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在研究新发现化石材料及对已有标本重新观察的基础上,讨论中国晚中新世至早上新世原大羚(?Protoryx)的系统分类位置。依据头骨、角心和牙齿等方面所具有的独特性状,建议把发现于中国北方曾归入?Protoryx的种类从旧大陆同时代或时代稍早的Protoryx Major, 1891中分出,另立2个新属:以?Protoryx yushensis Teilhard&Trassaert,1938为属型种的华北羚(Huabeitragus gen.nov.)和以?Protoryx shansiensis Bohlin,1935为属型种的粗壮羚(Macro- tragus gen.nov.)。华北羚体型中等大小,头骨窄长,颅轴与面轴夹角近直角,弯曲发生在角前的额部,眶上孔小,分开远,脑颅部短窄,间顶骨长方形,基枕骨方形,没有中纵沟或中嵴,角心长而纤细,基部靠近,向上分散度大,基部横断面近三角形,具前棱。Macrotragus头骨大而粗壮,其弯曲发生在角心基部之间的额面,弯曲度大于直角;角基之间的额中缝隆起呈脊;脸部宽;眶前窝浅;眶下孔后缘位于P3之上。眼眶向头骨两侧突出不明显,其前缘位于M3后缘之后;脑颅部短宽,呈简状,背面稍凸;枕面高、呈半圆形;基枕部为长方形或梯形,具中纵沟和弱的中纵脊;卵圆孔大,面向侧方;角心大而粗壮,内外侧扁,无棱,基部紧靠,向上的分散度小,基部横切面为椭圆形。无角后窝。前臼齿列退化。这两个属之间的主要不同在于前者的个体较小;角心较细弱,向上分散度大(前面观),具前棱,基部横切面呈次三角形;头骨窄,其弯曲位置靠前,弯曲接近90°;脑颅部窄;枕面平,为低矮的长方形;基枕部呈方形,面平,无中纵沟和中纵脊,前、后节结不发育和卵圆孔相对小等。在系统位置上,它们可能属于山羊亚科(Caprinae)。  相似文献   

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NEW MATERIAL OF HELICOPRIONID SHARK FROM LIANYUAN OF HUNAN   总被引:1,自引:0,他引:1  
Edestiforme8obruchev,1953HelicoprionidaeKarpinsky,l9llHunanohelicoprionxiandongensi8gen.etsp.nov.Ho1otypeApartoftooth-row,withtenteethfragments,no.XKG-F9lo1,Xi-angtanMininglnstitute.LocalityandhorizonXiandong,Lianyuan,Hunan;QixiaFormation,theLowerPermian.DisgnosisTeethofcuttingtype,lateral1ycompressed;cuttingb1adeserratedanteriorlyandposteriorly,serrationtiny;lateralwingrelativelywide,serrationontheanteriormarginirreguIarandnoserrationontheposteriormargin,"intermediateregionwithoutenam…  相似文献   

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Even after 200 years of study, some details of the cranial anatomy of ichthyosaurs, one of the most successful groups of marine vertebrates in the Mesozoic, are still unclear. New information on the braincase, palate and occiput are provided from three‐dimensional scans of an exceptionally preserved ichthyosaur (‘Hauffiopteryxtypicus) skull from the Toarcian (183–174 Ma, Lower Jurassic) of Strawberry Bank, England. This ichthyosaur has unusual, hollow, tubular hyoid bars. The occipital and braincase region is fully reconstructed, creating the first digital cranial endocast of an ichthyosaur. Enlarged optic lobes and an enlarged cerebellum suggest neuroanatomical adaptations that allowed it to be a highly mobile, visual predator. The olfactory region also appears to be enlarged, suggesting that olfaction was more important for ichthyosaurs than has been assumed. Phylogenetic analysis suggests this ichthyosaur is closely related to, but distinct from, Hauffiopteryx, and positioned within Thunnosauria, a more derived position than previously recovered. These results further our knowledge of ichthyosaur cranial anatomy in three dimensions and provide a platform in which to study the anatomical adaptations that allowed ichthyosaurs to dominate the marine realm during the Mesozoic.  相似文献   

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Abstract:  A new fauna of Lower Cretaceous (Albian) ichthyosaurs, which includes at least one new genus and species, was recovered from deposits of the Loon River Formation at Hay River, Northwest Territories, Canada. All Cretaceous ichthyosaurs have been referred to a single genus, Platypterygius . The Loon River Formation material, however, does not satisfy the diagnosis for Platypterygius , and it is distinctive enough to warrant the erection of a new genus and species of ichthyosaur. Maiaspondylus lindoei gen. et sp. nov. is distinguished from other ichthyosaurs by an extensive overlapping contact of the jugal and the maxilla; marginal teeth with smooth crowns, fluted cementum, lingual curvature and implantation in a dental groove; and humerus with isomorphic proximal and distal ends, featuring three distal articular facets in which the medial articular facet is smaller than the lateral facets. The holotype and referred material is described here, and the relationship of Maiaspondylus to other ophthalmosaurs is discussed.  相似文献   

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Abstract:  A new genus and species of an elasmosaurid plesiosaur, Futabasaurus suzukii , is described based on a partial skeleton from the Inoceramus amakusensis Zone (Lower Santonian, Upper Cretaceous) of the Irimazawa Member of the Tamayama Formation, Futaba Group, in Fukushima Prefecture, Japan. The new taxon is characterized by a number of characters such as the wide space between the orbit and external naris, posterior extension of the interclavicle, relatively long humerus and prominent femoral muscle scar. The holotype includes a partial skull and mandible, posterior cervicals to sacrals, ribs, clavicular arch, pelvic girdle and four limbs. The remains are mostly in articulation, and exhibit evidence of predation/scavenging by sharks. The distribution of elasmosaurid species in the circum-Pacific region remains unclear due to the lack of diagnostic materials. The occurrence of F. suzukii is geographically and stratigraphically significant, because it allows species-level comparison; as a diagnosable elasmosaurid specimen, F. suzukii is the first and the oldest from the northern Pacific.  相似文献   

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Abstract:  A new species of the genus Epiphaxum (family Lithotelestidae) is described and illustrated in detail, and compared to other species. Epiphaxum arbuscula sp. nov. has been collected from Upper Eocene (Priabonian), Upper Oligocene (Chattian) and Lower Miocene (Upper Burdigalian) deposits of the Aquitaine Basin, south-west France. Epiphaxum is a poorly documented genus but its fossil record extends back to the Late Cretaceous; it was previously known only from the Paleocene (Danian). Epiphaxum arbuscula differs from all others species of the genus in the form of its colony. In contrast to the creeping colonies of previously known species, it has branched colonies. It is very common at one Upper Oligocene outcrop from which an assemblage with submarine cave remains has been described. A close relationship between the three extant species (two from the Caribbean Sea and one from the Indo-West Pacific region) and the Paleogene species is also noted. These constitute a group that has not undergone any important morphological changes for the last 65 million years.  相似文献   

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Abstract:  Small reptiles from the Early Jurassic Pant 4 fissure fill in Glamorgan, South Wales (St. Bride's Island, Pant Quarry), were formerly provisionally attributed to three species of sphenodontian lepidosaurs. A re-analysis, aided by new material, has found this herpeto-fauna to consist almost exclusively of a single new species, Clevosaurus convallis sp. nov . , with only one specimen referable to Sphenodontia incertae sedis . Clevosaurus is known from the Upper Triassic and Lower Jurassic in various parts of the world, but C. convallis represents the first occurrence of the genus in the Jurassic of Britain. The material is fragmentary but includes numerous premaxillae, maxillae, dentaries and palatines, and the new species is distinguished by the unique combination of six large additional dentary teeth and a very short nasal process of the premaxilla, along with the diagnostic Clevosaurus features.  相似文献   

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Abstract:  The two plesiosauroid species from the Posidonia shale of Holzmaden, ' Plesiosaurus ' guilelmiimperatoris and ' Plesiosaurus ' brachypterygius , do not belong to Plesiosaurus but form new monotypic genera. The new genus Hydrorion is erected for ' P .' brachypterygius , and the genus Seeleyosaurus is re-established for ' P .' guilelmiimperatoris . The recently described species Plesiopterys wildii is regarded as a junior synonym of S. guilelmiimperatoris . A short phylogenetic analysis shows that S. guilelmiimperatoris and Muraenosaurus are basal elasmosaurs. H. brachypterygius , Occitanosaurus tournemirensis from France and Microcleidus homalospondylus from England form a monophyletic clade, which is the sister taxon to the Cretaceous elasmosaurs. A palaeobiogeographical comparison of plesiosaur localities in the Lower Jurassic shows distinct palaeobiogeographical zones for the Toarcian, with different plesiosaur taxa in England, Germany and France.  相似文献   

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New palaeothentid marsupial from the Middle Miocene of Bolivia   总被引:1,自引:0,他引:1  
The new species Acdestis maddeni is described based on the most complete skull of a palaeothentid marsupial known so far. The skull preserves most of the upper dentition and of the ear region, the latter an anatomical region previously unknown in this extinct family. A right maxillary fragment including C–M2 also belongs to the new species. The specimens come from the middle Miocene localities of Rosario and Quebrada Honda in southernmost Bolivia. The generic allocation of the new species is somewhat problematic, given that most of the taxonomy of fossil palaeothentids is based on lower dentitions. Compared to that of caenolestids, the skull of A. maddeni is much larger, with a shorter and broader snout, a proportionally narrower interorbital constriction, and a less globular, more triangular–shaped braincase in dorsal view.  相似文献   

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The oldest freshwater neritiliid, Neritilia bisinuata , is described from the Middle Eocene of the Loire Basin. Another European species, N. neritinoides , ranging from the Lower Oligocene to Lower Miocene (Upper Burdigalian) is recognized; its habitat appears to have been freshwater, but very close to the sea. Two new marine neritiliid species from the Aquitaine Basin are described: Bourdieria favia sp. nov. from the Upper Oligocene and Pisulinella aucoini  sp. nov. from the Lower Miocene. A third undescribed species from the Lower Miocene is referred to the same family and related to Pisulinella . The Oligocene species has a strong spiral sculpture, a character completely absent in previously known neritiliid species. The genus Agapilia , founded on juvenile N. neritinoides and adult Vitta picta , appears to be a junior synonym of the genus Vitta. The associated occurrence of shells of the families Neritiliidae, Neritopsidae and Pickworthiidae (well-known inhabitants of Indo-West Pacific submarine caves) at Peyrère suggest the first occurrence of a characteristic assemblage of dark submarine caves during the Oligocene. Both factorial analysis and relative abundance show that at Peyrère these families are associated with other cryptic fossils (various gastropods, bivalves, Brachiopoda, corals, Annelida). However, there are indications of other submarine cave assemblages in various Cenozoic deposits from the Palaeocene to the middle Miocene.  © 2004 The Linnean Society of London, Zoological Journal of the Linnean Society , 2004, 140 , 447–467.  相似文献   

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Abstract:  Phragmoteuthida from the Lower Jurassic (Upper Sinemurian) of Dorset, England, are fully described for the first time. Two species are recognized and described: Phragmoteuthis montefiorei (J. Buckman) and P. huxleyi sp. nov. Phragmocones have an apical angle of between 20 and 30 degrees, and relatively few chambers compared with belemnoids. The conotheca is multi-layered. The siphuncle is thick-walled. The pro-ostracum is long and three-lobed as in the Triassic species. Arms are short and bear pairs of slightly curved hooks. Shell and siphuncle structure do not indicate a close relationship with other coleoids.  相似文献   

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Abstract:  From 1935 to 1962 Maslov published two monographs and several other papers on the taxonomy of fossil calcareous algae of diverse ages from the large geographical area of the former USSR. Among many other taxa, he described five new genera ( Solenophyllum , Palaeophyllum , Mesolithon , Bicorium and Tomilithon , as a subgenus of Parachaetetes Deninger) which he attributed to the Corallinaceae, and the new genus Karpathia , which he included in the Squamariaceae. Type material of these taxa, except for Mesolithon , is housed in the Geological Institute of the Russian Academy of Sciences, Moscow. Re-examination of the genus types from a modern perspective of coralline algal taxonomy shows that Karpathia is a validly published genus of corallinaceans (subfamily Mastophoroideae) to which several Cenozoic fossil species can be assigned. In contrast, the holotype of P. elegans , the type species of Palaeophyllum , cannot be assigned to any subfamily within the Corallinaceae and its preserved features are inadequate for confidently delimiting a separate genus. Solenophyllum is a valid genus in which Palaeozoic representatives of Corallinales formerly attributed to Parachaetetes Deninger can be included. Tomilithon is considered a younger heterotypic synonym of Solenophyllum . The algal nature of the bioclasts in the type of Bicorium is uncertain.  相似文献   

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陕西蓝田灞河组陕西转角羚一新种(英文)   总被引:4,自引:4,他引:0  
1997~ 2 0 0 1年 ,中科院古脊椎动物与古人类研究所与芬兰赫尔辛基大学合作 ,于灞河组地层中发现哺乳动物化石计 45种。其中牛科化石种类与蓬蒂期典型三趾马动物群中的种类不同 ,至少存在分别归属于Shaanxispira、Protoryx、Dorcadoryx和Gazella的 4个新的种类。本文仅记述其中陕西转角羚羊一新种 :灞河陕西转角羚 (Shaanxispirabaheensissp .nov .)。该种主要鉴定特征 :角心长而直 ,粗壮 ,横断面呈圆形 ,具一条发育的棱 ,起始于前内侧 ,右角上呈顺时针方向旋转一周至一周半 (从角基部视 ) ,角心前视向两侧中等程度散开 ,侧视中等后倾 ,起始于眼眶之后 ,角柄短。鼻骨窄 ,向上隆起 ,其后缘延伸至眼眶前棱之后的位置 ;眶前窝较宽浅 ,面部长而窄 ;角前额顶宽、稍隆起 ,眶上孔成椭圆形 ,眶上窝发育 ,向前的沟槽可延伸至泪骨 ;额骨在角前轻微隆起 ;角后颅顶宽、短 ,颅轴与面轴交角约为 90°。枕面宽、平 ,面向后下方。牙齿齿冠中等高度。臼齿上底柱与肋皆不发育。p4的下后尖位于下原尖之后 ,在老年个体上与下内尖连接 ,封闭后内谷。与已知大型转角羚羊相比较 ,灞河组新发现的种类与周氏陕西转角羚最为接近。不同的是该新种角心只发育一条棱 ,而不是两条 ,角心横断面呈圆形 ;顶骨较短 ;臼齿列相对较短 ,下臼?  相似文献   

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Abstract  The third instars of the Australian species Hyphydrus lyratus Swartz, H. contiguus Wehncke, H. elegans (Montrouzier) and H. decemmaculatus Wehncke (Coleoptera: Adephaga: Dytiscidae) are described, including a chaetotaxic analysis of the cephalic capsule, head appendages, legs, last abdominal segment and urogomphi. Larvae of these species morphologically resemble other species of Hyphydrus Illiger for which the larvae have been described. A key to identify larvae of the Australian species of Hyphydrus is provided. Larvae of H. effeminatus Watts appear identical to those of H. decemmaculatus . A 822 bp fragment of the CO1 gene of larvae and adults of these species showed very slight differences, suggesting the possibility that, in Australia at least, H. decemmaculatus is polymorphic.  相似文献   

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An almost complete skull and a second partial skull of Bohlinia attica (Artiodactyla: Giraffidae) from the late Miocene of Maragheh in northwestern Iran is described along with a complete upper dentition from Samos in Greece. These specimens enrich what is known of this species. The skull has massive bent ossicones. The braincase is horizontal to the face in lateral view. The upper premolars have strongly curved styles. The metapodials of this species are elongate with a deep plantar trough. The braincase is short and the occipital does not extend caudally. The type material of Bohlinia is from Pikermi but this taxon is also known from localities in F.Y.R.O. Macedonia, Greece, Turkey, Bulgaria, Iraq, and Iran. The new specimens are similar to others except that the ossicone terminates in a knob and the palatine choanae are positioned caudally. This taxon along with Honanotherium, which is most similar but with shorter metapodials and simpler premolars, can be placed in the subfamily Bohlininae.  相似文献   

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详细记述了发现于云南武定早泥盆世晚期利沃鱼(Livosteus)的新材料.依据其特征,建立一个新种.这是该属除在拉脱维亚以外在世界上的第二个种,因此具有重要的古生物地理意义.  相似文献   

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重新描述了江苏句容与安徽和县的裂齿鱼类化石 ,将华南下扬子区已记述的 2属 5种裂齿鱼类化石初步修订为 1属 2种———江苏张氏鱼 (Zhanginajiangsuensis)和扬子张氏鱼(Zhanginayangtzensis)。由于目前对裂齿鱼类的系统发育关系所知甚少 ,张氏鱼属的系统位置以及相关的动物地理分布问题仍有待深入的研究。  相似文献   

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