Revision and phylogeny of Syrphetodes (Coleoptera: Ulodidae): implications for biogeography,alpinization and conservation

The genus Syrphetodes Broun is revised to include a total of 13 species. Most of the species are restricted in their distributions, are rarely collected and have been attributed conservation status in New Zealand. Eleven species are described as new: three from Northland (S. relictus sp.n ., Te Paki; S. insularis sp.n. , Three Kings Islands; S. magnus sp.n. , Hokianga), one from the central North Island (S. obtusus sp.n. ), one from Central Otago (S. nunni sp.n. , Waikaia Bush), and seven from the southern Alps (S. cirrhopogon sp.n. , Aspiring National Park; S. occiduus sp.n. , Westland; S. melanopogon sp.n. , Mt Dewar, Paparoa Range; S. defectus sp.n. , northern Paparoa Range; S. marrisi sp.n. , Mt Domett, Northwest Nelson; S. carinatus sp.n. , Victoria Range). Eleven synonymies are proposed: S. crenatus Broun (= S. dorsalis Broun, syn.n .), S. marginatus Pascoe (= S. bullatus Sharp, syn.n. ; S. sylvius Broun, syn.n. ; S. cordipennis Broun, syn.n. ; S. punctatus Broun, syn.n. ; S. simplex Broun, syn.n. ; S. nodosalis Broun, syn.n. ; S. truncatus Broun, syn.n. ; S. variegatus Broun, syn.n. ; S. pensus Broun, syn.n. ; S. thoracicus Broun, syn.n. ). The phylogenetic relationships among the species were reconstructed using morphological (25 adult characters) and DNA sequence (nuclear 28S rDNA and mitochondrial cytochrome c oxidase subunit I) data. A morphological analysis rooted with Trachyderastes resulted in a split between lowland and high‐altitude species and a well‐supported group from Northland. Molecular trees rooted with representatives of Trachyderastes Kaszab (New Caledonia), Meryx Latrielle (Australia), Ulodes Erichson (Australia) and three New Zealand genera (Arthopus Sharp, Brouniphylax Strand, Exohadrus Broun) resulted in the following tree: ((Ulodes, Brouniphylax) (Exohadrus, Arthopus)) (Syrphetodes (Meryx, Trachyderastes)). Species relationships within Syrphetodes included a strongly supported northern North Island clade and an alpine clade either as sister taxon to S. crenatus and S. marginatus or sister remaining lowland lineages. Combined phylogenetic analyses also showed paritial congruence with separate partitions. The distributions of the lowland species, in particular those from the North Island, correspond to islands that existed in the Pliocene. The alpine, black‐coloured lineage, found above the treeline, is monophyletic based on several characters (e.g. lack of abdominal flanges and reduced scalation) and, in some reconstructions, the tan‐coloured S. cirrhopogon is sister taxon to the remaining black‐coloured species. This published work has been registered in ZooBank, http://zoobank.org/urn:lsid:zoobank.org:pub:697E68E8‐EE90‐46C1‐A009‐78A794E0EF4F .     

Gondwanan Gymnochilini (Coleoptera: Trogossitidae): generic concepts,review of New Zealand species and long‐range Pacific dispersal

Members of Leperina Erichson (Trogossitidae: Gymnochilini) from New Zealand, New Caledonia and Lord Howe Island are morphologically similar to members of the endemic Juan Fernandez Island genus Phanodesta Reitter, sharing at least one obvious character, elytral carinae that are beaded and contain well‐defined punctures. To test the monophyly of Leperina and Phanodesta, we reconstructed phylogenetic relationships of the genera of the tribe Gymnochilini by a cladistic analysis of 22 terminals and 47 adult characters rooted with one genus of trogossitine. Leperina is rendered paraphyletic by the placements of Seidlitzella Jakobson and Phanodesta. Kolibacia n.gen. (type species Leperina tibialis Reitter) is described for east Palaearctic species included formerly in Leperina (two new combinations); New Zealand Leperina and other species from New Caledonia and Lord Howe Island (Ostoma pudicum Olliff) are transferred to Phanodesta (six new combinations); and the remaining species are retained in Leperina. The following species are described as new: Phanodesta carinata n.sp., P. manawatawhi n.sp., P. oculata n.sp. and P. tepaki n.sp. Leperina ambiguum Broun is transferred to Grynoma Sharp resulting in a new combination and three new synonymies for New Zealand trogossitines: Leperina interrupta Brookes n.syn. and Leperina sobrina (White) n.syn. [= Phanodesta farinosa (Sharp)], and Trogosita affinis White n.syn. (= Tenebroides mauritanicus Linnaeus). A key to the New Zealand species and a checklist for the species of the Kolibacia, Leperina and Phanodesta are provided. The derived placement of Juan Fernandez Phanodesta in the phylogeny is evidence for long‐distance dispersal from Australasia. A tally of all Juan Fernandez Islands Coleoptera shows derivation mostly from Chile and South America, with few from the southern Pacific region, rarely from Australasia.     

Pselaphotumulus,a new genus of pselaphine endemic to New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Pselaphitae)

ABSTRACT

Pselaphotumulus Owens and Carlton, gen. nov., is described as the sixth genus in the tribe Pselaphini known from New Zealand. Three new species are described: Pselaphotumulus aorerei, sp. nov., Pselaphotumulus dubius, sp. nov. and Pselaphotumulus unus, sp. nov.. Three species, Pselaphus cavelli (Broun 1893), Pselaphus oviceps (Broun 1917) and Pselaphus urquharti (Broun 1917) are transferred to Pselaphotumulus, nov. combs. Lectotypes from type series in the New Zealand Broun Collection (Natural History Museum, London) are designated for these three species. Habitus photographs, distributional maps and line drawings of diagnostic characters are provided for each species. A key to species is provided. Searches of museum collections have not yielded representatives outside of New Zealand, suggesting that this is the first endemic genus in the tribe Pselaphini described from the New Zealand’s main islands, specifically, the South Island. Pselaphotumulus species exhibit restricted distribution patterns that approximately coincide with the Pacific/Indo Australian Plate boundary.     

A synopsis of the New Zealand Anillini (Coleoptera: Carabidae: Bembidiinae), with descriptions of new genera and species

Abstract

Pelodiaetodes n.gen. (type-species P. prominens n.sp.) and Hygranillus n.gen. (type-species H. kuscheli n.sp.) are described, and the following new synonymies are established: Nesamblyops oreobius (Broun, 1893) = N. coriaceus (Broun, 1908); Zeanillus pallidus (Broun, 1884) = Z. marginatus (Broun, 1914). Nine species of New Zealand Anillini are now recognised; they are distributed between five genera.     

The family and subfamily classification and New Zealand genera of Pythidae and Scraptiidae (Coleoptera)

ABSTRACT. The family Pythidae is defined on adult and larval characters to include the subfamilies Pythinae and PllipalPlnae. Trachelostenidae stat.n. is excluded. Relationships of Pythidae, Boridae, Trictenotomidae and SalPlngidae (s.l.) are analysed. Strong resemblances between some PllipalPlnae and Pyrochroidae appear to have arisen through convergent evolution. Adults and larvae of the New Zealand genera Techmessa, Techmessodes and Exocalopus are described. In Scraptiidae, adults of the New Zealand genera Nothotelus and Phytilea , and the larva of Nothotelus , are described. Type data are given for all described New Zealand species of these families. The following new synonymies are established: PllipalPlnae (Abdullah, 1964)=Techmessinae Paulus, 1971. Exocalopus pectinatus Broun, 1893= nitidiceps Broun, 1910. Techmessa concolor Bates, 1874= attenuata Broun, 1893= rugicollis Broun, 1910= unicolor Paulus, 1971. Techmessa telephoroides Bates, 1874= varians Broun, 1893. Techmessodes Plcticornis (Broun, 1880)= distans (Sharp, 1882). Techmessodes versicolor Broun, 1893= cephalotes Broun, 1910. The following genera are reassigned to the families indicated: Ischyomius to Trictenotomidae (from Pythidae or Melandryidae). Phytilea to Scraptiidae (from Oedemeridae or Anthicidae). Pseudananca to Aderidae (from Oedemeridae). Scraptogetus (= Metasclera ) to Aderidae (from Scraptiidae, Oedemeridae or Anthicidae).     

Siteroptes (Siteroptoides) species with Pediculaster-like phoretomorphs (Acari: Tarsonemida: Pygmephoridae) from New Zealand and Polynesia

Abstract

Six species of Siteroptes subgenus Siteroptoides—kneeboni (Wicht), mesembrinae (Canestrini), microsaniae n.sp., morelliae (Rack), muscarius n.sp., and portatus n.sp.—have been found in New Zealand. S. (S.) mesembrinae occurs on Niue Island also, and a further new species, pacificus, is recorded from Tonga. The phoretomorph females of all 7 species, the normal females of 4, the males of 3, and larvae of 2 are described and compared. The similarities and differences between normal and phoretomorph females are particularly noted. Besides kneeboni, 2 species described by Wicht but not occuring in New Zealand—athiasae and flechtmanni— are redescribed from type material and transferred to Siteroptes (Siteroptoides). Keys are given for normal females and Pediculaster-like phoretomorph females of Siteroptoides. Siteroptes mesembrinae subsimilis and S. ignotus altaicus are raised to species status. Diagnostically important measurements are given to help define the variation within and between species and morphs.     

Five new species of Thripidae (Thysanoptera) endemic to New Zealand

Abstract

Three new species of genus Thrips are described—austellus, coprosmae, and phormiicola—which are closely related to the common New Zealand species T. obscuratus (Crawford). The systematic and zoogeographic significance of this endemic species-group is discussed. Physemothrips hadrus n.sp. is described from the South Island; the only other member of this genus is from Macquarie Island. Anaphothrips zelandicus n.sp. is described from the South Island and the Chatham and Antipodes islands, A. woodi Pitkin is recorded from New Zealand, and the relationships of these two species to the Australian fauna are discussed.     

Anobiidae (Coleoptera) of the Kermadec Islands

Abstract

Lasioderma serricorne (Fabricius), Leanobium marmoratum (Lea), Leanobium flavomaculatum n.sp., Serianotus punctilatera kermadecensis n.ssp., and Dorcatomiella ornata Español are recorded from the Kermadec Islands. This fauna is relatively poor, probably because of the small size and limited diversity of the islands. Affinities are with New Zealand (L. flavomaculatum), Norfolk Island (L. marmoratum), and Lord Howe Island (S. punctilatera kermadecensis). D. ornata is strongly endemic, though other species of Dorcatomiella occur on the Marianas and Tahiti.     

New genera and species of Leptophlebiidae (Ephemeroptera) from New Zealand

Abstract

New genera Isothraulus, Arachnocolus, and Penniketellus are established for three species of leptophlebiid mayfly from New Zealand. Each genus is monotypic and endemic to New Zealand. Isothraulus and Arachnocolus are known only from the northern North Island, and Penniketellus is known only from the Arthur's Pass area of the central South Island. The male and female imago, nymph, and egg of Isothraulus abditus n.sp., the male imago, male subimago, and nymph of Arachnocolus phillipsi n.sp., and the male and female imago, female subimago, and egg of Penniketellus insolitus n.sp. are described. The relationships of each genus and the ecology of nymphs of each species are discussed.     

Two new genera and twelve new species of Anthuridea (Crustacea: Isopoda) from off the West Coast of New Zealand

Abstract

From the shelf and upper slope off the West Coast of the South Island of New Zealand, the following new taxa of Isopoda Anthuridea (Crustacea) are described: Kupellonura proberti n. sp. (Hyssuridae); Quantanthura pacifica n. sp., Haliophasma platytelson n. sp. and H. novaezelandiae n. sp. (Anthuridae) Albanthura stenodactyla n. gen. et sp., A. rotunduropus n. sp., Bullowanthura crebrui n. sp., Leptanthura tmncatitelson n. sp., L. exilis n. sp., L. profundicola n. sp., Paranthura longa n. sp., and Psittanthura egregia n. gen. et sp. (Paranthuridae). Most of these are blind, unpigmented species, adapted to life in deep water. Albanthura n. gen. is blind, has a single telsonic statocyst, short flagella of the antennae, a maxilliped with 3 palpal articles, and a triangular carpus on pereopods 4 to 7. Psittanthura n. gen. is also blind, with very unusual, enlarged subchelae of pereopods 1 and 2, a large, rectangular telson with a single statocyst; the maxillipedal palp is composed of 1 large and 1 small, distal article; a mandibular palp is absent.     

Two new species of Colobopyga (Homoptera: Halimococcidae) from the Australian region

Abstract

Two new species of Colobopyga Brèthes are described and illustrated from adult and second-instar females: C. hedyscepes n.sp. from Auckland, New Zealand, on Hedyscepe canterburyana (a palm native to Lord Howe Island); and C. australiensis n.sp. from Australia, on “Kentia palm”. A key is given to the 10 known species of Colobopyga.     

Descriptions of new taxa of the Clausiliidae from Turkey (Mollusca: Stylommatophora)

Summary

Several new taxa of the Clausiliidae were described From Turkey: Dobatia multidentifera n. sp., Albinaria monocristata n. sp., Albinaria basalifera n. sp., Roseniella (Chavchetia) n. subgen., Roseniella (Chavchetia) rufina n. sp., Armenica euprepes truncata n. ssp., Armenica multispirata n. sp., Armenica bicarinata grisea n. ssp., Kazancia n. gen., and Kazancia monticola n. sp. Investigations were made of the genital morphology of some species, and the arrangement of the taxa in Nordsieck’s classification (H, NORDSIECK 1979) is discussed.     

Typhlocybinae of New Zealand (Homoptera: Cicadellidae)

Two tribes of the subfamily Typhlocybinae are represented in New Zealand, as follows. Erythroneurini: the cosmopolitan genus Zygina is represented by 4 endemic species—toetoe, ramsayi n.sp., dumbletoni, and agni n.sp.—plus zealandica, which occurs in Australia also. Typhlocybini: Typhlocyba froggatti, T. lethierryi, Ribautiana tenerrima, Eupteryx melissae, Kybos smaragdula, and K. betulicola are introduced Northern Hemisphere elements; Matatua montivaga n.gen.&sp. is endemic; M. maorica (formerly Dikraneura maorica), also endemic, is of uncertain identity in the absence of male specimens. Keys are given for the separation of all taxa, the genera and species are described, and all species except Matatua maorica are figured.     

Pseudoexeirarthra,a new genus from New Zealand (Coleoptera,Staphylinidae, Pselaphinae), with descriptions of seven new species

A new endemic genus and seven new species of New Zealand pselaphine staphylinid beetles of the supertribe Faronitae are described as follows: Pseudoexeirarthra Park & Carlton, gen. n. (type species: Sagola spinifer Broun); Pseudoexeirarthra sungmini Park & Carlton, sp. n.; Pseudoexeirarthra kwangguki Park & Carlton, sp. n.; Pseudoexeirarthra youngboki Park & Carlton, sp. n.; Pseudoexeirarthra seiwoongi Park & Carlton, sp. n.; Pseudoexeirarthra parkeri Park & Carlton, sp. n.; Pseudoexeirarthra hlavaci Park & Carlton, sp. n.; Pseudoexeirarthra nomurai Park & Carlton, sp. n. Three species, Sagola spinifer Broun, Sagola colorata Broun, and Sagola puncticollis Broun, are transferred to the genus Pseudoexeirarthra. Six species are synonymized: Sagola dilucida Broun, Sagola guinnessi Broun, Sagola longicollis Broun, Sagola longula Broun, and Sagola rectipennis Broun under Pseudoexeirarthra spinifer (Broun); Sagola insueta Broun under Sagola colorata (Broun). A lectotype is designated for Pseudoexeirarthra spinifer (Broun). A key, habitus photographs, line drawings of diagnostic characters, and distribution maps are provided for each species.     

New species,new genera,and new records in Cerambycidae from Peru (Coleoptera)

Three new species and two new genera of longhorn beetles are described from Peru: Cosmoplatus polis n. sp. (Pteroplatini), Hemilocallia grishami n. gen., n. sp. (Calliini) and Arabela ramirezi n. gen., n. sp. (Hemilophini). Esmeralda laetifica Bates, 1869 (Prioninae), Stratone transversalis (Chevrolat, 1862) (Cerambycinae), Gymnocerina cratosomoides (Bates, 1862) (Lamiinae), and Trachysomus verrucosus (Olivier, 1795) (Lamiinae) are new records for Peru.     

A Contribution to the Taxonomy of the Genus Xiphinema Cobb, 1913

In a comparative study of different populations of Xiphinema, the significance of the female gonad structure in species determination is stressed, and monodelphic, pseudomonodelphic, and didelphic forms in the genus are defined. X. chambersi is redescribed and illustrated, X. ensiculiferum and X. krugi are redefined, and X. ensiculiferoides n.sp. and X. orthotenum n.sp. are described. Proposed new synonymies are: X. itanhaense = X. brasiliense; X. bulgariensis and X. conurum = X. italiae; X. ifacolum = X. basiri; X. macrostylum = X. ensiculiferum; X. truncatum = X. elongatum; and X. vulgare = X. setariae. X. yapoense is considered species inquirenda and X. obtusum a nomen dubium. The following eight subgenera of the genus Xiphinema are proposed: Radiphinema n. subg., Krugiphinema n. subg., Xiphinema n. subg., Elongiphinema n. subg., Halliphinema n. subg., Basiphinema n. subg., Rotundiphinema n. subg., and Diversiphinema n. subg. A key to the subgenera of Xiphinema is presented, plus a list of 50 species in the genus, their synonyms, and 10 species inquirendae.     

Taxonomy, host specificity and dietary implications of Hurleytrematoides (Digenea: Monorchiidae) from chaetodontid fishes on the Great Barrier Reef

Five new and five previously described species of Hurleytrematoides are reported from 19 of 34 chaetodontid species examined from the Great Barrier Reef; new species are H. faliexae n. sp., H. galzini n. sp., H. loi n. sp., H. morandi n. sp., and H. sasali n. sp. Previously described species are H. coronatum, H. fijiensis, H. prevoti, H. bartolii, and H. zebrasomae. The genus is rediagnosed in the light of morphological variation of the new species; the degree of spination and shape of the terminal genitalia distinguish individual species. Species of Hurleytrematoides infect almost every clade of the family Chaetodontidae found on the Great Barrier Reef, but obligate corallivores are not infected. All ten species were found at Heron Island on the southern Great Barrier Reef, but only six at Lizard Island on the northern Great Barrier Reef. For three of the four species not present at Lizard Island, the absence appears to be statistically significant. Although all species are apparently restricted to chaetodontids on the GBR, specificity within the family varies from oioxenous to euryxenous; a core/satellite host paradigm explains the distribution of several species.