal barriers for the Ectateus generic group: (1) the Sahara (northern barrier); (2) the dry ecosystems of Botswana, Namibia, and South Africa (southern barrier); and (3) the Congolian rainforests (internal distributional gap). The ancestor of the taxon probably originated in East African ecoregions that predominantly contained wattletrees (acacias) and Commiphora Jacq. Moreover, past climate changes seem to have had a great impact on the observed generic distribution. © 2015 The Linnean Society of London     

First phylogenetic analysis of the tribe Oligaphorurini (Collembola: Onychiuridae) inferred from morphological data,with implications for generic classification

Oligaphorurini represent tribe of the subfamily Onychiurinae, which currently comprises 5 genera and 53 species. The present study evaluated the monophyly of Oligaphorurini genera. We investigated phylogenetic relationships among 39 species, representing all extant genera of Oligaphorurini. Both equal- and implied-weighting parsimony analyses were used in phylogenetic reconstruction. The cladistic analyses were based on comprehensive survey of adults’ morphological characters because specimens suitable for molecular studies were not available for the majority taxa. The phylogenetic analysis resulted in the recognition of a monophyletic Chribellphorura, and strongly supported non-monophyly of the previously recognized genera Archaphorura, Dimorphaphorura, Micraphorura, and Oligaphorura. The following new synonymy is recognized: Oligaphorura = Dimorphaphorura syn. nov., = Micraphorura syn. nov., = Archaphorura syn. nov. The general classification of Oligaphorurini is followed by the diagnoses of genera and key to the all known species.     

Jumping plant lice (Homoptera: Psylloidea) of the temperate neotropical region. Part 1: Psyllidae (subfamilies Aphalarinae, Rhinocolinae and Aphalaroidinae)

The psyllid-fauna of temperate and subantarctic South America comprises members of three families: Calophyidae, Triozidae and Psyllidae. Three subfamilies of the Psyllidae are revised in this paper: the Aphalarinae are represented by two species in two genera, one of which develops on Aquifoliaceae; the Rhinocolinae are represented by two congeneric species on Anacardiaceae while the Aphalaroidinae contain 38 species in seven genera trophically linked to the Compositae, Euphorbiaceae, Leguminosae, Myzodendraceae, Rhamnaceae, Solanaceae and Zygophyllaceae. The family Psyllidae (= Aphalaridae syn. nov. , =Spondyliaspididae syn. nov. ) and the constituent subfamily Aphalaroidinae (= Arepuninae syn nov. , = Ciriacreminae auct. pp) are redefined. Three genera and 30 species are described as new and two new generic, two new specific synonyms, and five new combinations are proposed. Information on larvae and host plant relationships is also given. Lectotypes are designated for eight species and a type-species is fixed for one genus. Keys are provided for the identification to species.     

The subfamily classification of the Micropezidae and the genera of Eurybatinae (Diptera: Schizophora)

The classification of the family Micropezidae is re-examined and a key is given to the five subfamilies now recognised, of which Calycopteryginae is described as new and Eurybatinae is raised from tribal rank. Metopochetini is a new tribe of Eurybatinae. A key is given to the nine genera now included in the Eurybatinae, of which Papeza is described as new. P.szentivanyi is a new species from Papua. Notes on the taxonomy and distribution of genera are included. The position of Anaeropsis Bigot in the Micropezidae is confirmed. New synonymies are established as follows: Crepidochetus Enderlein (= Gongylocephala Czerny syn. n.); Metopochetus bivittatus (Macquart) (= Calobata tenuipes Walker syn. n.). New generic combinations are as follows: Metopochetus terminalis (Walker) and M.compressus (Walker) from Calobata; Crepidochetus nigrifemur (Czerny), C.pallidus (Steyskal), and Cater (Steyskal) from Gongylocephala; Crosa fragilis (Walker) from Micropeza; Crosa nigriventris (Enderlein) and C.uneifera (de Meijere) from Eurybata.     

A revised generic classification of the tribe Sileneae (Caryophyllaceae)

A reclassification of the tribe Sileneae compatible with molecular data is presented. The genus Eudianthe ( E. laeta and E. coeli-rosa ) is restored. Viscaria, Ixoca (= Heliosperma ), and Atocion together form a well supported monophyletic group distinct from Silene and Lychnis , and are recognized at generic level. With Agrostemma and Petrocoptis , the number of genera in the tribe sums up to eight. The new combinations Silene samojedora, Silene ajanensis, Lychnis abyssinica, Atocion asterias, Atocion compacta, Atocion lerchenfeldiana , and Atocion rupestris are made.     

Phylogenetic relationships in the tribe Oxyptilini (Lepidoptera,Pterophoridae, Pterophorinae) based on morphological data of adults

The monophyly of the tribe Oxyptilini and phylogenetic relationships of the genera embraced in this tribe were examined using 171 (75 binary and 96 multistate) characters of adult morphology. The study material included 98 species of 30 genera, representing all previously recognized genera of Oxyptilini, together with the genera Sphenarches, Antarches, Diacrotricha, and Cosmoclostis, four species of Oidaematophorini, three species of Platyptiliini, as well as three and two other species belonging to Pterophorini and Exelastini respectively. Two Agdistis species were used as outgroups. The cladistic analysis resulted in six equally parsimonious trees. A majority of the recovered synapomorphic characters have previously been used in the taxonomy of the subfamily. However, 25 novel characters were found. The monophyly of Oxyptilini was supported, although only with homoplastic characters and low amounts of tree confidence; the genera Capperia, Procapperia, Paracapperia, Oxyptilus, Megalorhipida, and Trichoptilus were found to be nonmonophyletic; Sphenarches and Antarches were recovered as members of Oxyptilini; the two genera Cosmoclostis and Diacrotricha were placed out of Oxyptilini, inside the tribe Pterophorini; and close affinity of the genus Dejongia to Stangeia, Stenodacma, Megalorhipida, Trichoptilus, and Buckleria species was revealed. Four new combinations, Cosmoclostis lanceata (Arenberger) comb. nov. , Nippoptilia regulus (Meyrick) comb. nov. , Capperia tadzhica (Zagulajev) comb. nov. , and Buckleria negotiosus (Meyrick) comb. nov. are proposed; Capperia insomnis Townsend was considered as a senior synonym of Procapperia hackeri Arenberger syn. nov. , Buckleria negotiosus (Meyrick) as a senior synonym of Buckleria vanderwolfi Gielis syn. nov. , and Oxyptilus variegatus Meyrick syn. nov. as a junior synonym of Oxyptilus secutor Meyrick. © 2011 The Linnean Society of London, Zoological Journal of the Linnean Society, 2011, 163 , 484–547.     

A phylogenetic study of the Plantaginaceae

In a study based on morphological, embryological and chemical data of the Plantaginaceae, within the subclass Sympetalae or Asteridae, the superorder Lamianae is shown to be monophyletic. However, it was not possible to reconstruct the phylogeny within Lamianae or to find a sister-group for the monophyletic Plantaginaceae – Hydrostachyaceae for the latter is rejected. Three or rarely four genera have previously been recognized within Plantaginaceae, but in both cases Plantago appears as polyphyletic, which is considered unacceptable. Six clades are recognized as subgenera within Plantago: subgen. Plantago (c. 131 species), subgen. Coronopus (c. 11 species), subgen. Littorella (three species), subgen. Psyllium (c. 16 species), subgen. Bougueria (one species), and subgen. Albicans (51 species). Within P. subgen. Plantago, the paraphyletic sect. Plantago (c. 42 species) is found in all parts of the world except South America, New Zealand, Australia, and New Guinea, where sect. Oliganthos and sect. Mesembryniae vicariate. It is not possible to infer the phylogeny within sect. Plantago, for example between the endemic species from distant Pacific Islands; their common ancestor might be a species that once had a very wide distribution. Keys to genera, subgenera, sections and series are given. Only one genus, Plantago L., is recognized. The following proposals are made in the Appendix: P. ser. Oliganthos Rahn, ser. nov.; P. ser. Carpophorae (Rahn) Rahn, stat. nov. (= sect. C.); P. ser. Microcalyx (Pilg.) Rahn, stat. nov. (= sect. M.); P. unibrackteata Rahn, nom. nov. (=P. uniflora Hook.f. non L.); P. subgen. Littorella (P. J. Bergius) Rahn, stat. nov. (=Littorella P. J. Bergius); P. araucana Rahn, nom. nov. (=Littorella australis Griseb. non Plantago australis Lam.); P. americana (Fernald) Rahn, comb. nov.(=Littorella a.); P. subgen. Bougueria (Decne.) Rahn, stat. nov. (=Bougueria Decne.); P. nubicola (Decne.) Rahn, comb. nov. (=Bougueria n. Decne.); P. subgen. Albicans Rahn, subgen. nov. Ten lectotypes are selected.     

The cladistics and higher classification of the Pimpliformes (Hymenoptera: Ichneumonidae)

The monophyly of the ichneumonid clade Pimpliformes is established and the phylogenetic relationships of the eight component subfamilies are resolved. The clade (Acaenitinae + (Diacritinae + (Cylloceriinae + (Diplazontinae + Orthocentrinae)))) is the sister-lineage to the clade (Pimplinae + (Rhyssinae + Poemeniinae)). The Nearctic genus Cressonia Dasch is transferred to the Diacritinae from the Orthocentrinae. Tribes are not recognized in the Acaenitinae as the Coleocentrini (sensu Townes, 1971) is paraphyletic with respect to the Acaenitini. The Cylloceriinae is recognized as comprising three genera, Cylloceria Schiødte, Allomacrus Förster and Sweaterella gen.n. The Orthocentrinae, including the Helictinae of authors, is shown to be monophyletic, but the latter is clearly shown to be paraphyletic if the Orthocentrus genus-group is excluded. The Pimplinae comprises four monophyletic tribes: the Delomeristini, consisting of Delomerista Förster and Atractogaster Kriechbaumer; the Perithoini trib.n., which includes only Perithous Holmgren (= Hybomischos Baltazar syn.n.); the Pimplini, which includes the Theronia genus-group as well as the Pimpla genus-group; and the Ephialtini, which includes the Polysphinctini syn.n., a monophyletic group that previously rendered the restricted Ephialtini paraphyletic. The tribe Delomeristini is the sister-group to the clade (Ephialtini + (Perithoini + Pimplini)). The subfamily Poemeniinae is recognized as comprising three tribes: the Pseudorhyssini (trib.n.) which includes the single Holarctic genus Pseudorhyssa Merrill; the Rodrigamini (trib.n.) which includes only the Costa Rican genus Rodrigama Gauld; and the Poemeniini. The tribe Pseudorhyssini is the sister-group to the clade (Rodrigamini + Poemeniini). The phylogenetic inter-relationships of the genera of Poemeniini are resolved. A new genus from South Africa, Guptella (gen.n.) is described, and Achorocephalus Kriechbaumer is shown to be a synonym of Eugalta Cameron (syn.n.). The evolution of biological traits within the Pimpliformes is discussed with reference to the elucidated phylogeny, and zoogeographic patterns are outlined.     

Systematic revision of the genus Psithyristria Stål (Hemiptera: Cicadidae) with seven new species and a molecular phylogeny of the genus and higher taxa

The cicada genus Psithyristria Stål (Hemiptera: Cicadidae), endemic to Luzon, the Philippines, is reviewed, and its higher taxonomic affiliations are determined using morphology and molecular phylogenetics. Seven new Psithyristria species, P. grandis, sp.n. , P. albiterminalis, sp.n. , P. incredibilis, sp.n. , P. paraspecularis, sp.n. , P. peculiaris, sp.n. , P. moderabilis, sp.n. and P. genesis, sp.n. , are described. A key to the 12 known species is provided. The molecular phylogenetic analyses of Psithyristria and putatively related taxa showed some unexpected generic relationships, leading to changes in higher taxonomic placement for a number of Asian cicada genera. Psithyristria falls within the tribe Cicadini Latreille, 1802, and the tribe Psithyristriini Distant, 1905 syn.n. is therefore synonymized with Cicadini. The subtribe Psithyristriina Distant, 1905 stat.n. is recognized within Cicadini and defined for the genera Psithyristria, transferred from Psithyristriini, and Basa Distant, Pomponia Stål and Semia Matsumura, transferred from Cicadina, synonymizing Pomponiina Kato, 1932 syn.n. with Psithyristriina. The Cicadini subtribe Leptopsaltriina Moulton, 1923 is redefined to include the genera Terpnosia Distant, Euterpnosia Matsumura, Leptosemia Matsumura, Neocicada Kato, Puranoides Moulton, Leptopsaltria Stål, Tanna Distant, Purana Distant, Formosemia Matsumura, Maua Distant, Nabalua Moulton, Taiwanosemia Matsumura, Gudaba Distant and Calcagninus Distant, synonymizing Terpnosiina Kato, 1932 syn.n. The tribe Cicadatrini Distant, 1905 is recognized and redefined to include the genera Cicadatra Kolenati, Psalmocharias Kirkaldy, Mogannia Amyot & Audinet‐Serville, Nipponosemia Kato and Emathia Stål, synonymizing Moganniini Distant, 1905 syn.n. The nine Psithyristria species included in the molecular analyses have an uncorrected mitochondrial genetic distance of 2–9%, and most species appear to be related through a single radiation event that occurred approximately 5–10 Ma. No phylogenetic structure was found in two unusually divergent characters, the forewing venation and the shape of the uncal lobes. Psithyristria is found in the northern Luzon mountains and is closely related to Semia of Taiwan and eastern China, Pomponia of east Asia and the Indonesian/Malaysian islands, and probably most closely to Basa of India. Psithyristria may be a cicadid example of a small number of Philippine organisms that trace their ancestry directly back to the temperate Asian mainland, rather than through the island chains to the south of the country.     

Phylogeny of the tribe Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) based on morphological characteristics,with reclassification of the Empoasca generic group

A morphology‐based phylogenetic analysis of the tribe Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) is presented for 58 of 83 formerly recognized genera based on 99 morphological characters of adults. The results support excluding the New World Beamerana generic group from Empoascini. The remaining genera of Empoascini were recovered as a monophyletic sister group of Dikraneurini. Previously recognized tribes Jorumini and Helionini are derived from within Empoascini and are considered synonyms of the latter tribe. Three previously recognized informal generic groups, the Empoasca group, Alebroides group and Usharia group were paraphyletic but the Ficiana group was recovered as monophyletic based on five synapomorphies. Genera previously placed in the Alebroides group represent at least six independent lineages, indicating that the hind wing character separating this group from the Empoasca group (CuA and MP veins free) is highly homoplasious. Empoasca (sensu lato) is also paraphyletic. Thus, twelve previously recognized subgenera of Empoasca are elevated to genus status and five species groups of Empoasca from the New World are recognized as separate new genera. Sikkimasca Dworakowska, 1993 is treated as synonym of Marolda Dworakowska, 1977 based on the phylogeny. Biogeographic analysis suggests that Empoascini most likely first evolved in the Oriental region and spread to other biogeographic realms more recently by multiple independent invasions.     

Cladistic analysis of Coenosiini (Diptera: Muscidae: Coenosiinae)

The phylogenetic relationships among world genera of Coenosiini (Diptera: Muscidae: Coenosiinae) were investigated using parsimony. The analysis involved forty‐six ingroup terminal taxa, representing 100% of the genera currently assigned to this tribe, three outgroups and sixty‐seven adult male and female morphological characters. The monophyly of Coenosiini is confirmed by the position of the three katepisternal bristles, equidistant from each other and placed at the points of an equilateral triangle. Genera Andersonosia, Agenamyia, Anthocoenosia, Drepanocnemis, Pachyceramyia and Rhabdotoptera are removed from Coenosiini and temporarily placed in Limnophorini. The other genera fall into two groups: the Lispocephala‐group, comprised of genera with mainly Old World species and the Nearctic genus Pentacricia; and the Coenosia‐group, with the highest level of generic diversity in South America. Each group is defined by synapomorphies of its constituent genera: Lispocephala‐group by the presence of a posteroventral apical seta on the hind tibia, the presence of two arms in male sternite 6 (not forming a ring) and the short or very short female ovipositor; the Coenosia‐group by the presence of a developed epiproct and narrow sternites 6 and 7 of the female ovipositor. The following new generic synonymies are proposed (junior synonyms in parentheses): Lispocephala Pokorny (=Pectiniseta Stein), Coenosia Meigen (=Tenuicosta Stein; Dexiopsis Pokorny), Neodexiopsis Malloch (=Haroldopsis Albuquerque), Pilispina Albuquerque (=Levallonia Albuquerque; Noelia Albuquerque; Parvomusca Medeiros; Cholomyioides Albuquerque), Apsil Malloch (=Raymondomyia Malloch), Stomopogon Malloch (=Angolia Malloch; Angolina Pont) and Pygophora Schiner (=Chouicoenosia Cui & Xue).     

A systematic study of Phytalmia (Diptera, Tephritidae) with description of a new genus

The tribe Phytalmiini is defined to include the genera Phytalmia Gerstaecker, Diplochorda Osten Sacken (=Nesadrama Perkins syn.n.) and Sessilina gen.n. (type-species Dacus nigrilinea Walker), with Ortaloptera Edwards as an outlying member; a key is provided to these genera. The tribe is nearly restricted to the Papuan Subregion; the record of Nesadrama petiolata Hardy from the Philippine Islands is questioned, and this species is synonymized with Diplochorda myrmex Osten Sacken syn.n. Nesadrama longistigma Perkins is a synonym of D.turgida (Walker) syn.n. Keys are provided to the six species of Phytalmia: alcicornis (Saunders), antilocapra sp.n., biarmata Malloch, cervicornis Gerstaecker (=prisca Enderlein syn.n.), megalotis Gerstaecker (= wallacei Saunders syn.n.), mouldsi sp.n.; also the three species of Sessilina, horrida sp.n., literata sp.n., nigrilinea (Walker). The enlarged cheek-processes may be greatly reduced in P.biarmata and cervicornis. The morphology of the internal reproduction system of mouldsi females is described.     

The tribe Bryocorini (Insecta: Heteroptera: Miridae: Bryocorinae): phylogeny,description of a new genus,and adaptive radiation on ferns

A cladistic analysis of the tribe Bryocorini based on 68 morphological characters is conducted. Bryocorini are supported as a monophyletic group with Eccritotarsini as their sister taxon. Based on the phylogenetic analysis, we redefine the tribe Bryocorini to contain the following seven genera: Bryocorella Carvalho, 1956, Bryocoris Fallén, 1829, Bryophilocapsus Yasunaga, 2000, Cobalorrhynchus Reuter, 1906 gen. dist., D iplazicoris gen. nov. , Hekista Kirkaldy, 1902, and Monalocoris Dahlbom, 1851. The genus Bryocorella is transferred to Bryocorini from the tribe Eccritotarsini. The subgenus Cobalorrhynchus is treated as a separate genus. Diplazicoris is described as monotypic to accommodate D iplazicoris lombokianus sp. nov. An updated diagnosis of the tribe, a key to genera, and a diagnosis of each recognized genus are presented. Selected photomicrographs, scanning micrographs, and illustrations of the pretarsus, metepisternal scent efferent system, metafemoral trichobothria, and morphology of head, pronotum, and male and female genitalia are provided. Mapping of the host data on the revealed tree shows that Bryocorini represent one of the very few currently known examples of the adaptive radiation of a fairly large insect group on ferns. © 2015 The Linnean Society of London     

蔺叶蜂亚科一新属新种（膜翅目：叶蜂科）

本文记述蔺叶蜂亚科１新属新种“朱氏背齿叶蜂Ｎｏｔｏｄｏｎｉｄｅａ ｃｈｕｉ ｇｅｎ．ｓｐ．ｎｏｖ．新属隶属于等角叶蜂族Ｐｈｙｍａｔｏｃｅｒｉｎｉ,与窝眶叶蜂属Ｒｈａｄｉｎｏｃｅｒａｅａ Ｋｏｎｏｗ及扁室叶蜂属Ｃｏｒｐｉｌｕｓ Ｍａｌａｉｓｅ较近似;窝眶叶蜂属触角粗短且简单,第２节长等于宽,唇基截型,颚眼距狭于１／２单眼直径,后眶沟深并具陷窝,爪具微小内齿,前足胫节内距分叉等易于新属区别;扁室叶蜂属     

Blackmania gen. nov. and the related genus Acaudella (Hemiptera: Aphididae: Aphidinae)

The aphid genus Blackmania gen. nov. is described. In the genus, B. eastopi sp. nov. associated with Polygonum equisetiforme (Polygonaceae) from Israel and Cyprus is described and illustrated. Morphologically, the new genus is similar to the genus Acaudella in respect to the lack of a cauda. Acaudella puchovi, associated with Atraphaxis caucasica (=A. buxifolia) and A. spinosa (Polygonaceae) from Uzbekistan and Israel, is re‐described and the apterous viviparous female is figured for the first time. The lectotype and paralectotypes of A. puchovi are also designated. An identification key to known species of the tribe Macrosiphini without cauda is provided. The morphological separation of B. eastopi gen. nov., sp. nov. from A. puchovi is visualized using principal component analysis.     

New gall midges of the tribe Leptosynini (Diptera,Cecidomyiidae) from the Late Eocene ambers and the classification of the supertribe Heteropezidi

Three new genera and 27 new species of gall midges are described from the Late Eocene ambers: Henria baltica sp. nov., Frirenia manca sp. nov., F. musicata sp. nov., Leptosyna samlandica sp. nov., L. fastosa sp. nov. from Baltic amber and H. xystica sp. nov., H. liquida sp. nov., Stellasegna vlaskini gen. et sp. nov., S. vaporea sp. nov., S. nexa sp. nov., Rasnitsia verticosa gen. et sp. nov., F. rohdendorfi sp. nov., F. schevchenkoi sp. nov., F. melica sp. nov., F. lukashevichae sp. nov., F. leporidis sp. nov., F. marmarygma sp. nov., F. vesana sp. nov., Vincinescia alisae gen. et sp. nov., L. margarita sp. nov., L. munifera sp. nov., L. sukachevae sp. nov., L. assa sp. nov., L. larga sp. nov., L. vegeta sp. nov., L. vaticina sp. nov., and L. shcherbakovi sp. nov. from Rovno amber. Strobliella capitata Fedotova is redescribed as Henria capitata (Fedotova, 2004) (comb. nov.). Diagnoses of Henria (= Electroxylomyia Nel et Prokop, syn. nov.), Frirenia, and Leptosyna are revised. As a result, Henria comprises 3 extant and 5 extinct (Late Eocene) species, including H. eocenica (Nel et Prokop), comb. nov. (= Electroxylomyia eocenica), Frirenia comprises 1 extant and 10 Late Eocene species, and Leptosyna comprises 3 extant and 11 Late Eocene species. The tribe Heteropezini is elevated to the supertribal rank (Heteropezidi) and included in the subfamily Lasiopterinae. Leptosynini is treated as a separate tribe, and Lasiopterinae is considered as part of Cecidomyiidae s. str. (i.e., excluding Lestremiidae). Keys to the tribes and genera of Heteropezidi and to species of Henria, Stellasegna, Frirenia, and Leptosyna are provided. The gall midge faunas of the Rovno and Baltic ambers are compared. Phylogenetic relationships within the supertribe are hypothesized.     

Systematic revision of Aluntiini Emeljanov, 1979 (Hemiptera: Fulgoromorpha: Dictyopharidae: Dictyopharinae): reclassification,phylogenetic analysis,and biogeography

The dictyopharid planthopper tribe Aluntiini s.l. is revised and reclassified into two tribes: Aluntiini s.s. and Arjunini Song & Szwedo trib. nov. The tribe Aluntiini s.s. includes five genera: Aluntia Stål, 1866; D endrophora Melichar, 1903 stat. rev. ; Dictyomorpha Melichar, 1912; Indodictyophara Liang & Song, 2012; and Madagascaritia Song & Liang gen. nov. The new tribe Arjunini comprises two genera – Arjuna Muir, 1934 and Pippax Emeljanov, 2008 – both moved from Aluntiini s.l. Four new species – A luntia longicephalica Song & Szwedo sp. nov. , Madagascaritia angusta Song & Liang sp. nov. , Arjuna maai Song & Wang sp. nov. , and Arjuna muiri Song & Wang sp. nov. – are described. A morphologically based phylogenetic analysis is undertaken for Aluntiini, Arjunini, and the representatives of Dictyopharini, Hastini, Orthopagini, and the fossil Worskaitini within Dictyopharinae, all distributed in the Old World. A matrix of 129 characters of the habitus, coloration, head, thorax, and male and female genitalia of the adults was used for the cladistic analysis. The phylogenetic results show that Aluntiini s.l. as placed in Dictyopharidae is well supported, but it is distinctly paraphyletic and should be separated into two unambiguous tribes. A palaeotropical distribution pattern displayed by Aluntiini is suggested. The origin and diversification of Aluntiini are discussed preliminarily. © 2015 The Linnean Society of London     

Out of Africa? A dated molecular phylogeny of the cicada tribe Platypleurini Schmidt (Hemiptera: Cicadidae), with a focus on African genera and the genus Platypleura Amyot & Audinet‐Serville

The Platypleurini is a large group of charismatic cicadas distributed from Cape Agulhas in South Africa, through tropical Africa, Madagascar, India and eastern Asia to Japan, with generic diversity concentrated in equatorial and southern Africa. This distribution suggests the possibility of a Gondwanan origin and dispersal to eastern Asia from Africa or India. We used a four‐gene (three mitochondrial) molecular dataset, fossil calibrations and molecular clock information to explore the phylogenetic relationships of the platypleurine cicadas and the timing and geography of their diversification. The earliest splits in the tribe were found to separate forest genera in Madagascar and equatorial Africa from the main radiation, and all of the Asian/Indian species sampled formed a younger clade nested well within the African taxa. The tribe appears to have diversified during the Cenozoic, beginning c. 50–32 Ma, with most extant African lineages originating in the Miocene or later, well after the breakup of the Gondwanan landmass. Biogeographical analysis suggests an African origin for the tribe and a single dispersal event founding the Asian platypleurines, although additional taxon sampling and genetic data will be needed to confirm this pattern because key nodes in the tree are still weakly supported. Two Platypleurini genera from Madagascar (Pycna Amyot & Audinet‐Serville, Yanga Distant) are found to have originated by late Miocene dispersal of a single lineage from Africa. The genus Platypleura is recovered as polyphyletic, with Platypleura signifera Walker from South Africa and many Asian/Indian species apparently requiring assignment to different genera, and a new Platypleura concept is proposed with the synonymization of Azanicada Villet syn.n. The genera Orapa Distant and Hamza Distant, currently listed within separate tribes but suspected of platypleurine affinity, are nested deeply within the Platypleurini radiation. The tribe Orapini syn.n . is here synonymized while the tribe Hamzini is pending a decision of the ICZN to preserve nomenclatorial stability.