| 高等植物脱落酸生物合成途径及其酶调控 |
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| 引用本文： | 万小荣,李玲.高等植物脱落酸生物合成途径及其酶调控[J].植物学报,2004,21(3):352-359. |
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| 作者姓名： | 万小荣 李玲 |
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| 作者单位： | （华南师范大学生命科学学院 广州 510631） |
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| 基金项目： | 教育部重点科研项目,the,‘Thousand-hundred-ten,project',cultivating,excellent,personnel,from,the,Department,of,Education,of,Guangdong,province |
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| 摘 要： | 脱落酸（ABA）生物合成一般有两条途径：C15直接途径和C40间接途径, 前者经C15法呢焦磷酸（FPP）直接形成ABA；后者经由类胡萝卜素的氧化裂解间接形成ABA, 是高等植物ABA生物合成的主要途径。9-顺式环氧类胡萝卜素氧化裂解为黄质醛是植物ABA生物合成的关键步骤, 然后黄质醛被氧化形成一种酮, 该过程需NAD为辅因子, 酮再转变形成ABA-醛, ABA-醛氧化最终形成ABA。在该途径中,玉米黄质环氧化酶（ZEP）、9-顺式环氧类胡萝卜素双加氧酶（NCED）和醛氧化酶（AO）可能起重要作用。
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| 关 键 词： | 脱落酸 生物合成 NCED基因 AO基因 酶调控 |
| 收稿时间： | 2003-05-12 |
| 修稿时间： | 2003-09-13 |
Pathways and Related Enzymes of ABA Biosynthesis in Higher Plants |
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| WAN Xiao-Rong LI Ling.Pathways and Related Enzymes of ABA Biosynthesis in Higher Plants[J].Bulletin of Botany,2004,21(3):352-359. |
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| Authors: | WAN Xiao-Rong LI Ling |
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| Affiliation: | (College of Life Science, South China Normal University, Guangzhou 510631) |
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| Abstract: | Two pathways have been proposed for the biosynthesis of ABA: the ‘direct pathway’ and ‘indirect pathway’. In the direct pathway, which operates in some fungi, ABA is derived from farnesyl diphosphate; in indirect pathway, ABA is produced from the oxidative cleavage of carotenoids. The first committed step for ABA biosynthesis in plants is the oxidative cleavage of a 9-cisepoxycarotenoid(C40) to produce xanthoxin(C15) and a C25 by-product(Fig.2). Xanthoxin is then oxidized to a ketone by an NAD-requiring enzyme. Consequently, there is a nonenzymatic desaturation
of the 2'-3' bond and opening of the epoxide ring to form abscisic aldehyde. In the final step of the pathway, abscisic aldehyde is oxidized to ABA. These endogenous enzymes such as zeaxanthin
epoxidase(ZEP), 9-cis-epoxycarotenoid dioxygenase(NCED) and aldehyde oxidase(AO) may play key roles in the regulation of ABA biosynthesis in higher plants. |
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| Keywords: | , |
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