Polyploidy and variation in theCampanula rotundifolia complex. Part II. (Taxonomic) I. Revision of the groupsSaxicolae,Lanceolatae andAlpicolae in Czechoslovakia and adjacent regions

A survey is given of three natural groups of the subsectionHeterophylla (Witas.)Fed. of the genusCampanula L. Within theSaxicolae four taxa of higher rank have been revealed:c. xylocarpa Kovanda (2n=34),C. gentilis Kovanda (2n=34),C. moravica (Spitzner)Kovanda subsp.moravica (2n=68), andC. moravica subsp.xylorrhiza (O. Schwarz)Kovanda (2n=102). BothAlpicolae andLanceolatae are represented by a single species: the first byC. cochleariifolia Lam. (2n=34), the latter byC. serrata (Kit. ap.Schult.)Hendrych (2n=34). Cytology, ecology and geographical distribution of all these taxa have been reviewed, and relationships to the other members of the complex discussed. Infraspecific variation within each species has also been examined.     

A revision ofTaraxacum sect.Piesis (Compositae)

On the basis of allozyme and cultivation data, and of additional herbarium material, a taxonomic and nomenclatural revision ofTaraxacum sect.Piesis A.J. Richards exKirschner et?těpánek is provided. The section is made up of halophilous, sexually reproducing taxa. InT. stenocephalum Boiss. etKotschy,T. pindicum Kirschner et?těpánek, sp. nov., andT. perenne Kirschner et?těpánek, sp. nov., a tetraploid chromosome number has been recorded, representing the only known case of sexuality at the tetraploid level in the genus. The complex ofT. stenocephalum, includes some geographically and morphologically extreme populations treated as subspecies: subsp.gumusanicum (Soest)Kirschner et?těpánek, comb. nov., subsp.magnum Kirschner et?těpánek, subsp. nov., and subsp.daralagesicum (Schischk.)Kirschner et?těpánek, comb. nov. In addition toT. bessarabicum (Hornem.)Hand.-Mazz., a widely distributed Eurasian species,T. stenocephalum, a complex centred in Transcaucasia and Anatolia, andT. pachypodum H. Lindb., a North African endemic, four new species are described:T. salsum Kirschner et?těpánek, sp. nov., a diploid endemic confined to E Crimea,T. perenne Kirschner et?těpánek, sp. nov., a tetraploid sexual species known only from SW Crimea,T. pindicum Kirschner et?těpánek, sp. nov., a remarkable tetraploid endemic to the Pindos Mts., Greece, andT. salsitatis Kirschner, ?těpänek etYirdirimli, sp. nov., an Anatolian diploid species. Furthermore, a hybrid betweenT. salsum andT. bessarabicum from Crimea (documented on the basis of allozyme data elsewhere) is given a binomial,T. xmesohalobium Kirschner et?těpánek, nothosp. nov.     

Upper Triassic (Keuper) non-marine trace fossils from the Haßberge area (Franconia, south-eastern Germany)

Various trace fossils from the Hassberge Formation and the Löwenstein Formation (Middle Keuper, Upper Triassic) of the Haßberge region are described. Twenty-three different forms have been identified, 17 of which are named, includingCruziana pascens n. isp.,Lockeia cunctator n. isp., andRusophycus versans n. isp.Lockeia siliquaria James, 1879,L. amygdaloides (Seilacher, 1953),L. triangulichnus Kim, 1994, andL. elongata (Yang, 1984) are revised and synonymized under the oldest available name,L. siliquaria James, 1879.Rusophycus eutendorfensis (Linck, 1942) andR. carbonarius Dawson, 1864 are revised. The diagnosis ofPolykladichnus Fürsich, 1981 is emended, and a diagnosis forHelminthoidichnites Fitch, 1850 is given for the first time. Among the described ichnotaxa,Skolithos ispp.,Rusophycus carbonarius, andTaenidium barretti are the most common forms. The trace fossil association is typical of theScoyenia ichnofacies, which indicates non-marine, periodically or completely inundated environments, such as floodplains and lake margins. Two palaeoichnocoenoses are identified. One ichnocoenosis, dominated byCruziana problematica, cf.Polykladichnus isp., andSkolithos isp. B characterizes margins of trough cross-bedded sandstones. Another ichnocoenosis, dominated byRusophycus versans n. isp.,Taenidium barretti,Scoyenia gracilis andSkolithos isp. A is related to ephemeral lake deposits. Taxonomic recommendations for the use of hitherto described and figured invertebrate Keuper trace fossils from Germany are given.     

Foraminifera from the Silberberg Formation («Lower Oligocene») at Silberberg,near Helmstedt (Germany)

From the Silberberg Quarry, type locality of the Silberberg Formation (Martini &Ritzkowski 1968), some planktonic foraminifera are described:Globigerina officinalis Subbotina, Globigerina ouachitaensisHowe &Wallace,G. praebulloides leroyi Blow &Banner, G.praebulloides occlusa Blow &Banner which taxa are here suggested to form intergrading morphotypes within the range of variation of a population dominated byG. officinalis. In addition,G. cf.danvillensis Howe &Wallace is found. The planktonic fauna, together with some elements in the benthonic fauna, suggests a possible correlation with the Danville Landing Member of the Jackson Formation, Louisiana, traditionally ascribed to the Upper Eocene. There is some similarity with European Uppermost Eocene — Lower Oligocene faunas described by various authors. The Silberberg Formation was suggested to be a Neostratotype for the Latdorfian Stage of the Lower Oligocene byMartini &Ritzkowski (1968). It is suggested that the Silberberg Formation may well be coeval with the Upper Priabonian as proposed byCita 1969.     

Verbeitung einiger seltenen und subendemischen Arten der griechischen Flora mit taxonomischen und chorologischen Betrachtungen

The taxa very rare in Greece,Gentianella bulgarica (Velen.)J. Holub andPrimula elatior (L.)Hill subsp.intricata (Gren. etGodron)Lüdi Balkan-South Carpathian and mountain subendemic taxon of South and South-Central Europe also the oreophytesGentiana asclepidaea L. andAndrosace villosa L. rare in Greece at high altitudes endemic species of Central Europe and arcto-alpic element both extending South as far as Central Italy and Central Greece as well as the relatively rare species in Greece,Crocus biflorus Mill. which exists in South Europe from Sicily and eastwards and finally four otherIridaceae, i.e.: the South Balkan Asia-Minor subendemic speciesCrocus olivieri Gay andC. pulchellus Herbert, the South Balkan endemic speciesC. veluchensis Herbert and the Mediterranean and mostly East-para-mediterranean subendemic speciesHermodactylus tuberosus (L.)Miller are reported and investigated. However new habitats and records in the Greek area of the investigated species are given, their distribution is discussed, while the existence of the taxa most rare in GreecePrimula elatior (L.)Hill subsp.intricata (Gren. etGodron)Lüdi inHegi,Gentianella bulgarica (Velen.)J. Holub,Gentiana asclepiadea L.,Androsace villosa L. andCrocus biflorus Mill. are presented in the form of a dot map and thus the southernmost limits of their spreading in the Balkan peninsula or in S.E. Europe are elucidated.     

New names inPhanerogamae 3

New nomenclatural combinations are proposed, resulting from the adoption of a narrower concept of the generic classification unit or from necessary changes of the taxonomic rank in several taxa. A new genus—Anemonidium (Spach)Holub—is proposed inRanunculaceae. Four genera are accepted, includingChamaepitys Hill (Ajuga L. p. p.),Cota J. Gay (Anthemis L. p. p.),Euchiton Cass. (Gnaphalium L. p. p.) andRostraria Trin. (Koeleria Pers. p. p.+Trisetum Pers. p. p.).Genistella Ortega is preferred toChamaespartium Adans. In all 95 new combinations are proposed (83 for species, 11 for subspecies, 1 for a variety), including 21 inCota, 13 inRostraria, 11 inChamaepitys and 10 inEuchiton.     

Reclassifications and new names for some European Phanerogams

On the basis of taxonomic and nomenclatural revisions of various plant groups 37 new nomenclatural combinations are proposed mostly for European phanerogamous taxa (21 for species, 14 for subspecies, 2 for hybrids.)Medicula Medicus,Pseudofumaria Medicus andThlasidia Rafin. are accepted as taxonomically justified genera and correct generic names.Dactylorhiza longebracteata (F. W. Schmidt)Holub is proposed instead ofD. fuchsii (Druce) Soó, andHylotelephium triphyllum (Haworth) Holub isstead ofH. purpureum (L.)Holub.     

LignicolousHyphomycetes from Czechoslovakia 6.Spadicoides andDiplococcium

Seventeen species belonging toSpadicoides Hughes andDiplococcium Grove collected in Central Europe mostly in Czechoslovakia are described and illustrated. Two new species are described inSpadicoides—S. carpatica Hol.-Jech.,S. penatium Hol.-Jech. and three new species inDiplococcium—D. bicolor Hol.-Jech.,D. insolitum Hol.-Jech. andD. parcum,Hol.-Jech.     

The stuttgart collection of oligocene primates from the fayum province of Egypt

In 1907, R. Markgraf made a small collection of Oligocene fossil primates from Egypt for the Staatliches Museum für Naturkunde in Stuttgart. Circumstances of discovery suggest that this collection came from the same stratigraphic horizon (the Upper Fossil Wood zone) and possibly the same locality as a similar collection made earlier that year byMarkgraf for the American Museum of Natural History in New York. Three new genera and species were described bySchlosser, based on the Stuttgart collection.Parapithecus fraasi Schlosser is a junior synonym ofApidium phiomense Osborn. “Parapithecus“grangeri Simons is placed in the new genusSimonsius. Propliopithecus haeckeli Schlosser andMoeripithecus markgrafi Schlosser have previously been recognized as synonyms. In addition,Aegyptopithecus zeuxis Simons is known from the Upper Fossil Wood zone.     

LignicolousHyphomycetes from Czechoslovakia 3.Sporoschisma,Sporoschismopsis andCatenularia

The species of three genera—Sporoschisma Berk. etBr.,Sporoschismopsis Hol.-Jech. etHennebert andCatenularia Grove are treated in this paper from Czechoslovakia:Sporoschisma juvenile Boudier,S. mirabile Berk. etBr.,Sporoschismopsis moravica Hol.-Jech. etHennebert andCatenularia cuneiformis (Richon) Mason. The studied fungi produce the conidia in basipetal succession, endogenously or subendogenously, quite internally in large phialides or only inside open swollen collarettes.     

Evolutionsmodelle für den Ursprung der Echinodermen: Zusammenfassung und Kritik

The origin of echinoderms is one of the most crucial questions within the evolutionary history of deuterostomes. An ancestral position was suggested byGarstang, Romer andNichols. They also assumed that hemichordates and chordates are sistergroups. In all other hypotheses the echinoderms took a more derived position.Gislén, Jefferies andHolland viewed the hemichordates as basal to the deuterostomes and postulated that echinoderms and chordates are sistergroups. According toJollie, Peterson et al. andMooi &; David echinoderms and hemichordates are sistergroups.Gudo andGutmann adopted the view ofMetschnikoff who combined the hemichordates and echinoderms in the Ambulacraria; they supposed that echinoderms were derived from pterobranchs. This variety of views is linked with different approaches to phylogenetic reconstruction utilized by each of the authors.Garstang, Romer, Jefferies andGislén compared morphological features, in the case ofGislén andJefferies with some attention to fossil evidence, whereasJollie, Holland andGislén also considerd embryological aspects.Mooi &;David as well asPeterson et al. used modern embryological (epigenetical) approaches.Nichols combined functional morphology and comparative anatomy. Evolutionary scenarios were reconstructed only by a few authors.Holland associated the development of echinoderms from pterobranch-like ancestors with repeated changes in feeding modes.Nichols envisioned that echinoderms had evolved from sipunculids that gained protection from predators through skeletal armor. In our own investigations based on constructional morphology echinoderms are interpreted as highly derived chordates.