Oceanographic significance of Pacific late miocene calcareous nannoplankton

An analysis of the variability in the composition and distribution of Pacific Late Miocene calcareous nannoplankton about their average biogeography shows that there are primarily two environmental factors causing that variability, climate and dissolution. Climate produces a latitudinal, biogeographic differentiation of the Late Miocene nannoflora, while selective dissolution superimposes a bathymetric differentiation of the nannoflora on that due to climate. Together, these two factors produce three distinct Late Miocene nannofloral assemblages, a high-latitude, temperate assemblage characterized by Reticulofenestra pseudoumbilica and Coccolithus pelagicus, and two tropical assemblages, their differences in composition depending on water depth and surface-water productivity: (1) in shallower water and beneath areas of higher organic production and sedimentation of calcite there is an undissolved assemblage characterized by sphenoliths, small elliptical placoliths and Coccolithus pataecus; (2) in deeper water and areas of lower productivity there is a dissolved assemblage dominated by discoasters.Selective dissolution produces most of the apparent biogeographic variation in Pacific Late Miocene nannoplankton compositions, the variation in compositions observed between the seventeen sites studied. Dissolution preferentially removes the more soluble constituents of the tropical nannoflora so that increasing dissolution tends to give tropical nannoflora a cooler, more temperate aspect. At the same time, selective dissolution shifts the composition of the warmer, tropical component towards its more resistant taxa.Nannoplankton records show a period of greatly decreased calcite dissolution in deep tropical and temperate South Pacific sites between about 8 and 10 m.y. ago. This decrease is strongly correlated with a temporary increase in the ¹³C composition of Pacific deep waters. Calcite dissolution increased during this same period in the deep North Pacific.Nannoplankton records of Late Miocene climate in the tropics are distinctly different from those at higher, south temperate latitudes. Tropical records show a sharp warming in the earliest Late Miocene after a generally cool late Middle Miocene. This was followed by a temporary cooling, nearly to Middle Miocene levels, about 7 m.y. ago. Toward the end of the Late Miocene, the tropical Pacific warmed again and remained warm into the Pliocene. Warming of temperate climates occurred much later. Not until latest Miocene did the southern the Pliocene. Warming of temperate climates occurred much later. Not until latest Miocene did the southern temperate latitudes warm appreciably. Southern subpolar climate cooled continuously through the Late Miocene. We attribute the resulting increases in the latitudinal climatic contrast across the southern Pacific Ocean to the development and migration of a strong subtropical convergence.On the basis of the nannoplankton oceanographic records we postulate that beginning about 10.5 m.y. ago Pacific surface circulation became primarily zonal and the production of deep and bottom waters in the Southern Ocean increased sharply. This produced a northward decrease in calcite preservation, an increase in benthic ¹³C, and a strong climatic gradient across southern latitudes. The period of most vigorous deep Pacific circulation ended 7 m.y. ago in response, we speculate, to the reduced ocean salinities during the Messinian.     

Cenozoic radiolarian paleobiogeography: Implications concerning plate tectonics and climatic cycles

A preliminary study of the paleobiogeographic patterns of radiolarian facies during the Paleogene and subsequent time shows that:(1) Through time radiolarian assemblages display distinct faunal provincialism reminiscent of modern faunal distributions correlated with planetary temperature gradients and surface oceanic conditions. The equatorial—tropical radiolarian fauna extended apparently unrestricted across the Pacific Ocean, the Caribbean Sea and the Atlantic Ocean through Early Miocene time. In the Caribbean Sea and the Atlantic Ocean, radiolarians reached their maximum abundance in the Eocene and Oligocene. Subsequently, they gradually declined to virtual disappearance in these areas in the early Miocene. Their Pacific counterparts remained practically undisturbed, except that post early Miocene assemblages there showed a marked trend toward decreasing test thickness. This trend has since been a worldwide characteristic of Neogene radiolarian assemblages and their modern equivalents. It is postulated that the disappearance of radiolarians in the Carribean Sea and the Atlantic Ocean at the end of the Paleogene is related to the onset of the emergence of the isthmus of Panama which interrupted the preexisting oceanic circulation between the Pacific and Atlantic Oceans.(2) Throughout the Paleogene there have been marked sequential fluctuations in the radiolarian assemblages of the Caribbean Sea which indicate intermittent incursions of higher-latitude fauna in this area. Associated with the faunal fluctuations are cyclic variations in the total carbonate of the sediment with patterns also comparable in duration to Pleistocene carbonate cycles in the equatorial Pacific known to have been induced by climatic changes. Based on similarities with Pleistocene climatic cycles in the equatorial Pacific and elsewhere, it is surmised that the faunal and lithologic fluctuations observed in Paleogene radiolarian sediments were also induced by the biologic and physico-chemical processes associated with worldwide changes in the climatic conditions of that time.     

The distribution of radiolarian assemblages in the modern and ice-age Pacific

Data on radiolarian abundances from several recent regional studies of the Pacific have been combined with new data from the temperate and tropical area to provide an ocean-wide view of radiolarian distributions. A Q-mode factor analysis of these data identified seven factors which have their areas of dominance in the following regions: tropics, western Pacific, subarctic, Antarctic, transitional zone, temperate regions, and the eastern central water masses. The distributions of these factors tended to follow those of surface water masses and major ocean currents. A more detailed analysis of the temperate and tropical region better delineated the complex flow and counter flow in this area. The relationship between the distribution of modern radiolarian assemblages and the surface circulation of the Pacific can be used to deduce the nature of oceanographic changes which occurred in the past. The modern radiolarian distributions are compared with those mapped at the 18, 000 B.P. level and reveal two major differences in the ice-age Pacific. The Tropical Factor, restricted primarily to the eastern half of the ocean in modern times, ranged across the entire ocean at 18, 000 B.P. and extended into the area of the western boundary currents. The Subarctic Factor, now found mainly in the western subarctic, expanded to the east and south at 18, 000 B.P. and had strong similarities to an assemblage found in the subantarctic area. The expansion of these two dominant assemblages was at the expense of the Western Pacific, Temperate, and Transitional Factors. These differences in the 18, 000 B.P. distributions are thought to be caused by an increase in the influence of arctic air masses in the North Pacific and by a general increase in the wind-driven zonal flow.     

Early Pleistocene glacial-interglacial radiolarian assemblages from the eastern equatorial Pacific

Polycystine radiolaria from ODP Hole 677A in the eastern equatorialPacific were examined at isotopically identified Early Pleistoceneglacial maxima and minima. Two distinct radiolarian assemblagesare recognized, characterizing glacial and interglacial optima.The Glacial Assemblage is characterized by high abundances ofTheocalyptra davisiana, Botryostrobus auritus, Anthocyrtidiumzanguebaricuim andHexacontium enthacanthum. The InterglacialAssemblage is characterized by Tetrapyle octacantha, Octapylestenozoa and Theocorythium vetulum. A comparison of these fossilassemblages with modern radiolarian distribution suggests thatthe Glacial Assemblage represents intensified upwelling of coldadvected water via the Eastern Pacific Boundary Current, whilstthe Interglacial Assemblage indicates climatic ameliorationin the eastern equatorial Pacific, with the prevalence of warm(>21C) tropical/subtropical surface waters. The recognitionof these radiolarian assemblages could be successfully appliedto studies of adjacent east Pacific areas where other palaeoecologicalindicators are lacking.     

Radiolarian faunal turnover across the Oligocene/Miocene boundary in the equatorial Pacific Ocean

The global warming trend of the latest Oligocene was interrupted by several cooling events associated with Antarctic glaciations. These cooling events affected surface water productivity and plankton assemblages. Well-preserved radiolarians were obtained from upper Oligocene to lower Miocene sediments at Ocean Drilling Program (ODP) Leg 199 Sites 1218 and 1219 in the equatorial Pacific, and 110 radiolarian species were identified.Four episodes of significant radiolarian faunal changes were identified: middle late Oligocene (27.5 to 27.3 Ma), latest Oligocene (24.4 Ma), earliest Miocene (23.3 Ma), and middle early Miocene (21.6 Ma). These four episodes approximately coincide with increases and decreases of biogenic silica accumulation rates and increases in δ¹⁸O values coded as “Oi” and “Mi” events. These data indicate that Antarctic glaciations were associated with change of siliceous sedimentation patterns and faunal changes in the equatorial Pacific.Radiolarian fauna was divided into three assemblages based on variations in radiolarian productivity, species richness and the composition of dominant species: a late Oligocene assemblage (27.6 to 24.4 Ma), a transitional assemblage (24.4 to 23.3 Ma) and an early Miocene assemblage (23.3 to 21.2 Ma). The late Oligocene assemblage is characterized by relatively high productivity, low species richness and four dominant species of Tholospyris anthophora, Stichocorys subligata, Lophocyrtis nomas and Lithelius spp. The transitional assemblage represents relatively low values of productivity and species richness, and consists of three dominant species of T. anthophora, S. subligata and L. nomas. The characteristics of the early Miocene assemblage are relatively low productivity, but high species richness. The two dominant species present in this assemblage are T. anthophora and Cyrtocapsella tetrapera. The most significant faunal turnover of radiolarians is marked at the boundary between the transitional/early Miocene assemblages.We also reviewed changes in other microfossil assemblages in the low latitudes during the late Oligocene through early Miocene. The microfossil assemblages of major groups show sequential changes near the Oligocene/Miocene (O/M) boundary (23.8 Ma). Many extinction events and some first occurrences of calcareous nannofossils and many occurrences of radiolarians are found from about 24.8 to 23.3 Ma, and first occurrences of planktic foraminifers and diatoms followed from 23.2 through 22 Ma. Hence, the O/M boundary is identified as a significant level for microfossil evolutions.     

Biochronology and paleoclimatic implications of Middle Eocene to Oligocene planktic foraminiferal faunas

Planktic foraminiferal assemblages have been analyzed quantitatively in six DSDP sites in the Atlantic (Site 363), Pacific (Sites 292, 77B, 277), and Indian Ocean (Sites 219, 253) in order to determine the nature of the faunal turnover during Middle Eocene to Oligocene time. Biostratigraphic ranges of taxa and abundance distributions of dominant species are presented and illustrate striking similarities in faunal assemblages of low latitude regions in the Atlantic, Pacific and Indian oceans. A high resolution biochronology, based on dominant faunal characteristics and 55 datum events, permits correlation between all three oceans with a high degree of precision. Population studies provide a view of the global impact of the paleoclimatic and paleoceanographic changes occurring during Middle Eocene to Oligocene time.Planktic foraminiferal assemblage changes indicate a general cooling trend between Middle Eocene to Oligocene time, consistent with previously published oxygen isotope data. Major faunal changes, indicating cooling episodes, occur, however, at discrete intervals: in the Middle Eocene 44-43 Ma (P13), the Middle/Late Eocene boundary 41-40 Ma ( ), the Late Eocene 39-38 Ma ( ), the Eocene/Oligocene boundary 37-36 Ma (P18), and the Late Oligocene 31-29 Ma ( ). With the exception of the boundary, faunal changes occur abruptly during short stratigraphic intervals, and are characterized by major species extinctions and first appearances. The Eocene/Oligocene boundary cooling is marked primarily by increasing abundances of cool water species. This suggests that the boundary cooling, which marks a major event in the oxygen isotope record affected planktic faunas less than during other cooling episodes. Planktic foraminiferal faunas indicate that the boundary event is part of a continued cooling trend which began during the Middle Eocene.Two hiatus intervals are recognized in low and high latitude sections at the Middle/Late Eocene boundary and in the Late Eocene ( ). These hiatuses suggest that vigorous bottom water circulation began developing in the Middle Eocene, consistent with the onset of the faunal cooling trend, and well before the development of the psychrosphere at the boundary.     

Biostratigraphy and paleoceanographic significance of Paleocene radiolarians from offshore eastern New Zealand

A 100-m-thick Paleocene sequence of mainly pelagic sediments at ODP Site 1121, on the eastern flanks of the Campbell Plateau, contains few to common radiolarians of relatively low diversity in the lower 40 m (Early to early Late Paleocene) and abundant, diverse radiolarian assemblages in the upper 60 m (mid-Late Paleocene). The 150 taxa recorded from the entire Paleocene interval are thought to under-represent the actual species diversity by at least one half as many morphotypes have not been differentiated below the level of genus. Assemblages in the lower 40 m are similar to those described from onland New Zealand and DSDP Site 208 (northern Lord Howe Rise); they are correlated with South Pacific radiolarian zones RP4 and RP5. Assemblages in the upper 60 m differ from other known Late Paleocene assemblages in the great abundance of plagiacanthids and cycladophorids. Similarities are noted with later Cenozoic cool-water assemblages. This upper interval is correlated with South Pacific zone RP6, as revised herein, based on comparison with faunas from Site 208 and Marlborough, New Zealand. The interval is also correlated with the upper part of North Atlantic zone RP6 (RP6b–c) based on the presence of Aspis velutochlamydosaurus, Plectodiscus circularis and Pterocodon poculum. Other species, such as Buryella tetradica and Buryella pentadica, are valuable for local correlation but exhibit considerable diachroneity between the Pacific, Indian and Atlantic Oceans. An age model for the Paleocene interval at Site 1121, based on well-constrained nannofossil and radiolarian datums, indicates that the rate of compacted sediment accumulation doubles from 15 to 30 mm/ka at the RP5/RP6 zonal boundary. In large part this is due to a sudden and pronounced increase in accumulation rates for all siliceous fossils; radiolarians and larger diatoms increase from <100 to >10 000 specimens/cm²/ka. This apparent increase in biosiliceous productivity is age-equivalent to a mid-Paleocene cooling event (57–59 Ma) identified from global stable isotope records that is associated with the heaviest δ¹³C values for the entire Cenozoic.     

Paleomagnetic and geochronologic calibration of latest oligocene to pliocene radiolarian events,Equatorial Pacific

Eight chronologically overlapping piston cores from the central Equatorial Pacific, which range in age from latest Oligocene to Pleistocene and for which the magnetostratigraphy was known, served as a basis for a paleomagnetic and geochronological calibration of 75 radiolarian first appearances and extinctions. The species studied are commonly accepted as useful stratigraphic indices.Comparison with results of detailed studies of tropical Pacific DSDP drill-sites reveals excellent agreement in the sequence and timing of first appearances, as well as concurrence of many extinctions. The density of radiolarian events is highest in the Early Miocene, where an average of 0.24 my elapsed between each event. In the Late Miocene the density of events is lowest: not only is each datum level separated by an average of 0.52 my in this time unit, but it also contains the longest eventless interval (2.5 my) of the last 27 my.     

Radiolarian-based transfer functions for estimating paleooceanographic conditions in the south Indian Ocean

A quantitative analysis of 37 radiolarian species in 58 deep-sea surface-sediment samples from the subtropical to the polar regions of the Indian Ocean produced four geographically distinct faunal assemblages (transitional, antarctic, subtropical, subantarctic). Geographic distributions of these assemblages coincide with present-day patterns of sea-surface temperature and water masses. The antarctic factor is almost exclusively found south of today's Antarctic Polar Front. Highest concentrations of the transitional factor are recorded at sites positioned between today's Subtropical Convergence and the Polar Front. The subtropical factor is dominant in sites north of today's Subtropical Convergence.Values of these four faunal assemblages in the surface-sediment samples were regressed onto present-day summer and winter temperatures of the surface waters overlying each of the core-top sites. Resulting transfer functions yield temperature estimates which compare favorably with observed (present-day) summer and winter sea-surface temperatures, with low standard errors of estimate (< ± 1.9°C) and no clear geographic pattern in maps of the residuals (difference between observed and estimated sea-surface temperature).     

Morphological characteristics of boreal radiolarians from the Triassic and Late Cretaceous: Comparative analysis

When comparing boreal radiolarian assemblages of the Triassic and Late Cretaceous, significant differences in their morphotypic composition are recognized. In particular, Triassic assemblages are dominated by spherical morphotypes, including pylomate forms of the genus Glomeropyle, while Late Cretaceous assemblages are dominated by prunoid morphotypes without polar spines (genera Prunobrachium and Amphibrachium). It is concluded that, in the course of evolution, high-latitude radiolarian assemblages considerably changed both taxonomically and morphologically. Therefore, it is impossible to determine the uniform morphological and taxonomic characters describing boreal assemblages of the entire Mesozoic. Features of boreal radiolarian assemblages are unique to each geological epoch.     

Ocean basin and depth variability of oxygen isotopes in cenozoic benthic foraminifera

One of the longest, most detailed quantitative records of oceanographic change in the Cenozoic is that provided by oxygen isotope measurements made on the tests of foraminifera. As indicated by measurements on benthic foraminifera, the deep waters of the world ocean have undergone an overall cooling of about 10°C in the Cenozoic. This change has been neither monotonic nor gradual. Rather, it is evidenced by a few, relatively rapid increases in the ¹³O content of the benthic shells. These “steps” in the isotopic record have been associated with major evolutionary changes in the mean state of the deep ocean. The variance around this mean state has also changed through the Cenozoic. From relatively high variance in the Middle Eocene, the oceans showed low variance in the Late Eocene and Oligocene. In the Miocene the variance of the isotopic measurement again increased, reaching a maximum of short duration in the Middle Miocene. This maximum as well as that which occurred during the Late Pliocene and Quaternary, may be attributable to fluctuations in the isotopic composition of the oceans caused by growth and decay of large ice sheets.In the Late Miocene the benthic oxygen isotopes in Atlantic sites less than 3000 m deep have a higher variance than sites at similar depths in the Pacific and Indian Oceans. It is thought that this high variance results from long-term changes in the importance of the cool and salty North Atlantic Deep Water relative to that of the warmer and less saline Antarctic Intermediate Water at Atlantic sites between 1000 m and 3000 m water depth. Such significant differences in benthic isotopic variance between the ocean basins have been demonstrated only in post-Middle Miocene intervals.     

Equatorial Pacific sea surface temperatures, faunal patterns, and carbonate burial during the Pliocene

A high-resolution record of radiolarian faunal abundances from the eastern equatorial Pacific is compared to records of carbonate and noncarbonate burial to examine the evolution of eastern tropical Pacific climate processes during the Pliocene. These data provide a means to evaluate the sensitivity of the equatorial Pacific to the onset of Northern Hemisphere glaciation around 2.8−2.5 Ma, to the closure of the Isthmus of Panama around 4.4−3.2 Ma, and to orogeny-related weathering changes before 4.0 Ma. Radiolarian faunal assemblages and sea surface temperature (SST) estimates indicate a gradual cooling from early to late Pliocene, but no significant changes occur near the onset of northern hemisphere glaciation. Records of carbonate and noncarbonate mass accumulation show a long term decrease from the Miocene/Pliocene boundary to the upper Pliocene. Greater carbonate burial in the early Pliocene relative to the late Pliocene parallels a gradual cooling from early to late Pliocene, and may reflect changes related to Isthmus closure or widespread orogeny. No significant time domain changes are seen in the eastern equatorial Pacific that could be related to the onset of Northern Hemisphere glaciation.Evolutive spectral analyses of these equatorial Pacific climate parameters indicate that variance in SST and seasonality commonly concentrate at frequencies not linearly related to orbital variations. Furthermore, cross spectral comparisons with a high resolution benthic δ¹⁸O record indicate that the surface ocean and carbonate flux share little coherent variance with high latitude climate processes during the Pliocene. Given the high degree of chronostratigraphic control in these records, these results suggest that Milankovitch-band surface ocean processes as well as carbonate burial in the equatorial Pacific are decoupled from high latitude climate processes during the Pliocene.     

High diversity Pleistocene rainforest Dasyurid assemblages with implications for the radiation of the dasyuridae

It is commonly accepted that dasyurids (Marsupialia: Dasyuridae) radiated in the late Miocene or early Pliocene in response to a drying trend in Australia's climate as evidenced from the high diversity of dasyurids from modern arid environments compared with Miocene rainforest assemblages. However, mid‐Pleistocene dasyurid assemblages from cave deposits at Mt Etna, Queensland are more diverse than any previously known from rainforest habitats. New taxa will be described elsewhere, but include three new genera as well as new species of Dasyurus, Antechinus and Phascogale. Comparison of dasyurids from Mt Etna sites that are interpreted as rainforest palaeoenvironments with fossil and extant assemblages indicate that they are at least as diverse as those from modern arid environments. Thus Neogene diversification of dasyurids occurred in both arid and rainforest habitats, but only the former survived continuing aridification. Hence, aridification cannot be invoked for the diversification of all dasyurid lineages.     

Sea-surface temperature estimates in the Southeast Pacific based on planktonic foraminiferal species; modern calibration and Last Glacial Maximum

Estimates of sea-surface temperatures based on foraminiferal faunal species suggest that the Eastern Equatorial Pacific Ocean was 3–5°C cooler during the Last Glacial Maximum (LGM) than at present. Analysis of new cores from the Southeast Pacific reveals a likely source of ice-age cooling in variations of the Peru Current. Off southern Peru, LGM ocean temperatures were 6–8°C cooler than at present, consistent with substantial cooling on land inferred from regional glacier advances and ice-core data. In the Southeast Pacific, ice-age foraminiferal assemblages have good modern analogs, and transfer functions that define assemblages based on ancient samples yield results similar to those based on coretop samples. During the LGM, subpolar species dominate the Eastern Boundary Current off Peru and extend to the equator. In contrast, the range of the equatorial upwelling species remains roughly constant. We infer from these data and a heat budget model that equatorward advection of cool water, more than equatorial upwelling, drove LGM cooling of the Eastern Tropical Pacific Ocean.     

Late Quaternary history of atmospheric and oceanic circulation in the eastern equatorial Pacific

Four radiolarian assemblages have been defined in recent seafloor sediments of the equatorial Pacific Ocean. The distribution of these assemblages corresponds to the modern pattern of oceanic circulation and water mass structure in this region: the eastern Pacific shallow permanent thermocline and the Equatorial Undercurrent; Peru Current upwelling and the oxygen minimum; the subtropical water mass; warm western tropical water and the North Equatorial Countercurrent. In twelve cores chosen to transect the region both longitudinally and latitudinally, the distribution of these four assemblages has been reconstructed for six time-intervals during the last 127,000 years: 18,000 B.P. (glacial Stage 2); 36,000 B.P. and 52,000 B.P. (interstadial Stage 3); 65,000 B.P. (glacial Stage 4); 82,000 B.P. and 120,000 B.P. (interglacial Stage 5). Atmospheric and oceanic circulation changes through time have been inferred from the reconstructed microfossil assemblage distributions. Changes in assemblage distributions indicate that variations in intensity, direction and mean position of the tradewinds caused marked changes in the oceanic circulation patterns through the last glacial cycle.Near the end of interglacial Stage 5, the disappearance of the North Equatorial Countercurrent from the eastern Pacific suggests that the mean position of the tradewinds was shifted to the south approximately 5° of latitude relative to the modern position, so that the Northeast trades prevented the flow of the North Equatorial Countercurrent into the eastern Pacific. Near the end of interstadial Stage 3, a change in wind direction occurred from predominantly zonal winds, which enhance equatorial divergence and surfacing of the Equatorial Undercurrent, to more meridional winds, which enhance coastal upwelling associated with the Peru Current.In the tropical Pacific Ocean, late Quaternary changes in atmospheric and oceanic circulation are linked with times of continental ice sheet growth in the Northern Hemisphere (i.e., the interglacial-to-glacial transitions across oxygen isotope stage boundaries $\text{5}\text{4}$ and $\text{3}\text{2}$). The major changes in circulation seem to occur a few thousand years in advance of the glacial episodes, at or near periods of ice sheet growth. This relationship indicates that changes in atmospheric circulation in the tropics led and influenced the development of conditions suitable for polar and continental ice sheet growth in the Northern Hemisphere.     

Late neogene biostratigraphy and paleoceanography of DSDP site 310 Central North Pacific and correlation with the Southwest Pacific

Late Neogene planktonic foraminifera have been examined at Site 310 in the Central North Pacific and their stratigraphic ranges and frequencies are presented here. Blow's (1969) zonation developed for tropical regions has been applied where applicable. Where tropical index taxa are rare or absent in this temperate region, Globorotalia crassaformis, and the evolutionary bioseries G. conoidea — G. conomiozea and G. puncticula — G. inflata have been found useful for zonal subdivisions. A correlation between stratigraphic ranges and frequency distributions of these species at Site 310 in the Central North Pacific, and Site 284 in the Southwest Pacific indicates that these species are relatively consistent biostratigraphic markers in temperate regions of both the North and South Pacific Oceans. An informal zonation for temperate latitudes of the Southwest Pacific has been established by Kennett (1973) and a similar zonal subdivision can be made at Site 310.Paleoclimatic/paleoceanographic interpretations based on coiling ratios, percent abundance, and phenotypic variations of Neogloboquadrina pachyderma indicate four major cold events during early, middle, and late Pliocene, and early Pleistocene. Faunal correlations of these events with similar events elsewhere in the Northeast and Southwest Pacific which have been paleomagnetically dated indicate the following approximate ages for these cold events: 4.7 Ma, 3.0 Ma, 2.6–1.8 Ma. and 1.2 Ma. Faunal assemblages have been divided into three groups representing cool, intermediate, and warmer water assemblages. Cool water assemblages are dominated by >60% N. pachyderma; intermediate temperature faunas are dominated by species of Globigerina and Globigerinita and contain between 20% and 30% N. pachyderma. Warmer water assemblages are dominated by species of Globorotalia and contain <10% N. pachyderma. Frequency oscillations within these groups, in addition to paleotemperature parameters evident in N. pachyderma, afford refined paleoclimatic/paleoceanographic interpretations.     

Late Miocene paleoclimatology: Subantarctic water mass,Southwest Pacific

The Late Miocene—Early Pliocene paleoclimatic history has been evaluated for a deep drilled sediment sequence at Deep Sea Drilling Project Site 281 and a shallow water marine sediment sequence at Blind River, New ealand, both of which lay within the Subantarctic water mass during the Late Miocene.A major, faunally determined, cooling event within the latest Miocene at Site 281 and Blind River coincides with oxygen isotopic changes in benthonic foraminiferal composition at DSDP Site 284 considered by Shackleton and Kennett (1975) to indicate a significant increase in Antarctic ice sheet volume. However, at Site 281 benthonic foraminiferal oxygen isotopic changes do not record such a large increase in Antarctic ice volume. It is possible that the critical interval is within an unsampled section (no recovery) in the latest Miocene.Two benthonic oxygen isotopic events in the Late Miocene (0.5‰ and 1‰ in the light direction) may be useful as time-stratigraphic markers. A permanent, negative, carbon isotopic shift at both Site 281 and Blind River allows precise correlations to be made between the two sections and to other sites in the Pacific region. Close interval sampling below the carbon shift at Site 281 revealed dramatic fluctuations in surface-water temperatures prior to a latest Miocene interval of refrigeration (Kapitean) and a strong pulse of dissolution between 6.6 and 6.2 ± 0.1 m.y. which may be related to a fundamental geochemical change in the oceans at the time of the carbon shift (6.3?6.2 m.y.). No similar close interval sampling at Blind River was possible because of a lack of outcrop over the critical interval.Paleoclimatic histories from the two sections are very similar. Surface water temperatures and Antarctic ice-cap volume appear to have been relatively stable during the late Middle—early Late Miocene (early—late Tongaporutuan). By 6.4 m.y. cooler conditions prevailed at Site 281. Between 6.3 and 6.2 ± 0.1 m.y. the carbon isotopic shift occurred followed, within 100,000 yr, by a distinct shallowing of water depths at Blind River. The earliest Pliocene (Opoitian) is marked by increasing surface-water temperatures.     

The Cenozoic vegetation of the Iberian Peninsula: A synthesis

The aim of this work is to provide a first approach to the evolution of Iberia's vegetation during the Cenozoic (with the exclusion of the Quaternary). The Palaeogene was floristically defined by Palaeotropical elements forming tropical/subtropical rainforests, mangrove swamps, edaphically-mediated laurophyllous forests and leguminous-sclerophyllous communities. During the Miocene, Iberian landscapes were drastically modified due to geographic and climatic changes (mainly cooling and aridification) changes. Open, steppe-like environments developed towards the interior of the peninsula and Arctotertiary elements invaded mountainous and riparian ecosystems, coexisting with or becoming part of evergreen, broadleaved forests of Palaeotropical species. From the Late Miocene onwards these forests suffered changes due to the extinction of taxa, the impact of environmental change on the survivors, and the perturbations caused by the arrival of further Arctotertiary elements. However, several Palaeotropical taxa overcame the environmental and climatic changes of the Miocene and Pliocene to form a part of the modern flora of the Iberian Peninsula.     

Distribution of Chironomidae (Insecta: Diptera) Head Capsules in Recent Sediments of Canadian Arctic Lakes

Thirty-two taxa of chironomid larvae were collected from the sediments of 50 lakes from across the Canadian Arctic Islands. Most chironomid taxa living in the Arctic have wide distributions, with only one taxon, Abiskomyia, showing a clear geographic limitation in this region. Many of these taxa have habitat preferences, among which lake morphometry, pH, nutrients and temperature are important. Due to the complex environmental patterns in the Arctic, lakes in both the northern and southern portion of the Canadian Arctic Archipelago have warmer temperatures and the chironomid assemblages of these two regions resemble each other more than those of the intervening central islands. Chironomid diversity is lowest in the central arctic islands, primarily Devon and Cornwallis Island, where the combination of low nutrients and cold temperatures provide the most severe environment for chironomid survival.     

Palaeoceanographic and palaeoclimatic inferences from Late Cretaceous planktonic foraminiferal assemblages from the Exmouth Plateau (ODP Sites 762 and 763, eastern Indian Ocean)

The evolution of planktonic foraminifera during the Late Cretaceous is marked in the Santonian by the disappearance of complex morphotypes (the marginotruncanids), and the contemporary increasing importance and diversification of another group of complex taxa, the globotruncanids. Upper Turonian to lower Campanian planktonic foraminiferal assemblages from Holes 762C and 763B (Ocean Drilling Program, Leg 122, Exmouth Plateau, 47°S palaeolatitude) were studied in detail to evaluate the compositional variations at the genus and species level based on the assumption that, in the Cretaceous oceans as in the modern, any faunal change was associated with changes in the characteristics and the degree of stability of the oceanic surface waters. Three major groups were recognised based on gross morphology, and following the assumption that Cretaceous planktonic foraminifera, although extinct, had life-history strategies comparable to those of modern planktonics: 1 – r-selected opportunists; 2 – k-selected specialists; 3 – r/k intermediate morphotypes which include all genera that display a range of trophic strategies in-between opportunist and specialist taxa. Although planktonic foraminiferal assemblages are characterised by a progressive appearance of complex taxa, this trend is discontinuous. Variation in number of species and specimens within genera has allowed recognition of five discrete intervals each of them reflecting different oceanic conditions based on fluctuations in diversity and abundance of the major morphotypes. Planktonic forms show cyclical fluctuations in diversity and abundance of cold (r-strategists) and warm taxa (k-strategists), perhaps representing alternating phases of unstable conditions (suggesting a weakly stratified upper water column in a mesotrophic environment), and well-stratified surface and near-surface waters (indicating a more oligotrophic environment). Interval 1, middle Turonian to early Coniacian in age, is dominated by the r/k intermediate morphotypes which alternate with r-strategists. These cyclical alternations are used to identify three additional sub-intervals. Interval 2, aged middle to late Coniacian, is characterised by the increasing number of species and relative abundance of k-strategists. After this maximum diversification the k-strategists show a progressive decrease reaching a minimum value in Interval 3 (early to late Santonian), which corresponds to the extinction of the genus Marginotruncana. In the Interval 4, latest Santonian in age, the k-strategists, represented mainly by the genera Globotruncana, increase again in diversity and abundance. The last Interval 5 (early Campanian) is dominated by juvenile globotruncanids and r-strategists which fluctuate in opposite phase. The positive peak (Interval 2) related to the maximum diversification of warm taxa (k-strategists) in the Coniacian seems to correspond to a warmer episode. It is followed by a marked decrease in the relative abundance of warm taxa (k-strategists crisis) with a minimum in the late Santonian (Interval 3), reflecting a decrease in temperature. Detailed analysis of faunal variations allows the Santonian faunal turnover to be ascribed to a cooling event strong enough to cause the extinction of the marginotruncanids.