Caulostrepsis spiralis isp. nov., Miocene grand bay formation of Carriacou (Grenadines,lesser antilles)

A new ichnospecies of Caulostrepsis, namely C. spiralis, is described from the Miocene Grand Bay Formation of Carriacou (Grenadines, Lesser Antilles). It occurs in association with C. cretacea and a common progenitor, most likely a spionid polychaete annelid, was most likely for its production.     

New records for prehistoric introduction of Neotropical mammals to the West Indies: evidence from Carriacou,Lesser Antilles

Aim This paper investigates the prehistoric introduction of five mammalian taxa to Carriacou (Lesser Antilles) and refines the known anthropogenic ranges for these fauna in the pre‐Columbian West Indies. The importance of such records for understanding the region’s historical biogeography and ecology is considered. Location Carriacou Island, Grenada (12°28′ N, 61°26′ W). Methods Zooarchaeological assemblages from Carriacou’s earliest documented prehistoric sites, Grand Bay and Sabazan, were analysed, and exotic taxa were identified and quantified. The timing of introductions was established based on multiple radiocarbon assays, including three new accelerator mass spectrometry (AMS) direct dates obtained on the bone of exotic taxa. Source species and location(s) are considered and compared with known prehistoric records for the Caribbean to synthesize anthropogenic distributions for the pre‐Columbian period. The contexts of the zooarchaeological remains are evaluated to better understand the nature and purpose of introductions. Results Zooarchaeological investigation on Carriacou reveals the occurrence of multiple mammal introductions from South American between c. ad 700 and ad 1400. This paper presents the first records for guinea pig (Cavia sp.), armadillo (Dasypus sp.), peccary (Tayassu/Pecari sp.), opossum (Didelphis sp.) and agouti (Dasyprocta sp.) from the island. Human‐mediated transport of these taxa is indicated by their absence from the record prior to human settlement of Carriacou. Several translocated species are either rare or entirely unknown for the region, and overall West Indian distributions are temporally and spatially discontinuous. Archaeological contexts indicate that mammalian introductions arose from human subsistence needs, but other social factors may have shaped the dispersal of these fauna. Main conclusions The taxonomic combination and richness of Carriacou’s introduced fauna are unusual within the region. Importantly, the new records significantly improve the known pre‐Columbian geographic range for peccary, guinea pig and armadillo. Integrated with regional records, these data augment our understanding of the Caribbean’s historical biogeography, and have the potential to improve our understanding of human mobility and anthropogenic environmental impacts in the West Indies prior to the arrival of Europeans.     

Taphonomy of the middle Cambrian (Drumian) Ravens Throat River Lagerstätte,Rockslide Formation,Mackenzie Mountains,Northwest Territories,Canada

The Middle Cambrian (series 3, Drumian, Bolaspidella Biozone) Ravens Throat River Lagerstätte in the Rockslide Formation of the Mackenzie Mountains, northwestern Canada, contains a Burgess Shale‐type biota of similar age to the Wheeler and Marjum formations of Utah. The Rockslide Formation is a unit of deep‐water, mixed carbonate and siliciclastic facies deposited in a slope setting on the present‐day northwestern margin of Laurentia. At the fossil‐bearing locality, the unit is about 175 m thick and the lower part onlaps a fault scarp cutting lower Cambrian sandstones. It consists of a succession of shale, laminated to thin‐bedded lime mudstone, debris‐flow breccias, minor calcareous sandstone, greenish‐coloured calcareous mudstone and dolomitic siltstone, overlain by shallow‐water dolostones of the Broken Skull Formation, which indicates an overall progradational sequence. Two ~1‐m‐thick units of greenish calcareous mudstone in the upper part exhibit soft‐bodied preservation, yielding a biota dominated by bivalved arthropods and macrophytic algae, along with hyoliths and trilobites. It represents a low‐diversity in situ community. Most of the fossils occur in the lower unit, and only the more robust components are preserved. Branching burrows are present under the carapaces of some arthropods, and common millimetre‐sized disruptions of laminae are interpreted as bioturbation. The fossiliferous planar‐laminated calcareous mudstone consists of chlorite, illite, quartz silt, calcite and dolomite and is an anomalous facies in the succession. It was deposited via hemipelagic fallout of a mixture of platform‐derived and terrestrial mud. Geochemical analysis and trace‐element proxies indicate oxic bottom waters that only occasionally might have become dysoxic. Productivity in the water column was dominated by cyanobacteria. Fragments of microbial mats are common as carbonaceous seams. Complete decay of soft tissues was interrupted due to the specific sediment composition, providing support for the role of clay minerals, possibly chlorite, in the taphonomic process.     

In deep water: a crinoid–brachiopod association in the Upper Oligocene of Antigua,West Indies

Extant brachiopods and stalked crinoids are found together in the deeper waters of the Caribbean Sea. Analogous brachiopod/crinoid associations have been reported from diverse palaeoenvironments in the Neogene of the region. Studied examples include the Pleistocene of Jamaica (deeper water fore reef), and the Miocene of Jamaica (island slope chalks), Barbados (accretionary prism) and Carriacou (turbiditic siliciclastic shelf). Comparison with analogous modern environments indicates deposition in 150+m water depth. This association has now been extended back into the Late Oligocene. Crinoids and brachiopods both occur in the Antigua Formation of Antigua; both occur high in the formation, implying deeper water in this retrograde succession. They have received little attention from systematists, although the brachiopods Cistellarcula dubia Cooper and Tichosina foresti Cooper have previously been described from the Antigua Formation; to these, we add Cistellarcula sp., Argyrotheca sp. and Tichosina sp. At Half Moon Bay in southeast Antigua, high in the Antigua Formation, we have found columnals of isocrinid crinoids (cf. Isocrinus sp.) associated with rare brachiopods (Terebratulina sp.) in island slope deposits. These taxa provide independent evidence for the deeper water aspect of this part of the Antigua Formation, in beds that also yield large, thin‐walled fossil sponges.     

Zooarchaeological analysis of prehistoric vertebrate exploitation at the Grand Bay Site,Carriacou, West Indies

This paper discusses zooarchaeological analysis of vertebrate faunal specimens from Grand Bay, a Ceramic Age (ca.ad 400–1300) site on the island of Carriacou in the Grenadines, West Indies. Using faunal data to assess subsistence patterns of vertebrate exploitation during late site occupation, we can begin to better understand Grand Bay procurement strategies and coral reef exploitation. Preliminary zooarchaeological results imply that Grand Bay vertebrate exploitation emphasized marine resources over terrestrial resources, with particular emphasis on coral reef habitats and fish. The faunal data are discussed in relation to common patterns of prehistoric vertebrate exploitation in the Caribbean and fishing strategies. The Grand Bay faunal sample also provides a foundation from which to formulate future research foci and question zooarchaeological approaches to understanding prehistoric coral reef exploitation in the Caribbean.     

Evolution of an isolated carbonate bank during Oligocene,Miocene and Pliocene times,Cayman Brac,British west Indies

Summary Modern carbonate sedimentation in the Caribbean Sea commonly occurs on banks that are surrounded and isolated by deep oceanic water. This depositional regime also occurred during the Tertiary, and many islands, such as Cayman Brac, have sequences that evolved in such settings. Cayman Brac is a small (about 39 km²) island, located on the Cayman Ridge, that has an exposed Oligocene to Pliocene succession which encompasses three unconformity-bounded formations. The upper Lower Oligocene Brac Formation is formed ofLepidocyclina limestones and sucrosis dolostones that locally contain numerous bivalves and gastropods. The overlying Lower to Middle Miocene Cayman Formation is formed of pervasively dolomitized mudstones to grainstones that contain an abundant, diverse biota of corals, gastropods, bivalves, foraminifera, and algae. Rhodolites are locally common. The Pliocene Pedro Castle Formation is formed of limestones, dolostones, and dolomitic limestones that contain a biota which is similar to that in the Cayman Formation. The unconformities between the formations represent substantial periods of time during which the previously deposited carbonates were lithified and eroded to produce karst terrains. All facies in the Brac, Cayman, and Pedro Castle formations on Cayman Brac developed on a bank that was no more than 20 km long and 3 km wide. There is no evidence of reef development other than isolated thickets ofStylophora and/orPorites and no systematic stratigraphic or geographic changes in the facies patterns of the formations. Comparison with modern Caribbean banks shows that the depositional regime was primarily controlled by water depth and energy levels. Limestones of the Brac Formation probably accumulated in low-energy conditions in water less than 10 m deep. The overlying Cayman Formation contains facies that formed in water 15 to 30 m deep with good cross-bank circulation. The Pedro Castle Formation formed in slightly shallower water (5–25 m) and lower energy conditions. The disconformities between the packages correlate with world wide eustatic drops in sea level.     

Composition and structure of Caribbean bat (Chiroptera) assemblages: effects of inter‐island distance,area, elevation and hurricane‐induced disturbance

Aim Although bats of the Caribbean have been studied extensively, previous work is largely restricted to zoogeography, phylogeography or the effects of island characteristics on species richness. Variation among islands in species composition that is related to geographical or environmental variation remains poorly understood for much of the Caribbean. Location Caribbean islands, including the Bahamas, Greater Antilles and Lesser Antilles. Methods Using presence–absence data, we assessed the extent to which island area, maximum island elevation, inter‐island distance and hurricane‐induced disturbance affected patterns of composition and nestedness for bats in the Bahamas, Greater Antilles and Lesser Antilles. Analyses were conducted for all species, as well as for two broadly defined guilds: carnivores and herbivores. Results For the Bahamas, only inter‐island distance accounted for variation in species composition between islands. For the Greater and Lesser Antilles, differences in island area and inter‐island distance accounted for differences in species composition between islands. Variation in species composition was not related significantly to differences in elevation or hurricane‐related disturbance. In general, results of analyses restricted to a particular broad guild (i.e. carnivores or herbivores) mirrored those for all bats. Bat species composition was nested significantly in each island group. Nestedness was stronger in the Greater Antilles and in the Lesser Antilles than in the Bahamas. Carnivore assemblages were nested significantly in the Greater and in the Lesser Antilles, but not in the Bahamas. In contrast, herbivore assemblages were nested significantly in each island group. Main conclusions Inter‐island distance had a greater effect on compositional similarity of Caribbean bat assemblages than did island area, elevation or disturbance related to hurricanes. Differential immigration and hierarchical habitat distributions associated with elevational relief are likely to be primary causes for nestedness of Caribbean bat assemblages.     

Deglaciation explains bat extinction in the Caribbean

Ecological factors such as changing climate on land and interspecific competition have been debated as possible causes of postglacial Caribbean extinction. These hypotheses, however, have not been tested against a null model of climate‐driven postglacial area loss. Here, we use a new Quaternary mammal database and deep‐sea bathymetry to estimate species–area relationships (SARs) at present and during the Last Glacial Maximum (LGM) for bats of the Caribbean, and to model species loss as a function of area loss from rising sea level. Island area was a significant predictor of species richness in the Bahamas, Greater Antilles, and Lesser Antilles at all time periods, except for the Lesser Antilles during the LGM. Parameters of LGM and current SARs were similar in the Bahamas and Greater Antilles, but not the Lesser Antilles, which had fewer estimated species during the LGM than expected given their size. Estimated postglacial species losses in the Bahamas and Greater Antilles were largely explained by inferred area loss from rising sea level in the Holocene. However, there were more species in the Bahamas at present, and fewer species in the smaller Greater Antilles, than expected given island size and the end‐Pleistocene/early Holocene SARs. Poor fossil sampling and ecological factors may explain these departures from the null. Our analyses illustrate the importance of changes in area in explaining patterns of species richness through time and emphasize the role of the SAR as a null hypothesis in explorations of the impact of novel ecological interactions on extinction.     

Cyclic sedimentation across a Middle Ordovician carbonate ramp (Duwibong Formation), Korea

Summary The Middle Ordovician Duwibong Formation (about 100 m thick), Korea, comprises various lithotypes deposited across a carbonate ramp. Their stacking patterns constitute several kinds of meter-scale, shallowing-upward carbonate cycles. Lithofacies associations are grouped into four depositional facies: deep- to mid-ramp, shoal-complex, lagoonal, and tidal-flat facies. These facies are composed of distinctive depositional cycles: deep subtidal, shallow subtidal, restricted marine, and peritidal cycles, respectively. The subtidal cycles are capped by subtidal lithofacies and indicate incomplete shallowing to the peritidal zone. The restricted marine and peritidal cycles are capped by tidal flat lithofacies and show evidence of subaerial exposure. These cycles were formed by higher frequency sea-level fluctuations with durations of 120 ky (fifth order), which were superimposed on the longer term sea-level events, and by sediment redistribution by storm-induced currents and waves. The stratigraphic succession of the Duwibong Formation represents a general regressive trend. The vertical facies change records the transition from a deep- to mid-ramp to shoal, to lagoon, into a peritidal zone. The depositional system of the Duwibong Formation was influenced by frequent storms, especially on the deep ramp to mid-ramp seaward of ooid shoals. The storm deposits comprise about 20% of the Duwibong sequence.     

Stratigraphy and sedimentology of Muschelkalk carbonates of the Southern Iberian Continental Palaeomargin (Siles and Cehegín Formations,Southern Spain)

The Triassic sediments of the External Zones of the Betic Cordillera were deposited on the Southern Iberian Continental Palaeomargin. Two coeval Ladinian formations, namely the Siles Formation and the Cehegín Formation, are described to illustrate the facies and lithostratigraphic variability in the Muschelkalk carbonates. There has been some dispute over the number of carbonate units present in the Siles Formation. Our studies assign a tectonic origin to these recurrent carbonate units. Both formations comprise only one carbonate unit, which is correlated to the Upper Muschelkalk of the Catalan and Germanic basins and some Iberian Range sections. To characterize the sedimentological features of these formations, 14 facies were defined. The most widespread sediment was originally lime mud, although bioclastic deposits are also common. In the facies succession, a main transgressive-regressive sequence could be identified. According to the facies model proposed here, a muddy coastal and shallow-water platform prograded over mid ramp deposits. There is no evidence for a seawards reefal or oolitic-bioclastic sandy barrier. The most significant feature of this sedimentary interpretation is that these carbonate facies show clear characteristics of an epicontinental platform.     

Bioclaustration trace fossils in epeiric shallow marine stromatolites: the Cretaceous‐Palaeogene Yacoraite Formation,Northwestern Argentina

The shallow carbonate facies at the top of the Yacoraite Formation (Late Cretaceous–Early Palaeocene) in the Metán sub‐basin, Salta Basin (Cretaceous‐Eocene), northern Argentina, have domal stromatolitic boundstones with peculiar cavities, interpreted here as bioclaustrations. The cavities appear to have been produced by organisms that lived within the microbial mat contemporarily with its growth, producing a distinctive ichnofabric. This is the oldest reported record of bioclaustrations in stromatolites, and the first in shallow marine environments. The interpretation of the facies suggests a stressed shallow, restricted setting with variations in salinity, represented by an intertidal environment with an extensive tidal flat. Bioclaustrations, stromatolites, endobiont Yacoraite Formation (Cretaceous‐Palaeogene), Northwestern Argentina.     

Upper Ordovician microphytoplankton of the Bill's Creek Shale and Stonington Formation,Upper Peninsula of Michigan,U.S.A.: Biostratigraphy and paleogeographic significance

An abundant, diverse, and well-preserved organic-walled microphytoplankton assemblage is described from the Upper Ordovician Bill's Creek Shale and the lower Stonington Formation (Bay de Noc Member) in the Upper Peninsula of Michigan, U.S.A. Based on graptolite and conodont evidence, the Bill's Creek Shale and Stonington Formation are Richmondian (=Ashgill) in age. The assemblage is dominated by acritarchs, which comprise 29 species (including the enigmatic palynomorph Gloeocapsomorpha prisca) assigned to 20 genera. The prasinophyte phycomata are represented by undifferentiated species of Leiosphaeridia and Tasmanites. In addition, chitinozoans are abundant, and scolecodonts and graptolite fragments are common. Paleontologic-palynologic and sedimentologic evidence indicates that the Bill's Creek Shale was deposited in a low-energy, shallow, nearshore marine environment. The overlying Bay de Noc Member of the Stonington Formation also accumulated in a low-energy, normal marine environment, but in a more offshore, somewhat deeper water setting. Both formations experienced minor transgressive and regressive episodes as indicated by fluctuations in the composition of the palynoflora. The combined Bill's Creek/Stonington acritarch assemblage closely resembles those described from the Richmondian-aged Maquoketa Shale (Missouri and Kansas), Sylvan Shale (Oklahoma), and Vauréal Formation (Anticosti Island, Québec, Canada). The overall composition of the acritarch assemblage from these four formations reflects a distinctive, recognizably Laurentian character. Nonetheless, many of the Bill's Creek/Stonington acritarchs have been reported from Upper Ordovician localities elsewhere, providing additional evidence for Late Ordovician cosmopolitanism of the marine microphytoplankton community. Additionally, the restricted stratigraphic range of many of the taxa further enhances their biostratigraphic application, both regionally and globally, and reaffirms the Richmondian (=Ashgill) age of the Bill's Creek Shale and Stonington Formation.     

Palaeoenvironmental implications of the ichnology and geochemistry of the Westbury Formation (Rhaetian), Westbury‐on‐Severn,south‐west England

Abstract: The Westbury Formation (Rhaetian) beds of Westbury Garden Cliff, Westbury‐on‐Severn, west of Gloucester, Britain, show an unusual combination of features. Both deep water and emergent characteristics are present within the sediments and the trace fossils. The ichnoassemblage consists of abundant Selenichnites, Planolites beverlyensis and Lockeia with rarer Oniscoidichnus, Chondrites, Rhizocorallium irregulare, Taenidium serpentium, an unusual form of Walcottia and Merostomichnites‐like traces. These trace fossils display an interesting relationship with the sediments: low‐energy Cruziana ichnofacies is found within high‐energy sandstones. The sandstones are interbedded with laminated mudstones, apparently deposited in deep water, but some aspects of the ichnoassemblage, preservation and sedimentation indicate shallower water. One new trace fossil, Radichnus allingtona igen. et isp. nov., closely resembles the traces of modern fiddler crabs and imply emergence, by analogy. This ichnofauna is similar to early stage disaster colonisation in recent experiments in Long Island Sound (south of Connecticut, USA) and with storm‐influenced deposits within the Cardium Formation (Seebe, Alberta, Canada). This indicates a lagoonal environment with influxes of sand and oxygen. Total organic carbon levels were found to fluctuate greatly between stratigraphic layers but remained relatively high. This implies low oxygen conditions. The abundance of sulphur (in pyrite) also supports an interpretation of anoxic conditions, and low sedimentation rates within the shale layers. A restricted shallow basin or lagoonal environment is proposed for the palaeoenvironment, with fluctuating oxygen influencing diversity.     

Significance of Middle Cambrian mixed carbonate-siliciclastic units for global correlation: southern Nevada, USA

Sequence-stratigraphic analysis of the Middle Cambrian Highland Peak, Bonanza King, Swasey, and Wheeler formations in the Great Basin refines platform-to-basin correlations and distinguishes local tectonic events from eustasy. This analysis provides a reliable sea-level history through the Ptychagnostus gibbus and Ptychagnostus atavus trilobite intervals and confirms that the Global Stratotype Section and Point (GSSP) at the first appearance datum (FAD) of P. atavus was deposited during an overall sea-level rise. Deposition during the Middle Cambrian Ehmaniella/Bolaspidella biozones in the western U.S. is represented by two lithologically distinct successions: (1) a poorly fossiliferous, shallow-water, mixed carbonate-siliciclastic succession that is widespread across southern Nevada and southeastern California and (2) a highly fossiliferous, deeper water, fine-grained, siliciclastic succession in central Nevada and western Utah. The deeper water succession was deposited within the fault-controlled House Range Embayment and contains the P. atavus GSSP. Correlation of these disparate successions had been hampered by a lack of high-resolution biostratigraphic data, and limited chemostratigraphy and sequence stratigraphy. In this study, sequence-stratigraphic analysis indicates that the Condor Member of the Highland Peak Formation and the “mixed unit” of the Bonanza King Formation are the shallow-water platform equivalents of the basal Wheeler Formation encompassing the P. gibbus and lower P. atavus zones. The deepening event that is recorded in the P. gibbus Zone represents a major flooding surface that may be used as an important event marker for regional and global correlation. The overlying P. atavus GSSP, however, is within the later stage of transgression and may represent a globally synchronous event that can be correlated from platform to basin.     

Taphonomy and palaeoecology of Hauterivian–Barremian nerineoid shell beds from the Neuquén Basin,west‐central Argentina

Nerineoid shell beds are described for the first time from Lower Cretaceous deposits of southern South America. These come from carbonates near the top of the Agrio Formation in southern Mendoza Province, west‐central Argentina. To envisage the origin of the nerineoid shell beds, a taphonomic study was carried out, which indicated that these represent within‐habitat time‐averaged, primary sedimentological concentrations with a secondary biogenic imprint related to a relatively high local production of nerineoid shells. The associated palaeoenvironments were studied through a facies analysis of the carbonate succession including the shell beds. The carbonates were deposited in a homoclinal ramp system and depict a shallowing upward trend from mid to inner ramp. The individuals lived and accumulated in oolitic shoals within the inner ramp, in a shallow, well‐lit, high‐energy setting above fair‐weather wave base. Substrate was oxygenated and loose. The nerineoids are shown to belong to one species of the genus Eunerinea, and through the functional morphology of the shells they are tentatively interpreted as infaunal or semi‐infaunal. It is suggested that the recorded monospecific nerineoid shell beds indicate that the palaeoenvironmental conditions may have been favourable for the development of abundant populations of these gastropods in the northern part of the Neuquén Basin during a short time interval in the Hauterivian–Barremian boundary. This could have been related to a brief warming episode, but other factors may have also been involved. □Argentina, Early Cretaceous, Gastropods, nerineoids, Neuquén Basin, shell beds, taphonomy.     

Integrated sedimentological,ichnological and sequence stratigraphical studies of the Koti Dhaman Formation (Tal Group), Nigali Dhar Syncline,Lesser Himalaya,India: paleoenvironmental,paleoecological, paleogeographic significance

Abstract

Integrated ichnology, sedimentology and sequence stratigraphy of the Lower Quartzite Member to the Arkosic Sandstone Member of the Koti Dhaman Formation (Cambrian Series 2, Stage 4), Tal Group, Nigali Dhar Syncline, Lesser Himalayan lithotectonic zone are presented. Trilobite traces of Gondwanan affinity i.e., Cruziana salomonis, Cruziana fasciculata, Rusophycus dispar and Rusophycus burjensis are recorded along with Arenicolites isp. and Skolithos isp. from the Lower Quartzite Member. A rich and diverse ichnoassemblage attributed to the Cruziana ichnofacies is described for the first time from the Arkosic Sandstone Member of the same formation. Seven ichnofossil assemblages, i.e., Cruziana-Rusophycus, Planolites-Palaeophycus, Cruziana problematica, Diplichnites, Cochlichnus anguineus, Bergaueria perata and Psammichnites gigas have been recognized in the Lower Quartzite to Arkosic Sandstone members of the Koti Dhaman Formation. Seven sedimentary facies i.e., sandstone–shale facies (FT1), cross-bedded (trough and planar) sandstone (FT2), bedded sandstone facies (FT3), shale facies (FT4), shale–sandstone facies (FT5), shale-rippled sandstone facies (FT6) and planar and trough cross-laminated sandstone (FT7) and four facies associations FA1-FA4 are identified in the Koti Dhaman Formation. The formation contains shallowing upward parasequences of a tidal flat complex. Overall, two major events are recognized: i) the break in sedimentation between the Lower Quartzite Member and the overlying Shale Member probably related to forced-regressive event and ii) the facies shift from FT6 to FT7 of the Arkosic Sandstone Member represents an erosive transgressive event; the surface is interpreted as wave ravinement surface, which also serves as a sequence boundary. Integrated ichnology, sedimentology and sequence stratigraphic studies indicate that the Lower Quartzite Member was deposited in a shallow subtidal sand sheet complex and tidal flat complex; the Shale Member was deposited in a mud flat setting of a tidal flat complex, and the Arkosic Sandstone Member in a mixed-flat (tidal flat complex) to sand sheet complex front and margin (subtidal sand sheet complex). Overall, the lower to middle part of the Koti Dhaman Formation represents a tide-dominated shallow subtidal–intertidal to mud-flat subenvironments of the tidal flat complex. A palaeogeographic reconstruction of lower Cambrian (516–514?Ma) is presented based on the distribution of trilobite traces from the Lesser Himalaya and the Bikaner–Nagaur area of Peninsular India (eastern Gondwana), Egypt, Jordan, Turkey (western Gondwana) and Canada (Avalonia).     

Adaptive radiation versus intraspecific differentiation: morphological variation in Caribbean Anolis lizards

Lizards in the genus Anolis have experienced adaptive radiation in the Greater Antilles, producing a suite of species morphologically adapted to use different parts of the environment. In the Lesser Antilles, adaptive radiation has not occurred, but on some islands, interpopulational variation is high and represents adaptation to different habitats. We compared the extent of morphological differentiation among Greater Antillean habitat specialists with that exhibited among populations of two species, Anolis marmoratus and A. oculatus, from the Lesser Antillean islands of Guadeloupe and Dominica. Although extensive, intraspecific divergence in the Lesser Antilles is substantially less in magnitude than the differences among habitat specialists in the Greater Antilles. All populations of A. marmoratus are most similar to Greater Antillean trunk‐crown habitat specialists, but populations of A. oculatus differ in their affinities: some are similar to trunk‐crown anoles, but others are more similar to trunk‐ground habitat specialists.     

Stratigraphy and geologic history of a southeastern salt marsh basin,North Inlet,South Carolina,USA

Forty seven vibracores and fifteen radiocarbon dates have beenobtained to outline the Holocene history of the North Inlet saltmarsh basin. Marsh deposits date from about 3500 years BP and havetransgressed over a Late Pleistocene beach-ridge terrain that waspartly eroded by Late Holocene tidal channel meandering. Marsh mudalso has prograded southward over shallow subtidal estuarine Macomamuds which date from about 4500 years BP and which are stillaccumulating in adjacent Winyah Bay. The southward migration of themarsh environment probably is due to the southward migration ofboth North Inlet and the mouth of Winyah Bay. The stratigraphy ofthe North Inlet basin offers no evidence for Late Holocene sea-level oscillations.Application of this model of marsh history to the study long-term ecosystem succession driven by slowly rising sea level isdiscussed.